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BIG IDEA #1 The process of evolution drives the diversity and unity of life. MECHANISMS OF VARIATION  Natural selection acts on phenotypic variations in populations  Sources of variation:      Mutation Random assortment during meiosis Crossing Over Random Fertilization Diploidy  Allele Frequencies can be altered by:  Gene Flow: Immigration and Emigration  Genetic Drift: small populations  Mating Patterns: Inbreeding and Sexual Selection T YPES OF SELECTION  Stabilizing Selection: favors intermediate phenotype (heterozygote advantage)  Directional Selection: favors one extreme over another  Disruptive Selection: favors both extremes over the intermediate Campbell, Neil A. Reece; Jane B., BIOLOGY, 6th Edition 2002 PATTERNS OF EVOLUTION  Convergent Evolution: two dissimilar populations evolve similar traits b/c of similar selective pressures. Ex: dolphin and shark  Parallel Evolution: similar to convergent however, organisms do not need to occupy the same niches. Ex: warning colors of many organisms  Divergent Evolution: organisms from a common ancestor become less similar (adaptive radiation) Ex: Galapagos Tortois EVIDENCE FOR EVOLUTION      DNA Amino acid sequence/similar proteins Analogous structures Vestigial structures Homologous structures CONDITIONS FOR HARDY-WEINBERG EQUILIBRIUM      Large population Random mating No mutations No gene flow No natural selection DETERMINING ALLELE FREQUENCIES  Frequency of dominant allele if frequency of recessive allele is given  p if q is given  Frequency of recessive allele if the % of the population with the recessive phenotype is given  q if q 2 is given  Calculate the % of the population with recessive allele if the % of the population expressing the dominant allele is given  q 2 if p 2 +2pq SPECIATION  Speciation occurs when populations accumulate enough changes over time to lead to the emergence of a new species.  Types:  Allopatric—geographic barriers  Sympatric—reproductive barriers  Polyploidy in plants leads to new species b/c the polyploids can not breed with the diploid ancestors MECHANISMS FOR REPRODUCTIVE ISOLATION  Prezygotic Isolating Mechanisms:       Geographic (Habitat) Isolation Ecological Isolation Behavioral Isolation Temporal Isolation Mechanical Isolation Sexual Isolation  Postzygotic Isolating Mechanisms:  Hybrid Sterility  Hybrid Inviability  Zygote Mortality PRE AND POST ZYGOTIC MECHANISMS FOR REPRODUCTIVE ISOLATION ORIGINS OF LIFE  Miller and Urey’s Experiment  Amino acid monomers, polymers, protobiont, first cells  Characteristics of the First Cells       Unicellular Heterotrophic Prokaryotic Simple lipid membrane Ribosomes RNA  Autotrophic prokaryotes would appear soon after ORIGINS OF COMPLEX CELLS  Theory of Endosymbiosis — Large eukaryotic cells evolved when a small prokaryotic cells was engulfed by a larger prokaryotic cell and they developed a symbiotic relationship where both benefitted.  Smaller one eventually evolves into mitochondria (in heterotrophs) or chloroplasts (in autotrophs).  Evidence: Mitochondria and Chloroplasts have their own DNA and ribosomes. They are about the size of prokaryotes. Their membranes are similar to prokaryotes. DIVERSIT Y OF LIFE  Three Domains  Bacteria  Archae  Eukarya  Six Kingdoms       Eubacteria Archaebacteria Protista Fungi Plantae Animalia CLADOGRAMS  Cladograms show relative relatedness between a group of organisms