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Plant Anatomy MUPGRET Workshop March 27, 2004 Definitions Anatomy Morphology Study of form and basic organization. Study of tissue organization. Physiology Study of normal organism function. Life Cycle Juvenile Vegetative Adult Reproductive Plant Structure Two main organ systems: shoot and root Shoot Above ground Leaves, buds, stems, flowers, fruits Root Below ground roots, tubers, rhizomes Plant Cells Formed at meristems. Mitosis in meristem produces new cells. Two types of meristems. Apical – produces primary growth, ex. tip of root or shoot. Lateral – produces secondary growth, ex. cambium. Apical Meristem Lateral Meristem Cells are grouped into tissues. Dermal Ground Vascular Cell Types Epidermal Ground Vascular Zea mays L. leaf cross section Dermal Tissue Covers outer surface of herbaceous plants. Composed of epidermal cells that secrete the waxy cuticle. Waxy cuticle protects against water loss. Ground Tissue Bulk of the primary plant body. Composed of parenchyma, collenchyma, and sclerenchyma. Parenchyma Parenchyma pineapple Collenchyma Celery Schlerenchyma hau – used to make rope Vascular Tissue Transports food, water, hormone, and minerals. Composed of xylem, phloem, parenchyma, and cambium. Vascular tissue Xylem Dutchmens pipe Dicot stem Vegetative Structures Roots Stem Leaf Roots Tap root Fibrous root Adventitious root Tuberous root Aerial root Roots Tuberous Taproot Fiberous Taproots Stem Anatomy Bud – undeveloped shoot. Node – Location of leaf or bud on stem. Internode – Space between nodes. Pith – Spongy tissue in the center of the stem. Lenticel – Pore in the outer layer of the stem. Stem Stem Modifications Tuber – underground stem with nodes Rhizome – underground stem with buds Stolon – aboveground stem with shoot buds Bulb – underground stem with fleshy leaves Corm – underground stem with papery leaves Modified Stem Tendril Thorn Onion set Rhizome Modified Stem II Corm Asparagus Stolon Growth Form Herb Shrub Tree Vine Growth Cycle Annual – single season Biennial – two seasons Perennial – multiple seasons Evergreen – leaves persist > 2 seasons Decidious – leaves die in cold or dry Leaf Parts Petiole Blade Stipule Axillary bud Leaf Structure Blade Stipule Axillary Bud Petiole Leaf Arrangement Alternate Opposite Whorled Leaf form Simple Compound Pinnate Palmate Vein Arrangement Pinnate Palmate Parallel Dichotomous Dichotomous Leaf Shape Ovate Elliptic Oblong Lanceolate Linear Orbicular Cordate Hastate Sagittate Peltate Perfoliate Terete Leaf Margins Entire Serrate Doubly serrate Dentate Crenate Undulate Revolute Crisped Lobed Misc. Vegetative Structures Thorns Spines modified stems (Honey Locust) modified leaves (Cactus) Prickles epidermal outgrowth (Rose) Reproductive Structures Flowers Fruits Floral Parts Pedicel Sepal Petal Perianth Stamen Carpel Pistil Flower Floral Symmetry Actinomorphic Many axes of symmetry, ie. the same wherever you cut it. Zygomorphic One axis of summetry, ie. can only cut one way. Inflorescence Types Solitary Spike Spadix Spikelet Whorl Panicle Raceme Umbel Compound Umbel Fruit Types Dry, indehiscent fruit Dry, dehiscent fruit Fleshy fruit Other Dry, indehiscent fruits Achene (lettuce) Samara (maple) Caryposis (wheat) Nut (almond) Dry, Dehiscent Fruit Legume (soybean) Capsule (tobacco) Silique (Arabidopsis) Schizocarp (maple) Fleshy Fruits Drupe (peach, nectarine) Berry (tomato) Pepo (cucumber) Hesperidium (citrus) Hip (rose) Pome (apple, pear) Other Fruit Types Aggregate Multiple mature ovaries from separate pistils of one flower (ex. raspberry) mature ovaries from separate pistils of several flowers (ex. pineapple) Accessory fruit is something other than ovary tissue (ex. strawberry is a swollen receptacle, seeds are achenes) Placentation Marginal Axile Free central Apical Parietal Basal Ovary Position Hypogynous (above calyx) Perigynous (within the floral cup) Epigynous (below calyx) Monocots vs. Dicots One cotyledon Parallel leaf veins Flower parts often in multiples of three Vascular bundles scattered in stem Two cotyledons Netlike leaf veins Flower parts often in multiples of 4 or 5 Vascular bundles in a ring in the stem