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Basal Tricolpates
Finally, true dicots
Basal Angiosperms, view 1
Basal Angiosperms, view 2
Characters of Tricolpates
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Tricolpate pollen
Floral parts whorled,
members alternating
Staminal filaments
slender
Sieve element plastids
with starch grains
Nuclear & chloroplast
sequence data
Basal Tricolpates (true dicots)
Basal Tricolpates – Utah View
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Ranunculales, order of buttercup family,
is important
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Ranunculaceae, Berberidaceae,
Papaveracae
Platanales of some interest
Proteales – fascinating but mostly
southern hemisphere
Ranunculales
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Mostly herbaceous
Toothed to lobed or compound leaves
Floral parts distinct, calyx, often many
stamens, ovary superior, seeds with tiny
embryos and much endosperm
Alkaloid berberine
Nuclear and chloroplast DNA
Papaveraceae (inc. Fumariaceae)
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Deciduous sepals
Carpels 2, connate
Fruit a capsule
Nearly all have
laticifers
Also has properties
of Ranunculales
Papaveraceae (2)
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Herbs or shrubs
Laticifers present (opium), alkaloids
Lvs simple but lobed or dissected, pinnate
venation
Flowers bisex, radial or bilat, Sepals 2,
deciduous, petals 4 or 6 (often crumpled),
sometimes united distally, stamens many to
2*3, carpels 2, united, superior ovary,
placentation parietal, fruit a capsule
Dendromecon
Dicentra
Corydalis
Lump or split?
Argemone
Ranunculaceae
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A family of exceptions
Mostly herbaceous, some shrubs, vines
Lvs usually alt or spiral, usually simple, oft lobed or
dissected, usually pinnate venation, usually no
stipules
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Flowers usually bisex, usually radial, usually not 3merous, tepals distinct, calyx usually distinct;
stamens usually many, carpels usually 5+, US
distinct, ovary superior
Fruit follicles (dehiscent, >1 seed) or achenes
(indehiscent, 1 seed)
Many stamens
No hypanthium
No stipules
Ranunculaceae
Berberidaceae
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Herbs or shrubs (ours), yellow wood
(berberine)
Lv usually alt to spiral, simple to compound,
entire to serrate or spinose-serrate
Flrs bisex, rad, usually 3 merous, calyx
distinct, inner petals (staminodes?) nectarproducing, stamens 4-many,anthers opening
by flaps, carpel 1, ovary superior; fruit usually
berry
Platanaceae
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Trees, monoecious
Branched hairs
Lvs alt or spir, usually palmate, gland
tooth, petiole covers axil bud, stipules
Flr unisex, reduced. Sep 3-7,pet 3-7, st
3-7, carp 5-9, distinct, ov sup. Fruit
achene, with bristles
web1.p15132387.pureserver.info/.../ Platane/
www.hope.edu/.../BasalEudicots/ Platanaceae
Platanaceae
www.hope.edu/.../BasalEudicots/ Platanaceae