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Biology 201 Dr. Edwin DeMont Digestion St. Francis Xavier University Feeding Animals exhibit a wide variety of feeding mechanisms and styles. Continuous feeders include aquatic suspension feeders (filter feeders); they are typically sedentary or sessile Discontinuous feeders have more specializations, as they “capture” their food, and attempt to avoid predation at the same time. They are often more active than suspension feeders. They gather and store the food, digesting it gradually. St. Francis Xavier University Digestion Digestive processes involve the uptake of food particles by a variety of processes and its subsequent enzymatic breakdown. Two different types of digestion exist: (1) Intracellular (2) Extracellular. St. Francis Xavier University Digestion Protists and sponges employ intracellular digestion—they do not have a gut. The possession of a gut allows extracellular digestion of food within the lumen of the gut cavity. St. Francis Xavier University Invertebrates Invertebrate digestive tracts range from gastrovascular cavities to complete, complex digestive systems. An incomplete digestive tract limits feeding, as the animal can only feed anew after undigested wastes have been expelled. St. Francis Xavier University Invertebrates When a complete gut is present there is a one-way flow of food through the animal. This makes possible a gut with distinct processing regions. And it works well St. Francis Xavier University Invertebrates Insects have a complete digestive tract with both storage and grinding regions. They use extracellular digestion. Digestion begins in the mouth and is usually completed in the gastric cecae. Most nutrient absorption occurs in the intestine and undigested foods pass to the rectum. Water and ions are absorbed from the fecal mass, and solid pellets pass out via the anus. St. Francis Xavier University Mammals The mammalian digestive system may be taken as an example of a typical (nonspecialized) vertebrate digestive system. Video St. Francis Xavier University Mammals The oral cavity consists of lips (sensory), teeth (grinding), an epiglottis (to divide food for the air headed to the lungs), and secretions by the salivary glands (to moisten food and add enzymes – eg. amylase.) Food (called a bolus) is then passed via the esophagus to the stomach. St. Francis Xavier University Mammals The stomach of a mammal stores food, mixes the food bolus, secretes enzymes, mucus, and hydrochloric acid, and controls the passage of the mixed, liquified food (chime) into the small intestine. St. Francis Xavier University Mammals There are 3 kinds of cells in the stomach: 1. parietal cells secrete hydrochloric acid; 2. chief cells secrete pepsinogen (pepsin); 3. numerous mucus cells secrete mucus to protect the gut from HCl. St. Francis Xavier University Mammals The small intestine is the primary site of digestion, as well as absorption; it has a lot of surface area. The length of the small intestine is related to the diet of the animal; herbivores have longer intestines. St. Francis Xavier University Mammals Villi (macroscopic) and microvilli (microscopic) both increase the absorptive surface area of the small intestine. Each villus has a capillary system and a lacteal to absorb nutrients. St. Francis Xavier University Mammals The pancreas secretes a number of enzymes into the small intestine facilitating digestion of nutrients and neutralizing acids. St. Francis Xavier University Mammals The liver functions in a variety of digestive ways, including breakdown of proteins, production of the plasma proteins, metabolism of carbohydrates and lipids and synthesis of some vitamins and bile salts. Further, the liver stores glycogen, which can be converted into glucose St. Francis Xavier University Mammals The large intestine primarily functions in absorption of water and minerals. Nutrients diffuse into the blood from the large intestine. St. Francis Xavier University Mammals The digestive tract is composed of an outer serosa, smooth muscle layers (typically one longitudinal and one circular), and the submucosa and the inner mucosa (connective tissues with blood and lymph vessels). St. Francis Xavier University Mammals Peristalsis (movement of food through the digestive cavity) and segmentation (churning) result from muscular movements within the gastrointestinal tube. Sphincters also regulate food movement, and close off certain gut regions. Control of gastrointestinal activity is regulated by the volume and composition of food in the lumen of the gut. St. Francis Xavier University