Download (Capparaceae). Malaysian species Malaysian species given. In

The genus Stixis
A
(Capparaceae).
M.
census
Jacobs
Rijksherbarium, Leyden
INTRODUCTION
For
long time, the
a
and
The
For
which is
genus,
Malaysia,
western
very
its
being
centre
a
in
of the
from
the
synonymy
name
have in the
species
species and
7
eastern
1
subspecies.
Himalayas
Hainan
to
Indo-China.
in the Flora Malesiana I,
and of the Malaysian species I refer
genus
revision; here only their
comprising
genus
extends
uniform,
Malaysian species have been treated
3
description
a
Stixis
been reduced, leaving
paper
The
genus
until Pierre monographed it in 1887. Several of Pierre’s
Roydsia,
present
Stixis has been known in the Indian Floras under the
and distribution
are
In the
given.
(i960)
5
95 —99.
the Flora Malesiana
to
key all
taxa
have
been included.
Thanks
at
my
are
due
the Directors of the following Herbaria for kindly putting material
to
disposal: Arnold Arboretum, Cambridge,
(BM), Bogor ("BO),
Herbarium,
Paris
Cambridge,
(P), Tokyo
U.S.A.
(CAL), Edinburgh
Calcutta
(GH),
U.S.A.
Leiden
(K),
Kcw
(A),
British Museum, London
Florence
(E),
(FI),
(TI), Berkeley (UC), Washington (US),
Vienna
(G), Gray
Geneva
Ann
(L),
Arbor
(MICH),
(W).
STIXIS
Lour., Fl.
Coch.
(1790)
5
(i960)
95,
1
f. 28,29 (map).
(1862)
110;
Type species:
Distribution:
S.
—
Stixis suaveolens
Seven
in
(Roxb.)
in India
species
China (Yunnan), Indo-China,
Gynophore (as
measured
2.
Sepals
reflexed
2.
Sepals
more
or
in
less
from the
anthesis.
in
3
Soc.
Linn. Paris
Jacobs,
201;
(1819) 87; Benth.
Fl. Br. Ind.
(1848)
1
Gynophore (as
3.
Adult leaves
4.
mostly
long,
Adult leaves
5.
from the
THE
torus) longer
shorter
all
mm.
than
or
in
axillary
12
at
cm.
the
reflexed.
shorter than 5
racemes
glabrous, occasionally
Style
none or
very
S.
Sepals
mm.
philippinensis
short, stigmas
reflexed in
S.
Gynophore glabrous. Ovary
longer
a
3,
distinct.
suaveolens
anthesis.
racemes,
2—3(—4)
rarely
in
a
terminal
than
few
12
cm.
hairs
glabrous, style longer
panicle,
3a.
mm
otherwise hirsute. Flowers in terminal
with
Siam,
3. S. ovata
axillary
Bracts
base,
N.
Ovary glabrous.
I.
not
Flowers in
—
SPECIES
than 6
Ovary hairy
over.
/,
409.
Malaysia.
western
undivided
torus)
underneath.
Gynophore glabrous only
cm
3.
hairy
Gynophore glabrous
cence
4.
measured
Hook,
&
307.
2.
1.
I,
Pierre.
TO
anthesis,
1
Fl. Mai.
(1872) 180, (1874)
(Sikkim, Assam, Chittagong), Burma,
Hainan, and
Style
spreading
1
17b (1936)
2,
Pi. Corom.
Hook./,
Arch.
KEY
1.
(1793) 361; Pierre, Bull.
Roydsia Roxb.
Hook./&Th.
Ned. Kruidk.
Covilhamia Korth.,
Willd.
in E.&P., Pfl. Fam. ed.
(1887) 652; Pax&Hoffm.
Gen. Pi.
ed.
295;
Bracts
on
the
than
(3—)4
mm.
panicles
3b.
ssp.
nerves.
the
ovary,
or
as
long.
the infloresssp.
about
ovata
12
—35
fasciculata
BLUMEA
6
6.
Leaves
times
i| —2
c.
long
as
VOL.
as
wide.
No.
XII,
I,
Inflorescences
1963
terminal.
Style 1—i£
6.
Leaves
c.
times
2—3
as
long
wide.
as
Inflorescences
axillary.
mm.
S.
4.
Style i|—2J
scortechinii
mm.
S.
5.
Fig.
454);
1.
b.
Ovary
8.
Ovary hairy. Style simple
glabrous. Style
obtusifolia (Hook.ƒ.
Sci.
d.
&
2
(1854)
329;
[Type specimen:
ƒ,
Fl.
Paris
Br.
1
Ind.
5
i
and
(i960)
F.-Vill.,
(1874)
—
stigmas
3
in Stixis.
c.
S.
a.
S.
—
S. scandens
g.
2
(1923)
213;
.
App. (1880)
S.
e.
S.
ƒ.
.
6.
11;
floribunda
ssp.
hookeri Pierre
(Poilane 10381),
(Philip.)
n.
35
S.
Roydsia floribunda
—
(Planch,
Merr.
scandens
(McGregor
fasciculata (King)
(Schmid 2);
all
x
(1906)
ex
341
See
Notes.
23
Laguna
Jacobs I.e. f.
1.
a
that it indeed
on
Basilan;
(map).
Seems
year
seems
s.n.
at
13,
t.
Planch,
Hook./) Pierre,
(CAL),
to
belong
both sides.
9j —10J
Racemes
mm,
mm,
to
20,
puberulous,
2
very
collected
6f. B.
ex
19th
—3
c.
8
cm.
filaments
—
Hook.
—
Fig.
Cotabato, and
ia.
(Luzon:
Davao
Prov.).
local.
in
northeastern Burma
century,
agrees
that species. The leaves
nerves
axillary,
stamens
be
97;
Bull. Soc. Linn.
(BM, G, K, L, W)].
Mindanao: Surigao,
to
the end of the
distinctly acuminate,
gynophore 3J
torus)
Prov.;
Pottinger
March of
obovate,
29
Philip.
72;
Distribution: Continental Asia: NE. Burma (see Note 1); Malaysia: Philippines
Bataan and
f. S.
4.
Soc. Nat. Moscou
Vidal, Sinopsis Atlas (1883)
[Type specimen: also Cuming
obtusifolia
S.
Erdtman, Pollen Morph. (1952)
Roydsia philippinensis Turcz., Bull.
(BM, G, K, L, W)].
—
Hall.
(Korth.)
Lour.
Govt. Lab. Publ.
Philip.
Nov.
distinct.
philippinensis (Turcz.)
ovata
(King) Jacobs (Lörzing 8683);
97.
341
409.
pistils
hookeri
ovary.
7.
(Turcz.) Merr.,
En.
the
but
short,
very
(Harmand M1);
Pierre
Cuming
(1887) 655.
or
than
(Clarke 37761);
Pierre
S. scortechinii
Th.)
Suppl. (1906) 58;
1
Jacobs, Fl. Mai. I,
27,
(Roxb.)
philippinensis
S.
J.
none
Androgynophores, gynophores,
S. suaveolens
shorter
stigmas
or
8.
Jacobs (Spire 1188);
I.
Style
Gynophore hairy.
5.
so
are
at
Bansparao
well with S.
c.
12—16
on
philippinensis
by 3!—6J
cm,
pairs, surfaces glabrous but full of pustules
Sepals reflexed,
c.
ovary 2 mm
10
in
mm;
9 —10
by ij—2
mm;
andro-
gynophore (as measured from the
diameter, glabrous, style
mm,
curved,
M.
pale, glabrous, distinctly 3-lobed.
Stixis
The genus
JACOBS :
(Capparaceae)
7
other collection thusfar is known
No
from outside
the Philippines.
(Roxb.) Pierre,
S. suaveolens
2.
I.-C.
(1908)
1
Fl. Gen. I.-C.
Gagn.,
277;
(1819)
87,
Hort. Suburb. Calc.
in
Hook./&Th.
(1886)
Suppl.
205;
4002)
1
[Type specimen:
c.
and less
or
f. 16,
1 —6.
(P)].
wide,
as
widest
(—121)
35—6\
cm;
pairs, thinly prominent,
in
panicle
(3 —)
sturdy
subglobose,
c.
white. Torus
c.
mm,
in the
ovate-acute,
606;
112
79.
—
I.-C.
Pierre
1
4022
on
sides.
wider
cm,
and after
anthesis,
glabrous, stigma
at
thin but
torus,
woody,
Asia:
splitting
India
North),
S.
China
into
(Sikkim;
(Upper and Lower Burma,
(two
a
10 mm,
base,
—6
4
(27—)
( —11)
pedicels
c.
by
2
once
2—3
mm,
40—43
mm,
3
very
few
c.
4
by
si
—
or
—10
a
3 —4
Buds
appressed
or
acute
top
to
subconical, passing
ovary
on
mm.
reported initially
mm,
( —50
7
downy-
to
2—4
more:
anthers elliptic,
Fruit
mm.
occasionally
tomentose
fulvously hairy,
f—1
above along
or
reflexed but spreading
not
the
ellipsoid,
orange,
branched
caducous;
mm,
—6
4
3,
triangular,
pustulate beneath,
pustules
some
and bracts
Androgynophore
styles
(io| —•)
less rounded
or
narrower
sometimes
sparsely
axes
Stamens
to
up
none,
or
glabres-
soon
above,
more
top
measured from the torus)
(as
—
subcoriaceous,
sometimes
to
sweet-scented, yellowish,
gynophore, glabrous.
the
3
c.
narrowed
both
654.
subdepressed above, prominent beneath,
wide, dish-shaped. Sepals
yellow. Gynophore
Distribution:
Burma
to
tomentose
measured from
endocarp
Gen.
(1887)
1
Leaves
cm.
rounded,
sometimes
Hooker/), glabrous, filaments
slightly during
\
(1934)
1
downy puberulous,
i£—2§
c.
\—1|
c.
long;
cm
diameter. Flowers
mm
into the
—100:
1
t.
Bot. Mag.
Assam
Linn. Paris
smooth except for
racemes
( —43)
13
elliptic-oblong,
densely
long, bright
by
mm
4
stamens,
gradually
axillary
bracts linear
puberulous,
pain;
Soc.
acute to
sometimes
Flowers
midrib.
obtuse,
Bull.
about the middle
base
abruptly acuminate, tip
more
7—9
the
Hook./,
(1887) 654; Gagn., Fl.
1
Petiole
cm.
the
to
Voigt,
Pi. As. (1854)
Ic.
409;
Corom. 3
202;
[Type specimen: Harmand hb.
—
Branchlets rather stout, terete,
internodes f—5
distinct; surfaces glabrous,
a
(1842)
Gen.
(1930)
6
lb.
Fig.
—
reticulation
in
I
Kanj. &Das, Fl.
blunt; midrib slightly sunken above, prominent,
acute to
nerves
15;
Soc. Linn. Paris
173,
4021
m.
long
as
( —30) by
13 —20
(1906) 36, f.
cochinchinensis Pierre,
var.
2—15
times
2—4
c.
Bull.
(1939)
Pierre
red-brownish;
cent,
(1854) 578;
4
(1872) 180, (1874)
1
J.
Roydsia suaveolens Roxb., Pi.
Pi. As.
Not.
Sci.
Lingn.
(1832) 643; Walp., Rep.
2
Br. Ind.
—
(1887) 654; Gagn., Fl.
1
35; Merr.,
(P)].
Stixis suaveolens
Climber
173.
(1921)
plate, I.e.].
flavescens Pierre,
Stixis
(1939)
Griff,
75;
Hook./, Fl.
Roxburgh's
(1908)
(not
(1845)
1
Carey
6881; Brandis, Ind. Trees
t.
[Type:
Suppl.
289; Fl. Ind., ed.
t.
Bull. Soc. Linn. Paris
Bot. Centralbl. 39
Hall./,
204;
mm,
—
J
8
mm
stretching
ellipsoid, i|
stipe
mm
sparsely
cm,
Gagne-
J
c.
—
long
1\
as
warted;
valves (Hooker /).
common
collections),
collections from
in Assam;
Siam
Yunnan),
(one
Manipur, Chittagong),
collection from Lampang
Hainan
China (all countries, several collections). Of all the species,
this
Indo-
(very common),
one
is
far the
by
com-
monest.
Ecology:
rocky soil,
Light forest,
once
reported
Uses: The fruit is
used
as
often
along
from the
reported
to
be
sea
rivers,
shore,
eaten
thickets,
to
1500
secondary
m
forest,
on
loamy
or
altitude.
in Assam and in Hainan. The leaves
may
be
tea.
Notes:
1.
and styles.
Occasionally
there
are
plants
with
the
pistil composed
of 4
or
5
carpels
8
BLUMEA
The specimens
2.
VOL.
and
Clemens 3664
No.
XII,
from
4113
1963
I,
Merrill
Annam, by
reckoned
to
scandens, actually belong here.
S.
3. S.
Hall,
(Korth.)
ovata
f., Beih.
Bot. Centralbl.
(1921)
ii
39,
35.
Covilhamia
ovata
Jacobs, Fl. Mai. I, 5 (i960) 97, f. 28, f. 29 (map: 2b).
3a. ssp.
—
—
(1921) 381.
Merr., En. Born.
fasciculata
S.
[Lectotype
name
140A.
(1906)
only).
Roydsia
—
37.
Gallatly
S.
—
499
Pax
201;
Pierre
Soc.
Bull.
stat. nov.
As.
Soc.
Pfl.
P.,
203; Pax
HofFm. I.e.
Gen.
Suppl.
I.-C.
Bull. Herb.
Hall,
(1939)
1
fasciculata
S.
I.e.
f.
f.
97,
212.
—
201.
—
(map:
29
(1896)
2
5,
Ind.
Brandis,
397;
(1908)
I
(i960)
5
Calc.
Gard.
(1889)
ii
Gén. I.-C.
121,
Trees
[Type specimen:
Fl. Gén.
(1887) 656; Gagn.,
i
ed.
17b
2,
(1904)
4
171.
Boiss. ii
f,
(1936)
202.
1072;
I.-C.
(1908)
1
[Type specimen:
—
4711
(1904)
4
Gagn., Fl.
Gén. I.-C.
(A;
P)].
type;
39
(1921)
Balansa 4073
specimen:
Fl.
Gagn.,
1071;
Beih. Bot. Centralbl.
[Type
—
(1936)
tristis Gagn., Fl.
var.
112
B-J.
Climbing shrub
soft stellate,
i
|—3(—15)
—14(—17) by
top
rounded and rather
surfaces
distinct;
—35(—40)
rather
cm
(1908)
1
Gen. I.-C.
1
Gagn., Fl.
35;
BO;
(A,
3
—6
reflexed, inserted
hairy
axillary
densely flowered, densely hairy
pedicels thin,
on
both
glabrous.
mm
on a
glabrous
diameter,
at
nearly
style
Fruit
with
G,
type;
3o( —40),
torus)
1
2
or
mm at
corky
mm,
c.
4
hirsute
by
warts.
3
cm,
soon
acutish
c.
to
glabrescent
blunt;
except
the main
(8 —)
panicle
3 —4 mm, narrow,
occur;
caducous;
long. Flowers fragrant. Sepals
mm,
lanceolate
Androgynophore
(i^
)i\
—
ovary
obovate,
c.
—
Gyno-
mm.
to
subglobose,
c.
35
i£ —2
the base, otherwise glabrous,
pericarp
above,
cuneate,
reticulation
especially along
branched leafy
filaments
at
to
sometimes
anthesis, later stretching
yellow-brown hirsute;
—1£
firmly herb-
Leaves
the middle
narrow,
by i\—2§
acutish.
glabrous,
2\
—
densely lenti-
sparsely branched panicles
mm
—6
with small
set
more
depressed above,
pustulate,
sparsely
4
top
densely
at
Hoffm.
&
Pax
ic.
obtuse,
above
hairs,
4 —5
wide,
mm
—
ellipsoid,
many
I
to
cm,
the twigs; bracts
as
3 —5-nerved,
otherwise
also hirsute;
spotted
)i6
—
from the
the base,
obscurely 3-lobed.
surface
torus
(20
Stamens
acute
hardly
or
racemes
long. Buds obovoid,
sides, obscurely
phore (as measured
mostly
terminal slender
a
2—9;
cm.
wide, widest
mostly sparsely
sometimes
18:
Fig.
i£—2^(—45)
as
not
pairs,
f.
—
glabrescent, and
late
brownish stellate
nerves,
mostly
long;
base
201,
(P)].
fairly straight,
terete,
long
as
cm;
6 —8
with light
Inflorescence
nerves.
1188
abruptly acuminate, tip f —I
nerves
the main
on
(1908)
I
Petiole terete,
times
—)3 —s|( —9)
midrib sunken above,
sometimes
Twigs
m.
simple, fulvous hairs,
sometimes
(1.5 —)2.3—2.6(—4.0)
aceous,
Gén. I.-C.
[Type specimen: Spire
—
cellate than in the other species.
12
(1953)
6
L, P)].
K,
11
Fl. Mai. I,
Bot.
[Type specimen: Balansa
—
(“longeracemosa”);
202
180;
type)].
longiracemosa A.DC,
S.
f1908)
Ann.
Beng. 58
Fam.
S. balansae A.DC., Bull. Herb. Boiss. ii
&
(1858)
L)].
G, K,
Jacobs,
—
King,
Linn. Paris
&
E
(G, K, L; P,
4023
BO,
2
1,
(L), Borneo].
s.n.
DD)].
type;
Hoffm. in
&
(A,
BM,
fasciculata (King) Gagn., Fl.
(CAL,
mollis Pierre,
S.
Jacobs,
King, J.
sp.
Bat.
Mitt. Bot. Staatss. Munchen Heft
Roydsia fasciculata
—
Ind.
Borneo.
fasciculata (King)
ssp.
2a,
t.
Malaysia:
Korthals
[Type specimen:
—
Clemens 26000
specimen:
Fl.
307; Miq.,
(1848)
1
borneensis Heine,
var.
Distribution:
3b.
Arch.
Ned. Kruidk.
Korth.,
ovata
2 —3
mm
thick,
spongy
mm,
mm
stigma
inside,
M.
Asia:
Distribution:
China
May,
Uses:
Forests, in hills
The leaves
scortechinii King,
120,
t.
139;
Mai. I,
ii
(1922)
1
Soc.
Petiole
i
j
up
—2
to
12
E.
&
Leaves
cm.
8J —12 by 3\ —5,)
cm;
subcoriaceous,
base rounded
sometimes brown and
blunt,
nerves
5—7
upper
side.
Inflorescence
Pfl.Fam. ed.
leathery
except for
1
—1|
tomentose
on
both sides.
filaments
glabrous,
2 —2j mm,
glabrous;
style persistent,
pale
at
green
January
3-lobed.
same
erroneous
(Hook. f.
S. obtusifolia
1
(1908)
36; Pax&Hoffm. in E.
f.
&
Th. in Hook,
(1877)
S.
harmandiana
205,
ed.
17b (1936)
2,
f. 18:
&
fl, Fl.
67; Brandis,
(1908)
201,
Ind.
by
1
—1|
1
I
—
10
mm,
mm
in
times
later
as
glabrous.
long
as
wide,
a
short,
glabrous (youngest
surfaces
the base of the midrib
long,
cm
or
more
stamens
J—4J by 2|—3j
3
a
subulate,
Torus
top
c.
obtuse,
(14 —)i6—22,
measured from
(as
the
densely
long.
mm
on
less
bracts
simple;
5
or
somewhat obovate,
sometimes with
a
[Type
ie.
the
diameter, glabrous, style (if
ellipsoid,
for
(1908)
1
—
narrowed with
glabrous;
mm,
—
Gynophore
Fruit
good
as
the
cm,
torus)
)2\
—
mm,
sometimes
few wart-like
lenticels;
when dried.
were
rare).
January 1867, and Thu-duc,
taken
probably
sheet in A,
locality
from the
labelled Pierre
491,
same
which is
for his number 1498.
Th.) Pierre, Bull.
&
in the
Br. Ind.
Trees
1
Linn. Paris
Soc.
key only; Hall, fl, Beih.
P., Pfl.Fam. ed.
2,
(1872)
(1906)
Suppl.
[Type specimen:
17b (1936)
180,
36.
Soc. Linn.
Fl. Gen. I.-C.
202.
m
1100
that number is the type of S. obtusifolia. In his publication,
Pierre, Bull.
12;
id.
Malaya
to
202.
unknown,
to
(Cambodia, Cochin-China;
holds
Pierre himself cited Thu-duc
Fl. Gen. I.-C.
near
sheets labelled Dong-nai River,
as
jungles,
Fig.
c. 2 —3
rounded
to
(glossy) brown
Indo-China
The
plant.
Fig.
—
midrib flattish above, prominent beneath,
few
pericarp leathery, thin,
opinion, the
considered to be
6.
top
number Pierre 1498,
1872, both under
individual
Kunstler
Fl. Gén. I.-C.
Buds ellipsoid-ovoid,
filiform.
maturity (ex coll.),
my
—6
subglobose,
ovary
Distribution: Asia:
Notes: In
5
nun,
—
396.
17b (1936)
—
the hairs in bundles
mm.
Androgynophore
s\
3
glabrous, stigma obscurely
the
US)].
obovate,
few pustules
raceme
3 —10
wide. Sepals reflexed,
mm
acutish,
acumen;
a
axillary
an
pedicels
caducous;
mm,
type;
2,
the youngest stage
ovate to
to
flowered, densely fulvous puberulous,
2—4
(1896)
5
Residency),
forest, landslides,
pairs, subprominent above, reticulation distinct;
unknown), smooth
stage
West Coast
(1887) 656; Gagn.,
1
P.,
Twigs fairly straight,
m.
Roydsia
—
Gard. Calc.
(1889)
ii
Beng. 58
(East and
specimen: Poilane 1498 (A, BM, G, K, L; P,
Climber
(map).
3
year.
Hoffm. in
&
Pax
29,
edible.
as
[Type specimen: King’s coll.
—
secondary
young
the
hookeri Pierre, Bull. Soc. Linn. Paris
5. S.
(" hookerii ");
f.
99,
recorded
are
Negri Sembilan, Penang).
places,
altitude. Fertile throughout
203
altitude. Most of the flowers
Ann. R. Bot.
397;
121.
Northern Sumatra
Selangor,
sunny
(i960)
5
(1889)
(1non Griff.) King, J. As.
Malaysia:
Dry
Pen.
m
G)].
type;
Perak,
Ecology:
Fl.
Beng. 58
Ridley, Fl. Mai.
Distribution:
(Wellesley,
Soc.
Indo-
collections),
fertile in other months.
of eye-sore. The fruits
case
Jacobs,
(King)
Roydsia parviflora
in
few
Tenasserim,
8oo( —1500?)
some
9
few collections from Laos and Cochinchina).
a
August, but also
applied
are
As.
J.
(BM; CAL,
8464
below
(Capparaceae)
Burma and.
(Upper
of the fruits in
most
scortechinii
S.
4.
Burma
Stixis
genus
in Tonkin and Annam;
(common
Ecology:
in
The
JACOBS:
—
Paris
I
1
(1939)
(1887) 654; Gagn.,
—
specimen:
ii
(1921)
Hook.
1
M’Clelland (K?, n.v.)].
(1887) 654; Gagn., Fl.
173;
Pierre 491
39
Roydsia obtusifolia
Fl. Br. Burma
409; Kurz, For.
(1874)
[Type
202.
1
Bot. Centralbl.
Pax&Hoffm. in E.
(K, L; P, type)].
—
Gen.
& P.,
I.-C.
1
Pfl.Fam.
Fig. if.
BLUMEA
10
VOL.
No.
XII,
1963
I,
Climbing shrub. Twigs fairly straight, mostly with
the petiole insertion,
(1 —)i|(—2J)
1.5 —1.8( —2.0) times
c.
—12^( —16) by 5^—7( —8)
10
base rounded
reticulation
abortive; midrib
distinct;
translucent pustules all
surfaces
in the axil of leaves
c.
12
by
8
flowers
by
to 20
the
over
tomentose,
ciliate,
reflexed,
nearly \
J—5
with
4
by i\
mm
mm,
minute
mm
f
mm,
(2 —)
3
cm,
prickles
Fruit
(Cambodia,
Ecology:
S.
Soc.
E.
Burma
P., Pfl.Fam.
Fl.
(1887)
1
ed.
37;
key only;
Malay
Stixis
1
Hall,
Cochinch.
f.,
densely
the base,
c.
older twigs,
long, bearing
yellowish
caducous,
soon
1 mm.
Buds globose,
Burma
3J
mm,
c.
both sides. Androgynophore
c.
observed in
thinly pubescent;
short
very
be
to
on
specimens
(in
Burma
mm
3 —5
18 —2i),
filaments
young
flowers).
subglobose,
ovary
specimens),
stigmas
c.
long, ellipsoid, (3 —)4f by
muricate with
long soft
—
Fig.
Twigs when
175
203; Pax
Suppl.
by
1
young
Ind.
655;
1
(1854) 578;
4
180,
For.
33;
(1922)
39
(1939)
171.
in E.
Pax
(A,
Ic. Pi. As.
(1854)
Fl.
32.
type; L, P,
&
1
1
607 f.
I.-C.
(1908)
1
in
P)].
Hook.
1;
descr.;
409,
(Griff.) Pierre,
Trees
Bull.
in the
200,
[Type specimen: Griffith
Burma
W)].
(1887) 655; Gagn.
P., Pfl.Fam. ed.
2,
in Lecomte,
17b
(1936)
[Type specimen: Harmand hb.
—
Hoffm.
(1877) 67; Brandis, Ind.
parviflora
Gén.
—
t.
&
type; K,
(1874)
Hook. /, I.e.
Stixis
—
(1912)
Bull. Soc. Linn. Paris
Hoffm.
206;
10381
Fl. Burma
120.
(1793) 361; Pierre, Bull.
(1908)
Poilane
1
name;
in Lecomte,
Gagn.
Willd.
[Neotype:
—
(1872)
(CAL, GH; K,
&
ed.
295;
Fl. Gen. I.-C.
202.
I
(1874)
densely
fulvous
few. Petiole
times
4—cm;
cm,
(1790)
Fl. Gen.
Gagn.,
202;
Pierre 4023
(K.;P,
ig.
(2.1 —)2-3 —2.9( —3.5)
i —!(—1)
(Central and eastern), Indo-China
Siam
June (Gagnepain).
Beih. Bot. Centralbl.
elongata Pierre,
(1908)
lenticels relatively
11 —13
or
Gagn.,
Not. Pi. As.
Mai. Pen.
(1887)
i
Peninsula
Fl. Gen. I.-C.,
type)].
tomentose
(in
woody stipe
17b (1936)
2,
Ridl., Fl.
Soc. Linn. Paris
I.-C.
none
on
cm
subulate, mostly
mm,
(especially
c.
—6
4
wide. Sepals sometimes coherent in twos,
26—4°
torus)
c.
the base; pedicels
at
mm
fruits in
656;
Kurz, J. As. Soc. Beng. 43 ii
and
between,
nerve
lower surface glossy; both
nerves,
towards
to
(Lower Burma),
Flowers in March,
scandens Lour.,
(1906)
I
top
thick
Th. in Hook, f, Fl. Br.
&
lesser
a
Cochin-china).
Roydsia parviflora Griff.,
f.
size;
which
with many small somewhat
rough, seed villose (?; Pierre), endocarp
pericarp
Linn. Paris
&
style
a
on
in
acumen
(Brandis).
Distribution: Asia:
7.
c.
stamens c.
from the
diameter, glabrous;
glabrous.
protrusions
below the
incrassate
above,
to
diminishing
leafless panicle lateral
a
laxly
narrowed, obtuse;
top
mm,
with
angular, mostly simple,
more
cup-shaped,
Gynophore (as measured
1
also
specimens)
the branches
long, glabrous,
sub-
Leaves
panicle, mostly terminal with often the basal branches
a
length but
2
often
glossy
the main
near
near
depressed above, prominent beneath,
less
or
when young; bracts up
diameter. Torus
mm
less
or
the margin,
especially
Burma
cm,
whole
especially
3
(in
or
10
tomentose,
2\
over,
glabrous. Inflorescence
incrassate.
below the middle
small thin-leathery brownish
a
more
protrusions
ferruginous pubescence,
or
lower part
wide, widest
as
more
near
surface
upper
the
cm,
minute
very
towards the inflorescence often
cm,
pairs connected
9—11
c.
2
ashen brown
an
long
as
rounded with
acute, top
to
sometimes remains
nerves
parts with
young
Petiole
glabrescent.
soon
coriaceous,
the
base
as
c.
long
acute
puberulous, the hairs simple
i£
cm.
as
to
Leaves
wide, widest
rounded,
blunt; midrib depressed above,
or
stellate, glabrescent;
firmly herbaceous
top
at
the middle
to
to
subcoriaceous,
mostly above,
rounded and acuminate,
prominent beneath,
nerves
the
5 —8
tip
pairs,
M.
subprominent
above,
glabrous with
a
but
a
connected
few
axillary,
wide. Sepals reflexed,
sides.
5
Androgynophore
by
—6
c.
—
ï
J
1
\
obovate,
mm,
a
thick
stalk
woody
rough with
pericarp
Notes:
of
S.
The
i.
scandens
flattish
was
apply
only
ovata ssp.
the leaves from his
Clemens
specimens
2
species
are
the sepals being
and
3774
this
ovary,
mm,
mm,
description
which could
that it
into
come
observed
stamens
that
unlikely
seems
would omit mentioning
These clearly
4113.
characters Loureiro's
and
rotunda",
3,
one
but there the smallest number of
"petala
not
not
and
more
the pubescence
than
revoluta",
the
mentions
174,
not
stamens
that
but
suaveolens,
with,
agree
viz.
in number
40—50
"germen pilosum".
26
a
(1935)
S.
to
free
styles, and
terminal inflorescence and
a
"stylus brevis,
not
simplicibus,
"racemis
not
stamens.
species which is definitely
this is
obtusifolia, but
belong
description does
Loureiro would have observed 3
axillaribus",
suaveolens and
mm.
Upper Burma),
given in Loureiro's
Loureiro's Flora Cochinchinensis
on
and
It is remarkable that Loureiro had
S.
2J —5
persistent style if
Burma (Northern and
Another
species.
the
on
glabrous and
ovary
stigmata
both
tomentose on
description.
appressed
obtusifolia,
crassius,
longis,
the
16,
not
In S.
mm
1
the type of S. elongata)
on
with
cm
distinctly hairy underneath,
so
couple of
a
possesses
by if
3),
Note
see
fasciculata,
in his Commentary
Merrill,
c.
warts.
to
Loureiro, who noticed the hairs
on
mm,
Torus
filaments
16
—24,
nerves.
c.
densely fulvous-hairy, style |
combination of characters
complete
and the leaves
20,
obtuse, densely
mm,
c. 2
above
Annam).
can
consideration is S.
surface
the main
on
long.
mm
4—5
stamens
1\
long, ellipsoid,
6 mm
Distribution: Asia: India (Assam;
Indo-China (Laos,
few pustules
a
distinct. Fruits (mature? described
to
many
top
glabrous;
mm,
—
on
with
long. Buds ovoid,
mm
Gynophore (as measured from the torus) f
glabrous, stigmas obscure
reticulation distinct;
of the midrib, underneath glabrous
part
nerves,
11
densely fulvous-puberulous, bracts subulate,
3 —6
c.
(Capparcaeae)
the margin,
near
the main
on
5 ——9) cm,
early caducous, pedicels
Stixis
along the basal
pustules
few hairs sometimes
Raceme
The genus
JACOBS:
hardly
an
argument
rare
and
not
the
stand against
to
common
careful
a
interpretation.
2.
A
sterile
The leaves
specimen collected
if
are
cm
petiolate,
towards the base and gradually acuminate
with
glabrous, above
and with
3.
A
a
glaucous
some
by J.
in Laos
lanceolate,
pustules
Vidal
obovate,
at
22
—
the top, with
by
3 —jf
cm,
narrowed
pairs of strong
8— 12
nerves,
the base of the midrib which is subprominent,
near
tinge underneath.
from the Naga hills in Assam, Meebold 7333,
specimen
here.
(P) probably belongs
s.n.
16
here, although the gynophore
is
as
long
as
most
probably
belongs
mm.
3
Index
Synonyms
the
are
in
italics; accepted
names
in normal
type;
new
names
in bold
type.
refer
The numbers
to
species.
Covilhamia
ovata
elongata
fasciculata King
obtusifolia
parviflora
3
b
Hook
f.
Hook,
Griff.
philippinensis
A.DC.
balansae
3a
Roydsia
floribunda
mollis Pierre
>
Stixis
Korth.
f
fasciculata
1
var.
&
Turcz.
King
suaveolens
Roxb.
Th.
6
var.
4
2
1
7
ovata
(King)
Gagn.
borneensis
tristis
flavescens
7
scortechinii
Pierre
obtusifolia
3b
Heine
Pierre
Pierre
longiracemosa
ssp.
Hall.
ovata
&
(King) Jacobs
Pierre
philippinensis (Turcz.)
Pierre
6
scandens
Lour.
scortechinii
suaveolens
5
A.DC.
1
3b
var.
Pierre 6
Th.)
/.
3a
parviflora (Griff.)
2
floribunda (Hook. /.)
hookeri
(Korth.)
ssp. fasciculata
3a
Gagn. 3b
Pierre
harmandiana
3b
3b
(Hook./.
7
Merr.
7
(King) Jacobs
(Roxb.)
cochinchinensis
4
Pierre
Pierre
2
2
1
3b
BLUMEA
12
Index
The
number
marked
is
specimen
d'AUeizette
Edano:
BNB For.
2.
mens
C/imh
446:
Eberhardt
FB
R.
2263
Gage
Meyer:
254: 2; 257: 2; 577:
the
A
text.
type
3807:
3b;
4071: 2; 4072:
Bon
3a.
3b (T);
4073:
4800: 3b;
5419:
3b (T); 47*9:
4074: 2; 4711:
3b;
BS
6179: 3b.
2.
-Ri»mw&
49079
P
3b;
13S3:
26605:
(T); 26647:
P
27060:
2;
1715:
26887:
3a;
3a;
Chevalier
3b; Fleuryhb.
37761:
2; 37753: 2;
305*0:
Cuming
3a.
32308: 3b;
431S0:
2;
541:1
2;
32354:
2.
43340:
Cle-
(T). Curtis3284: 4.
2.
FB
1;
F««sj
1.
3b
499:
FB
1;
20151:
(T). Galoengi
E.
Griffith
2.
Borden:
2326
6101
Hutchinson:
FB
1;
24199 Bawan:
1;
FB
26266
2.
517:
Bengal 175A:
Gamble
4.
2; 427: 2;
578:
2.
1044:
Griffith
2.
Gaudichaud
voy.
Burma and
Malay
Bonite
252:
Peninsula
2;
175:
(T).
1
Hallier I.
7
in
taxon
Buchanan 27: 7.
I.
3a
Mabesa:
Gallatly
2.
included
of the
4.
3b;
C/arfef
2.
26000:
FB
77:
ones
the number
2.
4743:
Mallonga: 1;
1963
I,
Malaysian
with
corresponds
3019 PokUn:
& Edaiio:
Dcn(
3a.
789:
1534:
De/>f
6444:
3774: 2; 4113: 2;
Darnton
the
specimens,
to,
Poilane hb. Chevalier
Chalterjee 86735: 2.
3b; 5259*: 3b.
referred
Alvins
49617 Ramos
1;
is
Balansa
148: 2.
255:
examined
No.
XII,
(T).
3b.
371:
BadulKhan
Beddome
to
specimen
a
VOL.
B
Jenkins,
2;
62410:
Lace
lilfer
11247:
4;
Maradjo
Parker
46*2:
2;
1498:
4024:
1521«:
3240:
2;
2.
2.
2.
Po
Rachmat
Talmy
750:
j*.
Kirfa/
Wallich
10413:
Yates
2;
4.
6. Harmand
J24:
72177: 2;
6; 4025:
745bis:
72594:
2.
2.
1CF.P
2.
53949 Sou':
8464:
4. /Cctt
King's
7004: 6.
(T). Kingdon-Ward 224,
4
1512:
coll.
3b.
295:
JCo
2;
52205:
60:
2;
J401: 2;
1664:
(with Jochems):
755*
2.
4;
Liang
7962:
62065:
4;
64722:
2;
«6«3:
4;
9696:
65394:
2;
4;
10052:
2.
4;
2;
2.
PNH
3b;
#060
7058:
DR.
2.
219:
2;
5;
Meebold
1.
Ta/my
1.
3b
(T);
3b; 10381:
7
/ifc.
comm.
(T);
7533:
Petelot
Pierre
354:
Anderson
11052:
coll
840:
2;
930: 2;
7.
966: 3b;
hb.
3b;
977:
2.
6.
Mouret
82:
itffip:
3b;
Pierre 491:
Pierre
11244:
24843: 3b; 24737A: 3b; 25830:
Prain's
2.
454:
Pasquier 3020: 3b.
4023:
10197:
23596:
195:
McGregor
rfM
Quisiimbing:
(T);
2
4022:
2; 20503: 2;
2;
2.
2930:
Parrey
(T);
4;
12:
7349:
4;
7688:
2039: 6; 2099: 6;
3b;
30519:
2452:
2;
12403:
2;
3b.
2204:
6
4023: 2;
3b;
(T);
ditto
3b;
13075:
25875: 2;
Purkayastha
26025:
Put
22: 2.
4
5560:
(T). S/jire
2.
Ridley
7355: 4;
913: 2; n#S:
9306: 3b. Toppin
3105:
7.
15634: 4.
3b (T).
Tiaii?
75:
Rhjc 417;
2.
S» Kbe 9093: 7.
2;
15576:
2;
16824:
2;
Watt
7336:
2;
3b.
2622:
2.
Rao
4.
5. Scortechiniis.n.:
Wang 32780:
2;
2.
6.
3b;
Thorel
11825:
172:
7321:
27347:
1232:
123: 6;
70761: 2;
1*02: 6.
Law
2944: 2;
3b.
2
J353:
20387:
11:
6.
4200:
1786:
5;
2543:
Si Boeea
I77«o: 2;
c.5
6534:
<>#2<>: 4;
2562:
4021:
Khant
Schmidfieldno.
Harmand
exp.
How 70420: 2;
2.
362:
2;
Kwjr
6.
1307:
7*51:
3b;
2815: 2;
(prob.);
6
Godefroy
123:
2;
(T).
3a
Maung Mya
Poilane 237:
31429:
253S:
70:
5775:
4222: 4;
Parkinson
15740: 2; 20297: 2;
2;
2;
2;
4.
4.
4*24:
(T);
5
s.n.:
Ldrzing
349:
122:
coll. Kunstler 1611: 4;
4225: 4;
Lakshnakara
2.
2.
28:
1730: 2;
Korthals
2.
2177:
223/I:
2;
King's
2.
430:
Hookerf.
2.
Anderson
comm.
Kanjilal 148P:
371: 2;
Mi: 6.
2579: 3a. Harmand
(T). Henry 13819:
2;
2.
Kitfa/
75605:
Williams
So/er
y
2;
964:
79158:
2718:
1.
2.
Winit
1.
Warburg 13483 (not 13183):
1923:
2.
1.