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SouthAmerica-IX' Studieson thefloraofnorthern H. A. MISCELLANEOUS NEW GLEASON SPECIES FROM BRITISH GUIANA Tetrapodenian. gen. Malpighiaceae; arborescent,the young branches stronglyflattened; stipules axillary, connate, erect; leaves opposite,petiolate,coriaceous,entire; flowersin minutely bracted, terminal racemes, on jointed pedicels; bracteoles 2, the lower glandular; calyx io-glandular; petals 5, clawed, fouressentiallyalike, the fifthmuch smaller,its blade bearing two pairs of latero-basal,stipitate,reniformglands; stamens IO, the fiveopposite the sepals the longer;filamentsshort,glabrous, and barelyconnate at base; anthers somewhattriangular-dilated or triangular-prismatic semicylindric simple, the stout, elongate, connectivestruncate to broadly rounded at the apex; pollensacs 4, elongate,strictlyparallel in the longerstamens,slightly divergentdistally in two pairs in the shorter;ovary depressedhemispheric,3-celled,glabrous; styles3, glabrous,bent near the tip at rightanglesand graduallytaperingdistallyto theobliquely truncateterminalstigma;fruitindehiscent,I-celled and I-seeded by abortion,subellipsoid,theexocarpthickand woody,the endocarp chartaceous; seed ellipsoid, the small cotyledon partly enclosed by the large. to The name is combinedfrom:rspa, 70'q, and ac6'v,referring the fourstalked glands on the fifthpetal. Tetrapodenia glandifera n. sp. Tree 4-6 m. high; young branches stronglycompressed,glabrous, later becoming terete and roughenedby elongatelenticels;stipulestriangular-subulate, erect, stiff,6-8 mm. long, densely pubescent above; petioles IO-I5 mm. long, glabrous,flattenedor barely channeled above; leaf-blades stiff,coriaceous, dull green, elliptic-oblong,IO-I7 cm. long, 4.5-8.5 cm. wide, abruptly acute or subacuminate, entire,broadly cuneate to obtuse or rounded at base, glabrous above, glabrous beneath (except under strong lens minutely lepidote) and biglandular at the base; veins and veinlets prominulouson both sides, conspicuouslyreticulate;racemessingle or in threes,terminatingthe branches,the axis somewhat flattened and minutelypuberulent,the bracts broadly ovate-triangular,I-2 mm. long; pedicels7-8 mm. long,glabrous,jointed in the middle,bearingop the lowersegmenttwo broadlyovate, amplexicaulbracteolesI.5 mm. long, the lowerof whichbears a conspicuousgland; calyx-glandsIO, cuneate-obovate,adnate to Contributionsfromthe New York Botanical Garden,no. 282. 289 290 BULLETIN OF THE TORREY CLUB [VOL. 53 the summit of the pedicel; sepals triangular,acute or barely retuse, exceeding the glands by I.3 mm.; petals dimorphic, clawed, the four larger with semiorbicular,subentire blade, cleftat base to the insertionof the claw, the fifthmuch smaller, obovate-oblong,with stout fleshyclaw, the basal pair of glands o.6 mm. in diameter,on stipes o.s mm. long, the distal pair half as large,on stipesone thirdas long; filamentsflat,glabrous,3 or 2 mm. long, dilated and barely connate at base; anthers stout 2-2.5 mm. long in bud, the simple connective somewhat surpassing the narrowlylinearpollen-sacs;fruitsmooth,20-25 mm. long by I2-IS mm. in diameter,rounded at base, subacuminate to the blunt apex whichis markedby 6 faintradial furrows. Type, La Cruz 3515, collected at Amakura, NorthwestDistrict, British Guiana, 23-30 March I923, and deposited in the herbariumof the New York Botanical Garden. La Cruz 2176, fromthe upper Mazaruni River,collectedthe precedingautumn, agrees with the type in all vegetative characters,except some small differences in texturedue to age, and exhibitsmaturefruit and old flowersfrom which the petals have fallen. La Cruz 2869, withmaturefruit,and 2013, sterile,bothfromtheMazaruni region,differfromthe firsttwo in theirlarger,proportionately narrower,and more sharply acute leaves which are distinctly shining above. The fruitis preciselythe same. The specific descriptionis drawn entirelyfrom the type, except that the lengthof the filamentsis measuredfromthe witheredflowersof 2i76. Duplicates of all four numbers are to be found in the leading Americanherbaria. Tetrapodeniais obviously closely related to Glandonia and Burdachia,two generaof the northernAmazonian forestswhich are still poorly known and have probably not been collected in recent years. The formeris monotypic,but both flowersand fruitare known and were carefullydescribed by Grisebach in the Flora Brasiliensis. The lattercontains two species, but the flowers of only one are known. Tetrapodenia differsfrom Glandoniain its hirsutestipules,the glands on the lowersurface of the leaf, the differentiated fifthpetal, the unappendaged anther-sacs,the shape of the fruit,and the presenceof two cotyledons. From Burdachia prismatocarpa,it is distinguishedby the glabrous,depressed-hemispheric ovary,the simpleblunt connectives,the characterof the fifthpetal, and the shape of the fruit. From Burdachia sphaerocarpa,the flowersof which are not known,it differsin its glabrousstipulesand the shape of the fruit. 19261 GLEASON: NEW SPECIES FROM BRITISH GUIANA 29I Te/rapodenia glandiferais representedin the herbariumof the Royal Botanic Garden at Kew by two collectionsby Jenman, 3779 and 6413. Combretum brunnescens n. sp. Shrubby vine, climbing 6 m.; stems glabrous, stronglyfurrowedand flattenedabove, becomingtereteand barely striatein age, the upper internodes 5 cm. long; petiolesstout, dark brown,6-IO mm. long, angular, glabrous;leaf-bladescoriaceous,olive-brownwhendried,broadly elliptic-obovate,70-85 mm. long, 43-58 mm. wide, obtuse to subrotundat base, entire,broadly roundedabove and minutely apiculate, shiningabove, glabrouson both sides; lateral veins 5 or 6 pairs,lightlyimpressedabove, prominentbeneath,ascending at an angle of 600 and nearly straight,the veinlets obscure; panicles freelybranched, terminal and from the upper axils, IO cm. long; spikes spreading,5-6 cm. long, the rachis strongly angled,glabrous; bracts subulate,puberulent,0.5-I.I mm. long; ovarysessile,linear-oblong, I.I-I.3 mm.long,0.4 mm. in dr- ameter,obtusely 4-angled, glabrous or minutelyvillous; calyx campanulate, glabrous,conspicuouslybut minutelyred-dotted, 1.7 mm. long, including the broadly triangular,acute teeth mm. long, glabrous within; petals white, transversely 0.4-0.5 elliptic,1.4 mm. wide,o.8 mm. long,glabrous,with a shortclaw 0.1 mm. long; stamens 8, insertednear the bottomof the calyx, the filamentswhite,glabrous, subulate, 4 mm. long, somewhat incurvedat three-fourths of theirlength,the anthersobcordate, 0.4 mm. long and wide; disk a minuteglabrousringat the bottom of the calyx; style straight,white,subulate, 2.8 mm. long. Type, La Cruz 3566, collected along the Amakura River, NorthwestDistrict, BritishGuiana, lat. about 8? IO' N., long. about 600 XV.,23-30 March 1923, and deposited in the herbarium of the New York Botanical Garden. The type of Combretum brunnescenshas been chosen arbitrarilyfromseveralspecimensin the herbariumof the New York Botanical Garden, all collected in the same general region. Duplicates of most of them have been distributedto the larger Americanherbaria. These are La Cruz iog4 and iiji fromthe Pomeroon District, 3425, 3585, and 3713 from the Northwest District, 1947 fromnear Bartica, 4258 fromKamakusa, upper Mazaruni River, and 4460 fromKaieteur Falls, Potaro River. All but the last two of these are in flowerand differfromthe type onlv in minordetails. In two fromthe PomeroonDistrict the largest leaves are only 6 cm. long; in all except I947 and 3425 the leaves are moreshiningabove; in 1141, 1947, 3585, and 3713 the terminalapiculum of the leaf is more developed and