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How Does a Population in “Hardy-Weinberg equilibrium” change ? (687)....or.......How does a population evolve ? Rattlesnake example...page 687 Mechanisms/Processes that drive change include...... 1. Mutations : 2. - might create NEW alleles for a trait - might provide an advantage to organisms carrying it, so they reproduce more and their alleles become more common in the population Ex. Bacterial resistance to antibiotics - provide more diversity to face environmental changes, what wasn’t an advantage before may become an advantage later Ex. Water fleas (“Daphnia”) mutated allele enables survival when water warms to 25 or 30 degrees, others die Genetic Drift : happens in really small populations....how common each allele is (it’s “frequency”) changes a lot by chance alone... 2 scenarios making this happen.... Bottleneck effect its less likely all of the alleles in the parent population will show up in the next generation. ( not all reproduce ). Some alleles become common quickly, while others are lost (drift)...less variation in the next generation Ex. Northern elephant seals hunted down to 20 animals in the 1800s, now bounced back to 30 000....but studies of 24 loci show NO variation. Whooping cranes in NWT : 12 to 177 Endangered species with really small populations Founder effect A small number of organisms colonize a new environment. Their gene pool doesn’t contain all the alleles of the parent population. (Drift) 3. Gene Flow : new alleles are brought into a gene pool, or some genes leave if there’s a significant amount of gene flow between two populations, the gene pool blends into one over time, and combined with selection, some diversity is lost Ex. Our own population ? 4. Non-Random Mating : inbreeding creates more homozygous organisms 5. - ex. self pollination in plants, or isolated populations - assortative mating, selecting a mate that has a similar phenotype as you, less diversity over repeated breedings ex. Dog breeding....may lead to increase in frequency of conditions like hip dysplasia Natural Selection : - 3 types........ Stabilizing selection - the environment favors the “in the middle” phenotypes, and selects out the extremes Ex. Human birth weight Directional selection - one extreme phenotype is selected over all others Ex. Small forest dwelling prehistoric horse species producing large open grassland horse species Fat on penguins Disruptive selection - 6. the extreme phenotypes at both ends of a population are favored, the “in the middle” phenotypes are removed Sexual Selection : smaller motile sperm (increased numbers) and enlarged nonmotile egg , creating competition between males that go looking for eggs. Leads to lots of diverse ways of attracting mates and species sexual behavior. Ex. Different bird plumage between genders (Sexual dimorphism) Courtship displays / rituals How Species Form...”Speciation” (page 708) A species is a group of related creatures capable of breeding to produce fertile offspring. New species may form by way of two main pathways....both due to natural selection working on a pop’n.... Transformation the build up of small changes over time Divergence A new species forms from a parent species that continues to exist. To remain a distinct species, a pop’n must have barriers to prevent interbreeding, like... Geographical barriers - a topographic feature keeps pop’ns separate....like a river or canyon ( allopatric speciation ) Biological barriers - Keep reproduction separate, even when living in the same area, like... Pre-Zygotic Barriers (prevent mating or egg fertilization) Behavioral isolation- songs, dances, pheromones are species specific Habitat isolation- even though they live together, habitat preferences mean different species encounter each other less often Temporal isolation- timing barriers ( day, season, year of mating varies) Mechanical isolation- your gear doesn’t mesh Gametic isolation- sperm / egg incompatible Post-zygotic Barriers - (Prevents hybrid zygotes from reproducing......) Hybrid inviability, sterility, or breakdown Processes Giving Rise to Species : Sympatric speciation - reproductive isolation. Ex. Plants with unique chromosomal structure, animals having non-random mating Polyploidy in offspring - will only successfully mate with another polyploid, new species that can’t breed with parent organism Allopatric speciation - population split into groups by geography Adaptive radiation, or divergent evolution a species colonizes new areas different enough for N.S. to form a variety of species from a common ancestor. Ex. Galapagos finches Convergent evolution Two unrelated species eventually turn out similarly in similar environments. Ex. Bird and bat flight Coevolution Two species interact to control the evolution of the other, like plants and pollinators, predators and prey, parasites and hosts. The speed of evolution : 2 theories : gradualism and punctuated equilibrium Gradualism slow accumulated changes, not supported as much in the fossil record. Punctuated equilibrium (Gould - Eldridge hypothesis) Sudden bursts of speciation ( < 50 000 yrs) interrupt long periods of time where species seem relatively unchanged. Theories on the Origins of Life : Chemical Evolution - pros/cons, page 730 ( the Haldane - Oparin hypothesis ) Inorganic chemicals could, under the right atmospheric conditions, become rearranged to make organic chemicals. Tested by Stanley Miller in 1953 methane, ammonia, hydrogen, and water inside a setup, sparks to mimic lightning...after a week, organic molecules present....including amino acids This is not life, but more tests show nucleotides and RNA form too. Self replicating molecules must have existed before any life forms. The Panspermia theory : life originated elsewhere, and traveled to Earth, introduced by meteors, or by other beings. The Gaia Theory : chemical evolution first, then the Earth living as a large organism controlling populations and maintaining homeostasis....James Lovelock...1960's Intelligent Design Theory : A supernatural being created all life, since life and all of its processes are too complex to be the result of random chance over long periods of time. ( Creationism ) Serial Endosymbiosis Theory ( SET ) : (The Lynn Margulis hypothesis) Eukaryotic cells included into their structure smaller free living bacteria like life forms, ( aerobic bacteria to mitochondria, photosynthetic bacteria to chloroplasts ), which allowed them to survive in more environments.