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.The Senses Special senses o Smell o Taste o Sight o Hearing o Equilibrium The Eye and Vision 70 percent of all sensory receptors are in the eyes Each eye has over 1 million nerve fibers Protection for the eye o Most of the eye is enclosed in a bony orbit o A cushion of fat surrounds most of the eye Accessory Structures of the Eye Eyelids and eyelashes Conjunctiva Lacrimal apparatus Extrinsic eye muscles Accessory Structures of the Eye Eyelids o Meet at the medial and lateral commissure (canthus) Eyelashes o Tarsal glands produce an oily secretion that lubricates the eye o Ciliary glands are located between the eyelashes Accessory Structures of the Eye Conjunctiva o Membrane that lines the eyelids o Connects to the outer surface of the eye o Secretes mucus to lubricate the eye and keep it moist Accessory Structures of the Eye Lacrimal apparatus lacrimal gland and ducts o Lacrimal gland—produces lacrimal fluid; situated on lateral aspect of each eye o Lacrimal canaliculi—drain lacrimal fluid from eyes medially o Lacrimal sac—provides passage of lacrimal fluid towards nasal cavity o Nasolacrimal duct—empties lacrimal fluid into the nasal cavity © 2015 Pearson Education, Inc. Accessory Structures of the Eye Function of the lacrimal apparatus o Protects, moistens, and lubricates the eye o Empties into the nasal cavity Lacrimal secretions (tears) contain: Dilute salt solution Mucus Antibodies Lysozyme (enzyme that destroys bacteria) Accessory Structures of the Eye Extrinsic eye muscles o Six muscles attach to the outer surface of the eye o Produce eye movements Structure of the Eye Layers forming the wall of the eyeball o Fibrous layer: outside layer o Vascular layer: middle layer o Sensory layer: inside layer Humors are fluids that fill the interior of the eyeball Structure of the Eye: The Fibrous Layer Sclera o White connective tissue layer o Seen anteriorly as the “white of the eye” Cornea o Transparent, central anterior portion o Allows for light to pass through o Repairs itself easily o The only human tissue that can be transplanted without fear of rejection Structure of the Eye: Vascular Layer Choroid is a blood-rich nutritive layer in the posterior of the eye o Pigment prevents light from scattering Modified anteriorly into two structures: 1. Ciliary body—smooth muscle attached to lens by ciliary zonule (suspensory ligament) 2. Iris—regulates amount of light entering eye Pigmented layer that gives eye color Pupil—rounded opening in the iris © 2015 Pearson Education, Inc. Structure of the Eye: Sensory Layer Retina contains two layers 1. Outer pigmented layer absorbs light and prevents it from scattering 2. Inner neural layer Contains receptor cells (photoreceptors) o Rods o Cones Structure of the Eye: Sensory Layer Signals pass from photoreceptors via a two-neuron chain o Bipolar neurons o Ganglion cells Signals leave the retina toward the brain through the optic nerve Optic disc (blind spot) is where the optic nerve leaves the eyeball o Cannot see images focused on the optic disc Structure of the Eye: Sensory Layer Neurons of the retina and vision o Rods Most are found toward the edges of the retina Allow vision in dim light and peripheral vision All perception is in gray tones Structure of the Eye: Sensory Layer Neurons of the retina and vision o Cones Allow for detailed color vision Densest in the center of the retina Fovea centralis–lateral to blind spot o Area of the retina with only cones o Visual acuity (sharpest vision) is here No photoreceptor cells are at the optic disc, or blind spot Structure of the Eye: Sensory Layer Cone sensitivity o Three types of cones o Different cones are sensitive to different wavelengths o Color blindness is the result of the lack of one cone type Lens Biconvex crystal-like structure Held in place by a suspensory ligament attached to the ciliary body © 2015 Pearson Education, Inc. Lens Cataracts result when the lens becomes hard and opaque with age o Vision becomes hazy and distorted o Eventually causes blindness in affected eye Risk factors include: o Diabetes mellitus o Frequent exposure to intense sunlight o Heavy smoking Two Segments, or Chambers, of the Eye Lens divides the eye into two chambers: 1. Anterior (aqueous) segment Anterior to the lens Contains aqueous humor 2. Posterior (vitreous) segment Posterior to the lens Contains vitreous humor Anterior Segment Aqueous humor o Watery fluid found between lens and cornea o Similar to blood plasma o Helps maintain intraocular pressure o Provides nutrients for the lens and cornea o Reabsorbed into venous blood through the scleral venous sinus, or canal of Schlemm Posterior Segment Vitreous humor o Gel-like substance posterior to the lens o Prevents the eye from collapsing o Helps maintain intraocular pressure Ophthalmoscope Instrument used to illuminate the interior of the eyeball and fundus (posterior wall) Can detect diabetes, arteriosclerosis, degeneration of the optic nerve and retina Pathway of Light Through the Eye Light must be focused to a point on the retina for optimal vision Light is bent, or refracted, by the cornea, aqueous humor, lens, and vitreous humor The eye is set for distance vision (over 20 feet away) © 2015 Pearson Education, Inc. Accommodation—the lens must change shape to focus on closer objects (less than 20 feet away) Pathway of Light Through the Eye Image formed on the retina is a real image Real images are: o Reversed from left to right o Upside down o Smaller than the object Pathway of Light Through the Eye The pathway of light through the eye: 1. Cornea 2. Aqueous humor 3. Through pupil 4. Aqueous humor 5. Lens 6. Vitreous humor 7. Retina Visual Fields and Visual Pathways Optic chiasma o Location where the optic nerves cross o Fibers from the medial side of each eye cross over to the opposite side of the brain Optic tracts o Contain fibers from the lateral side of the eye on the same side and the medial side of the opposite eye Visual Fields and Visual Pathways Overlap of the visual fields, and inputs from both eyes to each optic cortex provide for depth perception Pathway of Nerve Impulses into Brain The pathway of nerve impulses from the retina of the eye into the brain: 1. Optic nerve 2. Optic chiasma 3. Optic tract 4. Thalamus 5. Optic radiation 6. Visual cortex in occipital lobe of brain © 2015 Pearson Education, Inc. Eye Reflexes Internal muscles are controlled by the autonomic nervous system o Photopupillary reflex: bright light causes pupils to constrict through action of radial, circular, and ciliary muscles o Accommodation pupillary reflex: viewing close objects causes accommodation Viewing close objects causes convergence (eyes moving medially) A Closer Look Emmetropia—eye focuses images correctly on the retina Myopia (nearsightedness) o Distant objects appear blurry o Light from those objects fails to reach the retina and are focused in front of it o Results from an eyeball that is too long A Closer Look Hyperopia (farsightedness) o Near objects are blurry, whereas distant objects are clear o Distant objects are focused behind the retina o Results from an eyeball that is too short or from a “lazy lens” A Closer Look Astigmatism o Images are blurry o Results from light focusing as lines, not points, on the retina because of unequal curvatures of the cornea or lens Homeostatic Imbalances of the Eyes Night blindness—inhibited rod function that hinders the ability to see at night Color blindness—genetic conditions that result in the inability to see certain colors o Due to the lack of one type of cone (partial color blindness) Homeostatic Imbalances of the Eyes Glaucoma—can cause blindness due to increasing pressure within the eye Hemianopia—loss of the same side of the visual field of both eyes; results from damage to the visual cortex on one side only The Ear Houses two senses: 1. Hearing 2. Equilibrium (balance) Receptors are mechanoreceptors © 2015 Pearson Education, Inc. Different organs house receptors for each sense Anatomy of the Ear The ear is divided into three areas: 1. External (outer) ear 2. Middle ear (tympanic cavity) 3. Inner ear (bony labyrinth) The External Ear Involved in hearing only Structures of the external ear o Auricle (pinna) o External acoustic meatus (auditory canal) Narrow chamber in the temporal bone Lined with skin and ceruminous (wax) glands Ends at the tympanic membrane (eardrum) The Middle Ear (Tympanic Cavity) Air-filled cavity within the temporal bone Involved only in the sense of hearing Located between tympanic membrane and oval window and round window The Middle Ear (Tympanic Cavity) Two tubes are associated with the inner ear: 1. The opening from the auditory canal is covered by the tympanic membrane 2. The pharyngotympanic, or auditory, tube connects the middle ear with the throat Allows for equalizing pressure during yawning or swallowing This tube is otherwise collapsed Bones of the Middle Ear (Tympanic Cavity) Three bones (ossicles) span the cavity: 1. Malleus (hammer) 2. Incus (anvil) 3. Stapes (stirrup) Function o Vibrations from tympanic membrane move the hammer anvil stirrup oval window of inner ear Inner Ear or Bony Labyrinth Includes sense organs for hearing and balance © 2015 Pearson Education, Inc. Filled with perilymph Contains a maze of bony chambers within the temporal bone: o Cochlea o Vestibule o Semicircular canals Membranous labyrinth is suspended in perilymph and contains endolymph Organs of Equilibrium Equilibrium receptors of the inner ear are called the vestibular apparatus Vestibular apparatus has two functional parts: 1. Static equilibrium 2. Dynamic equilibrium Static Equilibrium Maculae—receptors in the vestibule o Report on the position of the head o Send information via the vestibular nerve Anatomy of the maculae o Hair cells are embedded in the otolithic membrane o Otoliths (tiny stones) float in a gel around the hair cells o Movements cause otoliths to bend the hair cells Dynamic Equilibrium These receptors respond to angular or rotary movements Crista ampullaris (in the ampulla of each semicircular canal)—dynamic equilibrium receptors are located in the semicircular canals o Tuft of hair cells covered with cupula (gelatinous cap) o If the head moves, the cupula drags against the endolymph Dynamic Equilibrium Action of angular head movements o The movement of the cupula stimulates the hair cells o An impulse is sent via the vestibular nerve to the cerebellum Organs of Hearing Spiral organ of Corti o Located within the cochlear duct o Receptors hair cells on the basilar membrane o Gel-like tectorial membrane is capable of bending hair cells o Cochlear nerve attached to hair cells transmits nerve impulses to auditory cortex on temporal lobe © 2015 Pearson Education, Inc. Mechanism of Hearing Vibrations from sound waves move tectorial membrane Hair cells are bent by the membrane An action potential starts in the cochlear nerve o Impulse travels to the temporal lobe Continued stimulation can lead to adaptation Mechanism of Hearing High-pitched sounds disturb the short, stiff fibers of the basilar membrane o Receptor cells close to the oval window are stimulated Low-pitched sounds disturb the long, floppy fibers of the basilar membrane o Specific hair cells further along the cochlea are affected Hearing and Equilibrium Deficits Deafness is any degree of hearing loss o Conduction deafness results when the transmission of sound vibrations through the external and middle ears is hindered o Sensorineural deafness results from damage to the nervous system structures involved in hearing o Ménière’s syndrome affects the inner ear and causes progressive deafness and perhaps vertigo (sensation of spinning) Chemical Senses: Taste and Smell Both senses use chemoreceptors o Stimulated by chemicals in solution o Taste has four types of receptors o Smell can differentiate a large range of chemicals Both senses complement each other and respond to many of the same stimuli Olfaction—The Sense of Smell Olfactory receptors are in roof of nasal cavity o Olfactory receptors cells (neurons) with long cilia known as olfactory hairs detect chemicals o Chemicals must be dissolved in mucus for detection by chemoreceptors called olfactory receptors Impulses are transmitted via the olfactory filaments to the olfactory nerve Interpretation of smells is made in the cortex Taste Buds and the Sense of Taste Taste buds house the receptor organs Locations of taste buds o Most are on the tongue © 2015 Pearson Education, Inc. o Soft palate o Cheeks Taste Buds and the Sense of Taste The tongue is covered with projections called papillae o Filiform papillae—sharp with no taste buds o Fungiform papillae—rounded with taste buds o Circumvallate papillae—large papillae with taste buds Taste buds are found on the sides of papillae Structure of Taste Buds Gustatory cells are the receptors o Possess gustatory hairs (long microvilli) o Hairs are stimulated by chemicals dissolved in saliva Structure of Taste Buds Impulses are carried to the gustatory complex by several cranial nerves because taste buds are found in different areas o Facial nerve (cranial nerve VII) o Glossopharyngeal nerve (cranial nerve IX) o Vagus nerve (cranial nerve X) Taste buds are replaced frequently by basal cells Taste Sensations Sweet receptors respond to sugars, saccharine, some amino acids Sour receptors respond to H ions or acids Bitter receptors respond to alkaloids Salty receptors respond to metal ions Umami receptors respond to the amino acid glutamate or the beefy taste of meat Developmental Aspects of the Special Senses Special sense organs are formed early in embryonic development Maternal infections during the first 5 or 6 weeks of pregnancy may cause visual abnormalities as well as sensorineural deafness in the developing child Congenital ear problems usually result from missing pinnas and closed or missing external acoustic meatuses Developmental Aspects of the Special Senses Vision requires the most learning The infant has poor visual acuity (is farsighted) and lacks color vision and depth perception at birth © 2015 Pearson Education, Inc. The eye continues to grow and mature until age 8 or 9 Developmental Aspects of the Special Senses Eye problems o Strabismus—“crossed eyes”; results from unequal pulls by the external eye muscles in babies o Ophthalmia neonatorum—conjunctivitis resulting from gonorrhea in the mother; baby’s eyelids are swollen, and pus is produced Developmental Aspects of the Special Senses Problems of aging associated with vision include presbyopia, glaucoma, cataracts, and arteriosclerosis of the eye’s blood vessels o Presbyopia—“old vision” results from decreasing lens elasticity that accompanies aging Developmental Aspects of the Special Senses The newborn infant can hear sounds, but initial responses are reflexive By the toddler stage, the child is listening critically and beginning to imitate sounds as language development begins Age-related ear problems: o Presbycusis—type of sensorineural deafness that may result from otosclerosis Otosclerosis—ear ossicles fuse Developmental Aspects of the Special Senses Taste and smell are most acute at birth and decrease in sensitivity after age 40 as the number of olfactory and gustatory receptors decreases © 2015 Pearson Education, Inc.