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I. Origin of Life on Earth C. Characteristics of Early Life 1. Self-Replication • • • • • 2. Proteins aren’t self-replicating RNA may carry out catalytic functions Ribozyme – Autocatalytic RNA RNA may have been first informational molecule DNA more stable; may have arisen from RNA Nutrition • • • • • • First cells likely heterotrophic No free oxygen in atmosphere of early earth First heterotrophs probably used anaerobic fermentation (less efficient than aerobic metabolism) First autotrophs may have used hydrogen sulfide (H2S) as hydrogen source (modern purple & green sulfur bacteria still get H from H2S) First autotrophs to split water for H probably ancestors of modern cyanobacteria (3.1 – 3.5 bya) Production of O2 had profound effects I. Origin of Life on Earth C. Characteristics of Early Life 3. Aerobes • • • • • 4. O2 abundant by 2.5 bya Replaced obligate anaerobes in most areas Aerobic metabolism much more efficient than anaerobic metabolism Stabilized concentrations of O2 and CO2 in atmosphere Development of ozone (O3) layer Eukaryotes • • • • Appeared ~2.1-2.2 bya How might eukaryotes have arisen from prokaryotes? Organelles (mitochondria, chloroplasts) may have originated from symbiotic relationships between prokaryote species • Chloroplasts closely related to cyanobacteria • Mitochondria closely related to alpha proteobacteria Serial endosymbiosis Fig. 25.8 I. Origin of Life on Earth C. Characteristics of Early Life 3. Aerobes • • • • • 4. O2 abundant by 2.5 bya Replaced obligate anaerobes in most areas Aerobic metabolism much more efficient than anaerobic metabolism Stabilized concentrations of O2 and CO2 in atmosphere Development of ozone (O3) layer Eukaryotes • • • • Appeared ~2.1-2.2 bya How might eukaryotes have arisen from prokaryotes? Organelles (mitochondria, chloroplasts) may have originated from symbiotic relationships between prokaryote species • Chloroplasts closely related to cyanobacteria • Mitochondria closely related to alpha proteobacteria Serial endosymbiosis Fig. 25.9 Fig. 25.9 Fig. 25.9 I. Origin of Life on Earth C. Characteristics of Early Life 4. Eukaryotes • a. b. c. d. Evidence for serial endosymbiosis Inner membranes of plastids & mitochondria have enzymes and transport systems similar to those of plasma membranes in modern bacteria Plastids & mitochondria replicate by binary fission process similar to that of bacteria Plastids & mitochondria each contain single, circular DNA molecule without histones or other proteins (similar to bacteria) Plastids & mitochondria have ribosomes that resemble prokaryotic more than cytoplasmic ribosomes (size, sequence, sensitivity to antibiotics) II. Geological Record • • Rocks, sediments, fossils – Occur in layers (strata) Oldest fossils – Stromatolites from 3.5 bya II. Geological Record A. Dating 1. Index Fossils • • • 2. Based on common species Useful for establishing relative ages Used by petroleum industry Radiometric Dating • • Technique for absolute dating Based on decay of radioactive elements in rocks Fig. 25.5 II. Geological Record A. Dating 1. 2. Index Fossils Radiometric Dating • • • Half-life unaffected by temperature, pressure, etc. Ex: 40K 40Ar with t0.5 = 1.3 billion years • Initial rock has 100% 40K and no 40Ar • Rock with 40K:40Ar = 1:1 is 1.3 billion years old • Rock with 40K:40Ar = 1:3 is 2.6 billion years old Commonly used radioisotopes 40K with t • 0.5 = 1.3 billion years 235U with t • 0.5 = 704 million years 14C with t • 0.5 = 5730 years II. Geological Record B. Geological Time Scale • Earth’s history divided into periods based on major geological, climatic, & biological changes Mass Extinction (K-T) First Bird (150 mya) ? Mass Extinction (96% of marine spp.) First Amniote Egg ? ”Age of Fishes” First Land Plants/Animals ? First Fishes - Siberian Volcanism - Increased CO2 - Altered Ocean Mixing Fig. 25.15 Cretaceous Mass Extinction Fig. 25.16 Time Scale: One Year Homo appears: Dec 31 @ 5 pm Fig. 25.7 Global Plate Tectonics Jurassic to Present Day By L.A. Lawver, M.F. Coffin, I.W.D. Dalziel L.M. Gahagan, D.A. Campbell, and R.M. Schmitz 2001, University of Texas Institute for Geophysics February 9, 2001 Earth – Future Drift