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Types of Muscle There are three types of muscle: voluntary or skeletal involuntary or smooth muscle cardiac muscle Their names variously suggest where they are found and the type of control we have over them. All of the muscle types contain contractile fibres made from actin and myosin, but there are structural and functional differences between them. Voluntary or skeletal muscle Voluntary or skeletal muscle is also known as striated or striped muscle because of its physical appearance – when looked at under the microscope, it has a striped appearance. Contraction of voluntary muscle is under conscious control, and moves the bones of the skeleton across joints. Skeletal muscle consists of fibres surrounded by a membrane or sarcolemma, within which there are several nuclei, many mitochondria and an extensive sarcoplasmic reticulum. The mitochondria provide the ATP energy for muscle contraction and the sarcoplasmic reticulum stores Ca2+ ions, essential for contraction of the muscle myofibrils divided into contractile units (sarcomeres). It is the regular arrangement of the actin and myosin filaments in the sarcomeres that give voluntary muscle its striped appearance. Contraction of voluntary muscle is rapid and forceful, though prone to fatigue. muscle, comprised of bundles of many fibres nucleus I-Band tendon, attaching muscle to bone part of muscle fibre A-Band myofibril Z-line Z-line sarcomere Involuntary or smooth muscle Contraction of smooth muscle is controlled by the autonomic nervous system and therefore is not under voluntary control. It is found in the walls of blood vessels, the gut and the urino-genital system. The cells are spindle shaped, have a single nucleus and are not arranged into fibres, so are not striated. Contraction is relatively slow and the muscle does not tire or fatigue. The diagram and light micrograph show smooth muscle in a transverse section of a trachea. Nucleus Myofibril Spindle-shaped cell Cardiac muscle As the name implies, this is found in the heart. It has some structural similarity to striped muscle – striations are visible in the muscle cells. However adjacent cardiac muscle fibres interconnect, their plasma membranes make very close contact and are freely permeable, so they allow electrical excitation (action potentials) to pass easily between cardiac muscle cells. Like smooth muscle, cardiac muscle is myogenic and will contract rhythmically without nervous stimulation. The net-like arrangement of cardiac muscle, close interconnections and the high conductivity of the muscle itself mean that contraction can spread quickly over the heart. Cardiac muscle contracts forcefully, but does not fatigue. The structure of cardiac muscle is shown in the diagram and micrograph: Myofibril Intercalated disc Striations Bridge Nucleus Space between fibres for blood vessels and connective tissue Note: The cells that make up the SAN, AVN and Purkyne tissue are modified cardiac muscle cells. They do contract, albeit weakly. However, they can generate and conduct impulses that control normal heartbeat. That said, the output of the heart is modified by the autonomic nervous system to suit the body’s requirements. Complete the following summary to compare voluntary, involuntary and cardiac muscle: Voluntary Muscle type Involuntary Cardiac Where found Characteristics of contraction (control, speed, force, if prone to fatigue) Structure Atrial and ventricular muscle SAN Function AVN Purkyne tissue