Survey
* Your assessment is very important for improving the workof artificial intelligence, which forms the content of this project
* Your assessment is very important for improving the workof artificial intelligence, which forms the content of this project
Chapter 9 Photosynthesis and Cellular Respiration Section 1: Energy in Living Systems Preview Chemical Energy Metabolism and the Carbon Cycle Transferring Energy Summary Chemical Energy Organisms require a constant source of energy. Energy is needed for organisms to maintain their homeostasis. Homeostasis is the process of maintaining internal order and balance even when the environment changes. Organisms use and store energy in the chemical bonds of organic compounds. Almost all of the energy in organic compounds comes from the sun. Chemical Energy Photosynthesis is the process by which plants, algae, and some bacteria use sunlight, carbon dioxide, and water to produce carbohydrates and oxygen. Organisms that are able to perform photosynthesis, such as plants, are autotrophs. Autotrophs make organic compounds that serve as food for them and for almost all of the other organisms on Earth, Visual Concept: Photosynthesis Chemical Energy, continued Organisms that cannot make their own food must absorb food molecules made by autotrophs, eat autotrophs, or eat organisms that consume autotrophs. Food molecules that are made or consumed by an organism are the fuel for its cells. Cells use these molecules to release the energy stored in the molecules’ bonds. The Visual Concept: Cellular Respiration Click above to play the video. Metabolism and the Carbon Cycle Metabolism involves either using energy to build organic molecules or breaking down organic molecules in which energy is stored. Organic molecules contain carbon. Therefore, an organism’s metabolism is part of Earth’s carbon cycle. Energy from the sun is converted to chemical energy in chloroplasts. Metabolism and the Carbon Cycle, continued Organisms extract energy in glucose molecules. Through the process of cellular respiration, cells make the carbon in glucose into stable carbon dioxide molecules and produce energy. Energy is also released and used to make ATP (adenosine triphospate), an organic molecule that is the main energy source for cell Visual Concept: Linking Photosynthesis and Respiration Transferring Energy In chemical reactions, energy can be absorbed and released during the breaking and forming of bonds. In cells, chemical energy is gradually released in a series of chemical reactions that are assisted by enzymes. Enzymes are proteins that act as catalysts in biochemical reactions. Transferring Energy, continued ATP When cells break down food molecules, some of the molecules is released as heat. Much of the remaining energy to make ATP. ATP is a portable form of energy “currency” inside cells. ATP is a nucleotide made up of a chain of three phosphate groups. When the bond of the third phosphate group is broken, energy is released, ATP Releases Energy Visual Concept: Comparing ATP and ADP Click above to play the video. Transferring Energy, continued In many cells, ATP synthase recycles ADP by bonding a third phosphate group to the molecule to form ATP. ATP synthase acts as both an enzyme and a carrier protein Transferring Energy, continued The flow of H+ ions through ATP synthase powers the production of ATP. In chloroplasts and mitochondria, a series of molecules, called an electron transport chain, pump H+ ions across the membrane to create a concentration gradient. The electron transport chain uses energy from released from electron carriers, such as NADH and NADPH, to pump hydrogen ions. Electron carriers Summary Organisms use and store energy in the chemical bonds of organic molecules. Metabolism involves either using energy to build organic molecules or breaking down organic molecules in which energy is stored. Organic molecules contain carbon. Therefore, an organism’s metabolism is part of Earth’s carbon cycle. In cells, chemical energy is gradually released in a series of chemical reactions that are Concept Check 1. What type of energy is used in cells, and what is the ultimate source of this energy? 2. How is an organism’s metabolism related to the carbon cycle? 3. How is energy released in a cell? Test Prep 1. What pigment causes a plant to look green? A. NADH B. NAPH C. carotenoid D. chlorophyll 2. What is the product of the electron transport chain of photosynthesis? A. water B. glucose C. pyruvate D. ATP and NADPH Section 2: Photosynthesis Preview Harvesting Light Energy Two Electron Transport Chains Producing Sugar Factors that Affect Photosynthesis Summary Harvesting Light Energy Photosynthesis is the process that provides energy for almost all life. Chloroplasts are the organelles that convert light energy into chemical energy. Within the inner membrane of the chloroplast, is the stroma which contains the thylakoid membrane. This membrane produces flat, disc-like sacs Thylakoids Harvesting Light Energy, continued Light is a form of electromagnetic radiation, energy that can travel through empty space in the form of waves. Sunlight contains all of the wavelengths of visible light which we see as different colors. A pigment is a substance that absorbs certain wavelengths (colors) of light and reflects all of the others. Harvesting Light Energy, Chlorophyll is a green pigment in chloroplasts that absorbs light energy to start photosynthesis. It absorbs mostly blue and red light and reflects green and yellow light, which makes plants appear green. Harvesting Light Energy, continued Plants also have pigments called carotenoids which help plants absorb additional light energy. When light hits a thylakoid, energy is absorbed by many pigment molecules and eventually transferred to electron carriers. Visual Concept: Spectrum of Light and Plant Pigments Two Electron Transport Chains Electrons from the electron carrier are used to produce new molecules, including ATP, that temporarily store chemical energy. During photosynthesis, one electron transport chain provides energy to make ATP, while the other provides energy to make NADPH. Both chains use energy from electrons excited by light. Two Electron Transport Chains, continued Producing ATP Step 1: Electrons excited by light leave the chlorophyll molecules. An enzyme splits water molecules to replace these electrons. Oxygen gas is formed and released into the atmosphere. Two Electron Transport Chains, continued Producing ATP Step 2: Excited electrons transfer some of their energy to pump H+ ions into the thylakoid. This process creates a concentration gradient across the thylakoid membrane. Two Electron Transport Chains, continued Producing ATP Step 3: The energy from diffusion of H+ ions through the channel portion of ATP synthase is used to catalyze a reaction in which a phosphate group is added to a molecule of ADP, producing Two Electron Transport Chains, continued Producing NADPH Step 4: Light excites electrons in another chlorophyll molecule. The electrons are passed on to the second chain and replaced by the deenergized electrons from the first chain. Two Electron Transport Chains, continued Producing NADPH Step 5: Excited electrons combine with H+ ions and NADP+ to form NADPH. NADPH is an electron carrier that provides high-energy electrons needed to Electron Transport Chains of Photosynthesis Click to animate the image. Producing Sugar The first two stages of photosynthesis depend directly on light because light energy is used to make ATP and NADPH. (a.k.a. Light Reactions) In the final stage of photosynthesis, ATP and NADPH are used to produce energy-storing sugar molecules from the carbon in carbon dioxide. Producing Sugar, continued The reactions that fix carbon dioxide are lightindependent reactions, sometimes called dark reactions. The most common method of carbon fixation is the Calvin cycle. Atmospheric carbon dioxide is combined with other carbon compounds to produce organic compounds. ATP and NADPH supply some of Visual Concept: Calvin Cycle Click above to play the video. Factors that Affect Photosynthesis Light intensity, carbon dioxide concentration, and temperature are three environmental factors that affect photosynthesis. Although different plants are adapted to different levels of light, the photosynthesis rate increases with increases in light intensity until all of the pigments in a chloroplast are being used. Photosynthesis is most efficient in a certain range of temperatures. Visual Concept: Environmental Influences on Photosynthesis Summary In plants, light energy is harvested by pigments located in the thylakoid membrane of chloroplasts. During photosynthesis, one electron transport chain provides energy used to make ATP, while the other provides energy to make NADPH. Summary, continued In the final stage of photosynthesis, chemical energy is stored by being used to produce sugar molecules from the carbon in the gas carbon dioxide. Light intensity, carbon dioxide concentration, and temperature are three environmental factors that affect photosynthesis. Concept Check 1. What is the role of pigments in photosynthesis? 2. What are the roles of the electron transport chains? 3. How do plants make sugars and store extra unused energy? 4. What are the three environmental factors that affect photosynthesis? Test Prep 3. The oxygen that is produced during photosynthesis comes directly from the A. absorption of light. B. mitochondrial membranes. C. splitting of water molecules. D. splitting of carbon dioxide molecules. Use the diagram of a chloroplast to answer the following question. 4. Which of the following correctly identifies the structure marked X and the activities that take place there? A. stroma-Calvin cycle B. thylakoid-Calvin cycle C. stroma-light reactions D. thylakoid-light reactions Section 3: Cellular Respiration Preview Glycolysis Aerobic Respiration Fermentation Summary Glycolysis The cells of most organisms transfer energy found in organic compounds, such as those in foods, to ATP. The primary fuel for cellular respiration is glucose. Fats can be broken down to make ATP. Proteins and nucleic acids can also be used to make ATP, but they are usually used for Glycolysis, continued In glycolysis, enzymes break down one sixcarbon molecule of glucose into two threecarbon pyruvate molecules. The breaking of a sugar molecule by glycolysis results in a net gain of two ATP molecules. This process of glycolysis is anaerobic, or takes place without oxygen. Visual Concept: Glycolysis Click above to play the video. Glycolysis, continued Glycolysis is the only source of energy for some prokaryotes. Other organisms use oxygen to release even more energy from a glucose molecule. Metabolic processes that require oxygen are aerobic. In aerobic respiration, the pyruvate product of glycolysis undergoes another series of Glycolysis Aerobic Respiration Organisms such as humans can use oxygen to produce ATP efficiently through aerobic respiration. The first stage of aerobic respiration is the Krebs cycle, a series of reactions that produce electron carriers. The electron carriers enter an electron transport chain, which powers ATP synthase. Up to 34 ATP molecules can be produced from one glucose molecule in aerobic respiration. Aerobic Respiration, continued Krebs Cycle Pyruvate (from glycolysis) is broken down and combined with other carbon compounds. Each time the carbon-carbon bonds are rearranged during the Krebs cycle, energy is released. The total yield of energy-storing products from one time through the Krebs cycle is one ATP, three NADH, and one FADH2. Visual Concept: Krebs Cycle Click above to play the video. Aerobic Respiration, continued Electron Transport Chain The second stage of aerobic respiration takes place in the inner membranes of mitochondria, where ATP synthase enzymes are located. Electron carriers, produced during the Krebs cycle, transfer energy through the electron transport chain. Energy from the electrons is used to actively transport hydrogen ions out of the inner Visual Concept: Electron Transport Chain Click above to play the video. Aerobic Respiration, continued Electron Transport Chain Hydrogen ions diffuse through ATP synthase, providing energy to produce several ATP molecules from ADP. At the end of the electron transport chain, the electrons combine with an oxygen atom and two hydrogen ions to form two water molecules. If oxygen is not present, the electron transport Fermentation To make ATP during glycolysis, NAD+ is converted to NADH. Organisms must recycle NAD+ to continue making ATP through glycolysis. The process in which carbohydrates are broken down in the absence of oxygen is called fermentation. Fermentation enables glycolysis to continue Fermentation, continued In lactic acid fermentation, pyruvate is converted to lactic acid. During vigorous exercise, lactic acid fermentation also occurs in the muscles of animals, including humans. During alcoholic fermentation, one enzyme removes carbon dioxide from pyruvate. A second enzyme converts the remaining Two Types of Fermentation Fermentation, continued Efficiency of Cellular Respiration In the first stage of cellular respiration, glucose is broken down to pyruvate during glycolysis, an anaerobic process. Glycolysis results in a net gain of two ATP molecules for each glucose molecule that is broken down. In the second stage, pyruvate either passes through the Krebs cycle or undergoes Fermentation, continued Efficiency of Cellular Respiration Cells release energy most efficiently when oxygen is present because they make most of their ATP during aerobic respiration. For each glucose molecule that is broken down, as many as two ATP molecules are made during the Krebs cycle. The Krebs cycle feeds NADH and FADH2 to the electron transport chain, which can produce Visual Concept: Comparing Aerobic and Anaerobic Respiration Summary The breaking of a sugar molecule by glycolysis results in a net gain of two ATP molecules. The total yield of energy-storing products from one time through the Krebs cycle is one ATP, three NADH, and one FADH2. Summary, continued Electron carriers transfer energy through the electron transport chain, which ultimately powers ATP synthase. Fermentation enables glycolysis to continue supplying a cell with ATP in anaerobic conditions. Concept Check 1. How does glycolysis produce ATP? 2. How is ATP produced in aerobic respiration? 3. Why is fermentation important? Test Prep 5. Which of the following is not involved in the aerobic part of cellular respiration? A. ATP B. glycolysis C. mitochondria D. the Krebs cycle The graph shows data on photosynthesis in one type of plant. Use the graph and your knowledge Photosynthesis not ofA.science todoes answer the following occur at 0 °C. question. B. The rate of at 40 °C is 6. photosynthesis Which statement is greater than the rate at 20 supported by the data? °C. C. The optimum temperature for photosynthesis is approximately 46 °C. D. The rate of photosynthesis increases as temperature increases from 25 °C to 30 °C. 7. The inner membrane of a mitochondrion is folded. These folds are called christae. How might cellular respiration be different if the inner mitochondrial membrane were not folded?