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Download Lecture 2: Wildlife Ecological Principles and Population Ecology Part 1
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Realm Biome Ecosystem ECOLOGY Population Organism Organ Tissue Taxonomy Anatomy Physiology Genetics Cell Organelles Molecules Atoms Biochemistry Genomics Molecular Biology More complex, Larger Less Abundant Community REALM BIOME ECOREGION LANDSCAPE ECOSYSTEM COMMUNITY POPULATION/SPECIES GENETIC Realms = large regions with ecosystems share a broadly similar biota. Mostly terrestrial and freshwater oriented Oceanic Antartic Australasian Nearctic Paleartic Afrotropic Indo-malay Biome (region of similar vegetation) Biomes are the major regional groupings of plants and animals discernible at a global scale. Their distribution patterns are strongly correlated with regional climate patterns and identified according to the climax vegetation type. However, a biome is composed not only of the climax vegetation, but also of associated successional communities, persistent subclimax communities, fauna, and soils. Biomes are determined by precipitation and temperature 2.4 Biomes and climate 867 Global Terrestrial Ecoregions Kentucky’s Major Recognized (Physiographic) Ecoregions Ecosystem Ecosystem = biological community + abiotic environment (the non-living things such as geology, water). Typically ecosystem is used to describe things at large scales. There are many examples of ecosystems -- a pond, a forest, an estuary, a grassland and their boundaries are not fixed in any objective way, although sometimes they might seem obvious. Often have complex food (trophic) webs and “self- contained” nutrient cycles such as nitrogen cycle Ecosystems Energy flow Nutrient cycles Absorption Plant By Remains Plants N cycle Bacteria Bacteria Nitrates Bacteria Ammonia Free N In Atmosphere Bacteria Chihuahuan Desert Ecosystem (example) Ecological Community A community is an assemblage of plant and animal populations that live in a particular area or habitat. The combinations of species within that community make it unique in the way it looks (composition and structure) and function (the ecological work it does). Community Properties Scale Spatial and Temporal Structure Species Diversity (Biodiversity) Trophic structure Succession and Disturbance Scale Size of community – water droplet to rotting log to forest floor Community scale dictated by the measurer/definer Spatial Structure Horizontal – Uniform, random, clumped Vertical – underground, forest floor, herb layer, shrub layer, mid-story, canopy canopy Mid-story Shrub Forest floor Below ground Herb 14,000 ft Vertical Community Spatial Structuring South Facing Slopes 12,000 ft North Facing Slopes Alpine Zone Alpine tundra Subapline Zone Spruce-fir Forest 10,000 ft 8,000 ft Foothills Zone Pinyon-Juniper Sagebrush Woodland 6,000 ft Upper Montane Zone – Mixed Conifer Forest Lower Montane Zone – Ponderosa Pine Forest Vertical Community Spatial Structuring Alpine Zone Alpine tundra Subapline Zone Spruce-fir Forest Upper Montane Zone – Mixed Conifer Forest Foothills Zone Pinyon-Juniper Sagebrush Woodland Lower Montane Zone – Ponderosa Pine Forest Temporal Structure Phenology = timing of appearance and activity of species (e.g. spring wildflower blooming period) Species ECOSYSTEM COMMUNITY POPULATION/SPECIES Population- is local occurrence of individuals of the same species; typically the unit of management used to maintain viability in natural patterns, abundances, and distributions. Species are defined as individuals that look identical or very similar and interbreed under natural conditions. E.g. Cheetahs of Serengeti National Park, Tanzania, Africa Species Diversity (Biodiversity) Species richness = # of species (simplest measure of biodiversity) Biodiversity Indices = # of species and their relative abundance within the community E.g. you count 100 birds in two forest blocks Block A has 95 starlings, 2 cardinals, 1 house wren, 1 blue jay, and 1 Carolina chickadee (a total of 5 species) Block B has 50 starlings, 20 cardinals, 10 house wrens, 5 blue jays, and 5 Carolina chickadees (a total of 5 species, but the individuals are more evenly distributed among species; therefore Block B would be considered more biodiverse) Trophic (feeding) levels Primary Producers (plants, phytoplankton, some bacteria – can make their own food – autotroph – self feeding) Heterotrophs – utilize autotrophs or others to obtain energy Primary Consumers (eat primary producers – herbivores – zooplankton, insects, rabbits, elephant) Secondary Consumer (carnivore – eat animals (called prey) microscopic predators to lions, tigers) Tertiary Consumer (shark, bass, kingsnake) Decomposers (bacteria, fungi;break down small bits of organic material and often recycle it back into soil, air, water) Omnivore – eats both plants and animals Herbivore – eats plants only Carnivore – generally solely a meat eater, or of the Order Carnivora Scavengers – (technically carnivores) eat dead animals Parasites – obtain nutrition from by living inside or on another organism Ecological Succession Ecological Succession: predictable change in species over time as new set of species modifies the environment to enable the establishment of another set of species. Each recognizable “step” in process is known as a seral stage. Disturbance Increasing Ecological Complexity and Stability Examples of Disturbance = Flood, Fire, Wind, Herbivory, Timber Harvesting, Mining, Road Building, Urbanization, Agriculture Succession Types Primary Succession – sequence of species (sere or seral stage) on landforms not influenced by a community, example, rock or retreating glacier Secondary Succession – vegetation partially or completed removed but seeds, seedlings, spores, etc. remain Climax community – self-perpetuating, mature community that is resistant to change and dominated by species that can tolerate competition – “final stage” Thinking About Succession Patterns and Processes Early in succession, species are generally excellent dispersers and tolerate harsh environments, but not the best inter-specific (between species) competitors. As ecological succession progresses, these are replaced with species which are better competitors, (but not as good at dispersing and more specialized to deal with the microenvironments created by other species likely to be present with them). Early species modify their environment in such a way as to make it possible for the next round of species. These, in turn, make their own replacement by superior competitors possible. Wildlife Habitat Wildlife Habitat - Place where organism lives Migration is a two way movement of an animal to and from an area with regularity or with changes in life history strategy – example Waterfowl, big game in Mountain west, African ungulate migration Niche (Elton 1927) Ecological Niche = abstract concept but in general means the role or function of organism in biotic community (it’s occupation) Multidimensional : where it seeks food, when it feeds, what size food it eats, where it reproduces Example elk and mule deer on winter range – overlap somewhat in habitat but elk hide in dense forest, mule deer in shrub cover Wading Birds Each has its own niche Bare Annual Ground Grass/forb Perennial Grass/forb Young Woodland Shrub Mature Woodland Bobwhite Quail Ruffed Grouse Cottontail Rabbit Gray Squirrel Wild Turkey Woodchuck White-tailed Deer Grass-forb Shrub Shrub – small tree Mature forest Generalized schema of some common terrestrial vertebrates along a successional gradient in an oak-hickory forest Fundamental niche – niche occupied by that organism when there is no competition from other species. Realized niche – niche occupied by species when some competition with other species is occurring. Niche segregation Tendency for species that live in same area and require similar resources to have niche requirements that differ in one or more dimensions. Niche segregation occurs when species that live in same area and require similar resources differ in one or more dimensions of habitat use (temporally or spatially). Eat same food base, great-horned owl active @ night, red-tailed hawk active @ day Eat same food base, coyote open habitat, bobcat forest habitat Competition Competition occurs when organisms in the same community seek the same limited resource; they have overlapping niche requirements. Resource may be but is not limited to such things like food, light, space, water, nest sites, etc. Interspecific competition: between individuals of two different species. Intraspecific competition: among individuals of the same species. Competitive Exclusion Principle No two species can simultaneously and completely occupy the same niche for an indefinite amount of time. “Styles” of Competition Exploitative competition occurs when individuals use the same limiting resource or resources, thus depleting the amount available to others. (e.g. squirrel eats all the acorns from a tree and leaves less or none for mice) Interference competition occurs when individuals interfere with the foraging, survival, or reproduction of others, or directly prevent their physical establishment in a portion of a habitat. (e.g. when coyote will not approach wolf kills or territory for fear of being killed) Outcomes of Competition Exploitative competition may cause the exclusion of one species. For this to occur, one organism must require less of the limiting resource to survive. The dominant species must also reduce the quantity of the resource below some critical level where the other species is unable to replace its numbers by reproduction. However, exploitative competition does not always cause the exclusion of one species. They may coexist, with a decrease in their potential for growth. For this to occur, they must partition the resource. Interference competition generally results in the exclusion of one of the two competitors.