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SBI3U Evolution – 2 Geologist Charles Lyell proposed that geological processes operated at the same rates in the past as they do today in a process called uniformitarianism. Lyell, therefore rejected catastrophism and theorized that slow, subtle processes could happen over a long period of time and could result in substantial change. For example, floods that happened in the past had no greater power than floods that occur today. This idea inspired naturalist Charles Darwin and others. If Earth is slowly changing, they wondered, could slow, subtle changes also occur in populations? French naturalist Jean-Baptiste Lamarck compared current species of animals with fossil forms and observed what he interpreted as a line of descent, or progression, in which a series of fossils (from older to more recent) led to a modern species that increased in complexity over time. Lamarck hypothesized that organisms could become progressively better adapted to their environments. For example, it was thought that giraffes stretched their necks to reach the foliage in tall trees and that this stretched neck condition was passed on to their offspring, resulting in tall giraffes that can eat from the tops of trees. Although this is not the way giraffes developed their long necks, Lamarck called this idea the inheritance of acquired characteristics. Lamarck also suggested that body parts not used would eventually disappear. At the time there was little understanding of cell biology and no understanding of genetics. Lamarck provided a hypothesis for how the inheritance of characteristics from one generation to the next might happen. More importantly, he noted that an organisms adaptations to the environment resulted in characteristics that could be inherited by offspring. In 1831, Charles Darwin sailed aboard the HMS Beagle, a British survey ship. The primary purpose of this voyage was to map the coast of South America but this journey provided Darwin with an opportunity to explore the natural history of various countries and geographical locations. At first, Darwin did not understand the significance of many of his observations but years later, many of them became important to his theory of evolution by natural selection. After his trip, Darwin began to propose answers to his questions and to compile his and others’ observations into a comprehensive theory to explain how species change over time. British naturalist Alfred Russel Wallace, reached conclusions that were similar to Darwin’s as they both accepted that populations changed as time passed, but they were unclear how they changed. An essay by economist Thomas Malthus called Essay on the Principles of Population, proposed that populations produced far more offspring than their environments could support. Malthus said that these populations were eventually reduced by starvation or disease. This gave Darwin and Wallace a key idea because according to them, individuals with traits that helped them to survive in their local environments were more likely to survive to pass on these traits to offspring. They reasoned that competition for limited resources between individuals of the same species would select for individuals with favorable traits – traits that increased their chances of surviving to reproduce. Therefore, these traits would appear in later generations, and as time passed, the entire population would have them. This is the idea of survival of the fittest, and Darwin called this process natural selection. Darwin proposed that all life descended from some unknown organism and as descendants of this organism spread out over different habitats over many years, they developed adaptations to help them better survive in their local environments. His ideas were published in 1859 in a book called The Origin of Species. Darwin’s theory of natural selection showed how populations of individual species became better adapted to their local environments over time. His ideas can be summarized as:  Organism’s produce more offspring than can survive. Therefore, organisms compete for limited resources.  Individuals of a population vary extensively, and much of this variation is heritable  Individuals that are better suited to local conditions survive to produce more offspring  Processes for change are slow and gradual. Darwin did not use the word evolution in the original copy of The Origin of the Species. Instead, he used the term descent with modification as he felt the word evolution implied progress. Darwin did not believe that natural selection demonstrated progress but rather is the result of the ability of certain individuals in any population to survive local environmental conditions and to pass on traits that helped them survive in the first place.
 
									 
									 
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                            