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Diversity and Evolution SC.912.L.151 SC.912.L.15.10 SC.912.N.1.3 SC.912.N.2.1 SC.912.L.15.8 Evolution Homozygous: both alleles are the same Heterozygous: each allele is different • Study of changes in populations, species, or groups of species • Occurs because populations VARY by the frequency (number of) heritable traits that appear from one generation to the next – Traits are represented by alleles • Fancy word for a version of a specific gene that controls a trait – Ex. You have a gene for eye color….you INHERIT different alleles for eye color…blue alleles or brown alleles (you get one from each parent) • Evolution is… – Changes in ALLELE frequencies in populations over time Two areas to study in evolution… • Microevolution •Macroevolution Microevolution • Describes the details of how populations of organisms change from generation to generation • Describes how a new species originates (speciation) • Change in gene frequency in a population • When evolution occurs quickly enough for scientists to observe Examples: • Bacteria – Reproduce rapidly – Trait for Resistance to antibiotics is passed down to offspring – This is how bacteria become resistant to certain medicines…NOT GOOD • Plants – Resistance to herbicides • Insects – Resistance to pesticides Macroevolution • Patterns of evolution for groups of species over extended periods of GEOLOGIC time • Patterns determine PHYLOGENY – The evolutionary relationships among species and groups of species • Remember our cladograms???? • 2 distinct macroevolution theories – Phyletic gradualism – Punctuated equilibrium • Gradualism – – – – – Two Ways MACROevolution is believed to have Occurred Evolution occurring over a long period of time Slow and constant change Small difference between generations This is how Darwin believed NATURAL SELECTION worked Evidence: • Fossil records only provide snap shots of evolutionary process; reveals only major changes in groups of organisms • Incompleteness of our fossil record explains the absence of the intermediate stages of development • Punctuated Equilibrium – Evolution occurs in spurts – Causes: • random mutations in DNA • Sudden Major changes in environment – Changes occur rapidly followed by long period without change – Evidence: • Fossil history consists of fossils of mostly extended periods of “stasis” (stability) with few (if any) fossils from short, rapid bursts of evolution. • Absence of fossils from intermediate stages of evolution confirms PUNCUATED EQUILIBRIUM MICROEVOLUTION Describes details of how POPULATIONS of organisms change from generation to generation and how NEW species originate Species: group of individuals capable of interbreeding and producing viable offspring Speciation: formation of a new species Before we get into speciation… • Speciation is getting one group of species to split into new, different groups • This large group MUST become separated or ISOLATED in some way so they can NO longer interbreed… • 7 types of isolation that can lead to speciation…. • Isolating Mechanisms How to MAINTAIN reproductive isolation • Review: What is a ZYGOTE??? – Pre-zygotic isolating mechanisms (before a zygote is even made) • Ways to PREVENT fertilization 1. 2. 3. 4. Habitat (geographical) isolation Temporal isolation Behavioral isolation Reproductive Isolation » Mechanical isolation » Gametic isolation – Post-zygotic isolating mechanisms (after zygote is made) • Ways to prevent the formation of FERTILE offspring 5. 6. 7. Hybrid inviability Hybrid sterility Hybrid breakdown Pre-zygotic isolating mechanisms (before a zygote is even made) 1. 2. 3. 4. Habitat isolation • Species A never Encounters species B • Species and mates/flowers at different time or season than species B Temporal isolation Behavioral isolation • Species A doesn’t recognize species B’s mating ritual • Mechanical isolation • Reproductive parts are incompatible Gametic isolation • Male gametes do not survive in the environment of the female gamete • Female gametes do not recognize male gametes Reproductive Isolation • How We Get New Species 1. Reproductive isolation – When organisms of the same species become genetically split and STOP breeding with each other – Gene pool split, two different groups can no longer interbreed • 2. 3. 4. Mechanical or gametic isolation Behavioral isolation – Groups of organisms of the same species develop different courtship rituals (or behaviors) over time and will not be attracted to others Geographic isolation – When two populations are separated by geographic barriers – Mountains, rivers, etc. – Doesn't always separate all species…river and birds vs. squirrels Temporal isolation – When two or more species reproduce at different times – Specific day/season when that species is able reproduce Post-zygotic isolating mechanisms (after zygote is made) 5. Hybrid inviability • Zygote fails to develop properly • Aborts or dies before reaching reproductive maturity (usually at embryonic stage) 6. 7. Hybrid sterility • Hybrid becomes functional adult but they are reproductively sterile • Eggs or sperm are non-existent or dysfunctional Hybrid breakdown • Hybrids produced actually DO produce offspring but their offspring have reduced viability or fertility • Microevolution: Observed Evolutionary Change – Describes details of how POPULATIONS of organisms change from generation to generation and how NEW species originate – Species: group of individuals capable of interbreeding and producing viable offspring – Speciation: formation of a new species 3 processes of speciation: 1.Allopatric Speciation 2.Sympatric Speciation 3.Adaptive Radiation 3 Processes of Speciation 1. Allopatric Speciation – Geographic barrier prevents interbreeding between members of the population – Barrier leads to reproductive isolation – Allele frequencies change (due to natural selection, genetic drift, or mutation – If gene pool changes too much, interbreeding between populations will no longer be possible= NEW SPECIES 3 Processes of Speciation 2. Sympatric Speciation – New species formed WITHOUT geographic barrier – Due to: • reproductive isolation • Behavioral isolation • Temporal isolation Allopatric speciation • speciation occurred in different regions • Key: geographical separation • Geographical isolation Sympatric speciation • one population of one species became two species while in the same geographic region with no physical separation • (temporal isolation, reproductive isolation, behavioral isolation) 3 Processes of Speciation 3. Adaptive Radiation – Rapid evolution of MANY species from a single, common ancestor – Occurs when ancestral species is introduced to an area where diverse GEOGRAPHIC or ECOLOGICAL conditions are available for colonization – Creates VARIANTS of ancestral species that are SPECIALIZED for particular environments 3. Adaptive Radiation Adaptive Radiation • Where species all deriving from a common ancestor have over time successfully adapted to their environment via natural selection Niche • Derived from Latin word meaning “nest” • The role an organism plays in its community or ecosystem • No two species can occupy the same niche in the same area at the same time • Think of a sports team…everyone has a specific role on the field – You can’t have two people playing short stop at the same time Patterns of Evolution Four Patterns Characterize the Evolution of Species 1. 2. 3. 4. Divergent Evolution Convergent Evolution Parallel Evolution Coevolution • Organisms from the same ancestor have developed different anatomies/behaviors (and eventually form new species) • Environment influences their new anatomy/behaviors 1. Divergent Evolution 2. Convergent evolution • organisms that descended from different ancestors • Evolve similar anatomies and/ or behaviors if they live in similar environments 3. Parallel Evolution • Two related species (from same lineage) make similar evolutionary changes after their DIVERGENCE from a common ancestor – Marsupial mammals and placental mammals – Independently evolved similar adaptations when ancestors encountered similar environments 4. Coevolution • Evolution of one species in response to the evolutionary adaptations made by another species • Examples – Predator and prey adaptations – Plants and insect adaptations – Pathogens and immune systems of animals Summary of Evolution • DNA is the blue-print for building ALL living things on planet Earth. • DNA sequences are changed by: • • • • • • • random mutations Radiation Viruses Chemicals sexual reproduction Migration geological events • As a result there is variation within populations of a species. • Those individuals of a particular species with a phenotype (as a result of their genotype) that is more fit to survive in a given environment has a better chance to reproduce. • Those individuals who reproduce more pass-on their genes at a higher frequency than those who do not. • EVERYTHING is trying to SURVIVE and REPRODUCE