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AP Environmental Science Mr. Grant Lesson 31 Ecological Communities © 2011 Pearson Education, Inc. Objectives: • Define the term trophic level. • Characterize feeding relationships and energy flow, using them to construct trophic levels and food webs. • Distinguish characteristics of a keystone species. • Characterize disturbance, succession, and notions of community change. • Perceive and predict the potential impacts of invasive species in communities. • Explain the goals and the methods of restoration ecology. © 2011 Pearson Education, Inc. Define the term trophic level. Trophic Level: Rank in the feeding hierarchy of a food chain. Organisms at higher trophic levels consume those at lower trophic levels. © 2011 Pearson Education, Inc. Characterize feeding relationships and energy flow, using them to construct trophic levels and food webs. • Energy is transferred among trophic levels in food chains. • Lower trophic levels generally contain more energy, biomass, and individuals. • Food webs illustrate feeding relationships and energy flow among species in a community. © 2011 Pearson Education, Inc. Ecological Communities Community = an assemblage of populations of organisms living in the same place at the same time Members interact with each other Interactions determine the structure, function, and species composition of the community Community ecologists are people interested in: Which species coexist, and how species relate to one another How communities change, and why patterns exist © 2014 Pearson Education, Inc. Energy passes among trophic levels One of the most important species interactions involves who eats whom. Matter and energy move through the community Trophic level = rank in the feeding hierarchy Producers (autotrophs) Consumers Detritivores and decomposers © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Producers Producers, or autotrophs (“self-feeders”) = organisms that capture solar energy for photosynthesis to produce sugars Green plants Cyanobacteria Algae Chemosynthetic bacteria use the geothermal energy in hot springs or deep-sea vents to produce their food © 2014 Pearson Education, Inc. Consumers Primary consumers = second trophic level Organisms that consume producers Herbivores consume plants Deer, grasshoppers Secondary consumers = third trophic level Organisms that prey on primary consumers Carnivores consume meat Wolves, rodents (that eat insects) © 2014 Pearson Education, Inc. Consumers Tertiary consumers = fourth trophic level Predators at the highest trophic level Consume secondary consumers Are also carnivores Hawks, owls Omnivores = consumers that eat both producers (plants) and consumers (animals) © 2014 Pearson Education, Inc. Detritivores and decomposers Organisms that consume nonliving organic matter enrich soils and/or recycle nutrients found in dead organisms Detritivores = scavenge waste products or dead bodies Millipedes, soil insects Decomposers = break down leaf litter and other nonliving material into simpler chemicals that can be used by plants Fungi, bacteria Enhance topsoil and recycle nutrients © 2014 Pearson Education, Inc. Energy, biomass, and numbers decrease at higher trophic levels Most energy organisms use is lost as waste heat through cellular respiration Less and less energy is available in each successive trophic level Each level contains only about 10% of the energy of the trophic level below it There are also far fewer organisms and less biomass (mass of living matter) at each higher trophic level A human vegetarian’s ecological footprint is smaller than a meat-eater’s footprint © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Food webs show feeding relationships and energy flow Food chain = a linear series of feeding relationships Food web = a visual map of feeding relationships and energy flow Includes many different organisms at all various levels Greatly simplified; leaves out most species Contains many food chains © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Distinguish characteristics of a keystone species. • Keystone species exert impacts on communities that are far out of proportion to their abundance. • Top predators are frequently considered keystone species, but other types of organisms also exert strong effect on communities. © 2011 Pearson Education, Inc. Some organisms play outsized roles in communities Community dynamics are complex Species interactions differ in strength Keystone species = a species that has a strong or wide-reaching impact far out of proportion to its abundance Removal of a keystone species has substantial ripple effects Significantly alters the food web © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Some organisms play outsized roles in communities Trophic cascade = a phenomenon in which predators at high trophic levels indirectly affect populations at low trophic levels Predators keep species at intermediate trophic levels in check, allowing growth of species at a lower level Extermination of wolves led to increased deer populations, which overgrazed vegetation and changed forest structure “Ecosystem engineers” physically modify the environment Beaver dams, prairie dogs, ants, zebra mussels © 2014 Pearson Education, Inc. Characterize disturbance, succession, and notions of community change. • Disturbances are varied, and communities respond to disturbance in different ways. • Succession describes a typical pattern of community change through time. • Primary succession begins with an area devoid of life. Secondary succession begins with an area that has been severely disturbed but where remnants of the original community remain. • Ecologist today view succession as being less predictable and deterministic than they did in the past. • If a disturbance is severe enough, communities may undergo phase shifts involving irreversible change – or novel communities may form. © 2011 Pearson Education, Inc. Communities respond to disturbance in various ways Disturbance = event that causes rapid changes in the environment that alters the community or ecosystem Tree falling, opening space in a forest Removal of keystone species, spread of invasive species Natural disturbances like tornadoes, hurricanes Human impacts cause major community changes Some species become adapted to disturbance Plants growing in fire-prone regions may have seeds that require fire to germinate © 2014 Pearson Education, Inc. Communities respond to disturbance in various ways Communities have different ways of maintaining themselves when disturbed Resistance = when a community of organisms resists change and remains stable despite the disturbance Resilience = when a community changes in response to a disturbance, but later returns to its original state A disturbed community may never return to its original state © 2014 Pearson Education, Inc. Succession follows severe disturbance Succession = the predictable series of changes in a community following a disturbance Primary succession = disturbance removes all vegetation and soil life Community is built from scratch Glaciers, drying lakes, volcanic lava Pioneer species = the first species to arrive in a primary succession area (e.g., lichens, mosses) © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Succession follows severe disturbance Secondary succession = a disturbance dramatically alters, but does not destroy, all local organisms The remaining organisms and soil form “building blocks” that help shape the process of succession Fires, hurricanes, farming, logging Climax community = community that remains in place with few changes until another disturbance restarts succession © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Communities may undergo shifts The dynamics of community change are more variable and less predictable than thought Competition may inhibit progression to another stage Chance factors also affect movement between stages Climax community may not be based on climate alone Phase (regime) shift = occurs when the overall character of the community fundamentally changes Some crucial threshold is passed, a keystone species is lost, or an exotic species invades Human activity may be creating novel or no-analog communities that are new mixtures of species that have not been seen before in nature © 2014 Pearson Education, Inc. Perceive and predict the potential impacts of invasive species in communities. • People have introduced countless species to new areas. Some of these no-native species may become invasive if they do not encounter limiting factors on their population growth. • Invasive species such as the zebra mussel have altered the composition, structure, and function of communities. • We can respond to invasive species with prevention, control, and education measures. © 2011 Pearson Education, Inc. Invasive species pose new threats to community stability Invasive species = non-native (exotic) organisms that spread widely and become dominant in a community Introduced species = species that were deliberately or accidentally brought from elsewhere Growth-limiting factors (predators, disease, competitors, etc.) are removed or absent Have major ecological effects Chestnut blight from Asia wiped out American chestnut trees Some species help people (e.g., European honeybees) © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc. Zebra Mussels… There are two sides to the story. Eutrophication has plagued the Great Lakes for decades. But by filtering phytoplankton and other suspended material from the water column, the nonnative zebra mussel has helped clean up Lake Erie and other parts of the Great Lakes, the Hudson River, and many more aquatic environments. These mussels are much more efficient at filtration than their native counterparts. Many birds feed on them, and the mussels’ excrement provides habitat for a food chain anchoring a great diversity of species. Biologists credit the zebra mussel with restoring native grasses and fishes. Were it native, the zebra mussel would be hailed as a savior, not reviled as a scourge. EXCERPT FROM Whole Earth Discipline Stewart Brand Category: Nature We can respond to invasive species through control, eradication, or prevention Techniques to control invasive species Removing them manually Applying toxic chemicals Introducing native predators or diseases Stressing them with heat, sound, electricity, carbon dioxide, or ultraviolet light Control and eradication are hard and expensive Understanding the biology of invasive species can help predict where they will be a problem Prevention, rather than control, is the best policy © 2014 Pearson Education, Inc. Explain the goals and the methods of restoration ecology. • Restoration ecology is the science of restoring communities to a previous, more functional or more “natural” condition, variously defined as before human or before recent industrial impact. • The growing practice of restoration ecology, informed by the science of restoration ecology, helps us restore ecological systems. © 2011 Pearson Education, Inc. Altered communities can be restored Humans have dramatically changed ecological systems Severely degraded systems cease to function Ecological restoration = efforts to restore communities Restoration is informed by restoration ecology = the science of restoring an area to an earlier condition to restore the system’s functionality (e.g., filtering of water by a wetland) It is difficult, time-consuming, and expensive It is best to protect natural systems from degradation in the first place © 2014 Pearson Education, Inc. Altered communities can be restored Prairie restoration involves replanting native species, controlling invasive species Nearly all tallgrass prairie was converted to agriculture by the 19th century The world’s largest project is the Florida Everglades Flood control and irrigation removed water Populations of wading birds dropped 90–95% It will take 30 years and billions of dollars to restore natural water flow © 2014 Pearson Education, Inc. © 2014 Pearson Education, Inc.