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Plant structure, growth and development Chapter 35 Plant Tissues • There are three categories of plant tissues – Dermal tissues • form the outer covering of the plant • serve a protective function • herbaceous plants – epidermis • Woody plant – periderm (outer bark) – Vascular Tissues • Function to transport materials through the plant body and provides support – Xylem – transport of water and dissolved minerals – Phloem – transport of dissolved sugars • Arranged into vascular cylinder in the root or vascular bundles in stems and leaves – Ground Tissues • • • • Neither dermal or vascular Pith – internal ground tissue (in vascular cylinder or stem) Cortex – ground tissue located towards the outside of the plants organs Functions for storage, photosynthesis Cell types composing plant tissues • Simple tissues – Collenchyma • Cell walls of irregular thickness • Flexible support – Parenchyma • Cell walls very thin to facilitate movement of particles through cell • Storage, photosynthesis – Sclerenchyma • Very thick cell walls (primary and secondary cell walls made of lignin) • Dead at maturity • Sclerenchyma fibers and sclerids • Rigid support Cell types composing plant tissues • Complex tissues – Xylem • Tracheids and vessels – Modified sclerenchyma fibers – Dead at maturity (hollow) – Phloem • Sieve tubes and companion cells • Companion cells alive • Sieve tubes dead at maturity (sclerenchyma fibers) Plant organs • Plant organs may be divided into two systems – Shoot system • Above ground (usually) • Stem • Leaf • Flower – Root system • Taproot (primary) • Lateral roots (secondary) • Adventitious roots (arise from stem) Growth of plant parts • Plant exhibit localized growth at meristems – Apical meristems – produce primary tissues and increase the length of plant parts • Found at the tips of roots (root apical meristem) and in buds on stems (terminal buds and axillary buds) – Lateral meristems are found in woody plants and increase the girth of plant parts • Vascular cambium – produces secondary xylem and phloem • Cork cambium – produces cork cells that will form periderm (outer bark) The root system • Functions of the root – – – – – Absorption of water and nutrients Anchorage Storage Soil erosion Other root modifications: Root modifications • Prop roots – Adventitious • • • • Storage roots Aerial roots Buttress roots Pneumatophores – Exchange gases Root anatomy • Zone of cell division – Root apical meristem – Protected by root cap • • Zone of elongation Zone of maturation – Contains mature tissues – Root hairs • Cytoplasmic extension of epidermal cells • Increase surface area • Layers of typical root – Epidermis – Cortex – Vascular cylinder • • • • Be sure to know the difference between monocot and eudicot roots Epidermis Cortex Vascular cylinder – – – – Endodermis Pericycle Pith (in monocots) Xylem/phloem Development of lateral (secondary) roots from pericycle • Notice that the vascular cylinder of the original root and the lateral root are connected The stem • Part of shoot system • Functions for support of leaves and flowers • Modifications allow for other functions as well – – – – Bulbs Stolons Rhizomes tubers Stem anatomy (external) • • • • • • Node Internode Axil Axillary bud Terminal bud Bud scale – Modified leaf to protect apical meristem • • Terminal bud scar Leaf scar Stem anatomy (internal) • Be sure to note differences between monocots and eudicots Secondary growth to produce woody tissues • • All woody plants exhibit primary growth to produce new shoots (herbaceous) Secondary growth at lateral meristems is responsible for the production of woody tissues and increases the girth of plant – Vascular cambium – Cork cambium • Wood (secondary xylem) – Heartwood vs. sapwood • Cork cells form periderm outer bark • Secondary phloem forms inner bark Secondary growth to produce woody tissues The leaf • Primary function of photosynthesis • Form = function – CO2 + H2O Æ C6H12O6 + O2 – Other function of leaves (leaf modifications) • • • • • Tendrils Spines Storage Bracts Reproductive leaves (asexual) Leaf anatomy (external) • • • • Axil Petiole Blade veins Leaf types • Simple – one blade per leaf • Compound – blade divided into leaflets – Palmate – Pinnate (once –pinnate and twice pinnate) Leaf anatomy (internal) Epidermis cuticle guard cells stoma Mesophyll palisade spongy Vein bundle sheath xylem phloem