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Taxonomy Notes Nowicki Biology, Chapter 17 With a partner discuss why/how we classify things: 17.1: The Linnaean System of Classification Linnaeus developed the scientific naming system still used today taxonomy – science of naming and classifying organisms - hierarchical – multilevel scale in which each level is contained or included in every level above it - based on the system developed by Carolus Linnaeus Figure 1: Animal Hierarchy Looking at the Figure 1: Animal Hierarchy to the right: 1. Dogs are _____________ and ____________________ 2. Are birds animals? ______________ 3. Are fish mammals? ___________ Animals Mammals Dogs Fish Birds Cats taxon – a level of classification; the basic level is species; taxa is the plural - the diagram below shows 8 taxa in the hierarchical system of classification we use most general/least specific (includes the greatest number of organisms) Domain Kingdom Phylum Class Order Family Genus most specific – contains only one single kind of organism Species Mnemonic (Dear King Phillip Came Over For Grape Soda) Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species Come up with your own mnemonic: 1 (Domain is not shown in the diagram to the left) When looking at the taxonomic hierarchy (Domain, Kingdom, etc), Which taxon has the most diverse group of organisms? __________________________ Which taxon is the least diverse? ___________________________ Scientific Naming - Using common names is confusing because many organisms may have several different common names. - The cougar is also known as the mountain lion, puma or catamount, thus the need for a scientific name. - We use binomial nomenclature to assign scientific names. The scientific name for a cougar is Felis concolor binomial nomenclature – a system of naming in which each species has a two-part scientific name - Two different species cannot have the same scientific name - Importance? Binomial nomenclature provides a standard of communication among biologists, regardless of their native language scientific name – genus and species names of an organism - basis for our system of naming organisms. - two parts: o genus – first part/word of the scientific name always capitalized and italicized (typing) or underlined (handwriting) can be used alone o species – second part/word of the scientific name never capitalized always italicized (typing) or underlined (handwriting) can never be used alone Helix pomatia Typed Written 2 EXAMPLE: One of the most commonly used scientific names is Escherichia coli. This is the scientific name of a rod-shaped bacteria that lives in your intestines, helping you digest your food and produce vitamin K (necessary for blood clotting). It is also used in research involving bacteria and is sometimes linked to outbreaks of food-poisoning. When the name has been typed, it should look like this: Escherichia coli When the name has been handwritten, it should look like this: Escherichia coli It is okay to abbreviate the full name like this: E. coli or Escherichia, but never like this: coli Why are all scientific names so weird? Why can’t they just be “in English”? Scientific names are written in Latin because it is not a language spoken in any country today; therefore, it is a nonpolitical choice. There are several divisions of the International Code of Nomenclature that govern the naming of all organisms. The Linnaean classification system has limitations - Linnaeus based his classification on physical similarities - today we have technology that allows us to look at molecular similarities (amino acid sequences in proteins that are based on nucleotide sequences in the genes of the DNA) - remember that organisms can look similar due to analogous characteristics resulting from convergent evolution (think back to evolution: bird wings and butterfly wings have the same function, different structure) - classification should be based on homologous characteristics resulting from divergent evolution (think back to evolution: homologous characteristics share the same structure, have a common ancestor but have a different function) - molecular homology is much more definitive in determining species relationships than physical analogy The more taxonomic levels that two organisms share, the more closely related they are. Which bears are most closely related? 3 Ursus maritimus Ursus arctos Ailuropoda melanoleuca Using the chart to the right 1. What type of animal is Musca domestica? 2. From the table, which 2 animals are most closely related? 3. At what classification level does the evolutionary relationship between cats and wolves diverge (become different)? dichotomous key – tool used by scientists to identify organisms in the field based on defining characteristics Dichotomous Key for Norns - arranged in pairs of 1. a. Has pointed ears……………………………………………………. go to 3 questions to which the b. Has rounded ears………………………………………………..... go to 2 answer is generally yes 2. a. Has no tail……………………………………………………...kentuckyus or no, followed by a b. Has tail……………………………………………………………...… dakotus direction of what pair of 3. a. Ears point upward……………………………………………....... go to 5 questions to go to next b. Ears point downward…………………………………………....… go to 4 4. a. Engages in waving behavior………………………………….... dallus until the organism has b. Has hairy tufts on ears………………………………………californius been identified - key to using this tool is always starting with the first pair of questions! Norns are a very rare and actually imaginary animals that belong to the genus Norno. There are 8 different species of Norno, generally located in specific regions of the world. Use the key to identify the Norns shown. 5. a. Engages in waving behavior………………………………. walawala b. Does not engage in waving behavior…………………....... go to 6 6. a. Has hair on head…………………………………………………. beverlus b. Has no hair on head (may have ear tufts)………………. go to 7 7. a. Has a tail……………………………………………………………….... yorkio b. Has no tail, aggressive……………………………………………….rajus A B C 17.2: Classificaiton Based on Evolutionary Relationships 4 phylogeny – evolutionary history for a group of species - can be shown as branching “family trees” - think back to evolution and phylogenetic trees: does this reflect convergent or divergent evolution? _________________________________ Cladistics is classification based on common ancestry cladistics – classification based on common ancestry - goal is to place species in the order in which they descended from a common ancestor cladogram – diagram based on patterns of shared, derived traits that shows the evolutionary relationships between groups of organisms - clade – group of species that share a common ancestor derived characters – traits that can be used to figure out evolutionary relationships among a group of species; traits are shared by some species but are not present in others - the more closely related species are, the more derived characters they will share outgroup – group of species that shares no derived characters with the other groups being studied 1. Which are the derived characters in the cladogram below? 2. Circle the Hagfish and label it, “outgroup.” Explain why it’s the outgroup. Complete Quicklab: Classifying – Construct a Cladogram, p. 511 5 1. Use the following words to appropriately label the 5 blanks at the top of the cladogram. airplane bicycle car motorcycle walking 2. What are the derived characters in this cladogram? 3. What mode of transportation may be considered an “outgroup?” 4. A species that has evolved a new trait is not better than a species without that trait. Each species is just adapted to a certain way of life. When might riding a bike have an advantage over flying in an airplane? Cladograms are important tools in classification because they represent true evolutionary relationships based on all available data, especially molecular homology--DNA &/or amino acids. Interpreting a Clagogram: CLADE – group of organisms that share certain traits derived from a common ancestor. On many cladograms, a clade is marked by lines looking like the letter V. The dotted line marks the amniote clade shown in this diagram DERIVED CHARACTERS – shared trait of all species above the name of the character or the hash mark for it NODE – intersection of 2 branch points representing the most recent common ancestor What derived traits do groupers have? _______________________________________________ Do Lancelet’s contain a vertebral column? _______________ What organism is closely related to the wolf? _________________________ 6 17.3: Molecular Clocks Molecular clocks use mutations to estimate evolutionary time molecular clocks – models that use mutation rates to measure evolutionary time - basis for using molecular data to determine relatedness of 2 organisms o more differences between the genes or proteins of 2 organisms means they are more distantly related they are (they shared a common ancestor a long, long time ago) o fewer differences between the genes or proteins of 2 organisms means they are more closely related (they shared a common ancestor more recently) (Read pages 516-518 for more information on Molecular Clocks) 17.4: Domains and Kingdoms Classification is always a work in progress: Two Kingdoms Three Kingdoms Five Kingdoms Four Kingdoms Six Kingdoms Current system of 3 domains and 6 kingdoms The three domains in the tree of life are Bacteria, Archaea, and Eukarya Bacteria – includes single-celled prokaryotes in the kingdom Bacteria - cell wall contains peptidoglycan - includes bacteria that live in/on organisms, soil decomposers, disease causing bacteria (everyday bacteria) Archaea – includes single-celled prokaryotes in the kingdom Archaea - cell wall does not contain peptidoglycan, has lipids - includes bacteria that live in extreme environments such as deep sea vents, hot geysers, salt lakes, etc. classifying Bacteria and Archaea is difficult because they so frequently transfer genes among themselves that defining a “species” is nearly impossible; this process is just beginning and will continue as we learn more about these organisms Eukarya – made up of all organisms with eukaryotic cells – distinct nucleus and membrane-bound organelles - can be single-cells, colonial, or multicellular - includes the kingdoms Protista, Plantae, Fungi, and Animalia 7 Fill in this graphic organizer for the 3 domains and the 6 kingdoms under each. 8 CHARACTERISTICS OF DOMAINS AND KINGDOMS DOMAIN KINGDOM Archaea Archaea Bacteria Bacteria CELL TYPE Prokaryotes BODY FORMS unicellular Prokaryotes unicellular CELL WALL cell walls with peptidoglycan heterotroph & autotroph everyday bacteria, includes decomposers and Nitrogen fixers E.coli, Strep, Staph NUTRITION FACTS EXAMPLES cell walls with lipids heterotroph & autotroph live in extreme environments thermophiles, halophiles Eukarya Plantae Protista Fungi Eukaryotes mostly unicellular some have cell walls heterotroph & autotroph live in water/moist places; produce about 1/3 of Earth’s oxygen algae, amoeba, paramecium Eukaryotes mostly multicellular cell walls with chitin heterotroph only decomposers; making food and medicine Eukaryotes multicellular mushrooms, yeast, mold grass, trees, shrubs Animalia Eukaryotes multicellular cell walls with NO cell walls cellulose autotroph only heterotroph only produce about most 2/3 of Earth’s physically oxygen diverse kingdom sponges, worms, insects, fish, birds, reptiles, mammals 9