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The competitive exclusion principle Two species with similar needs for the same limiting resources cannot coexist in the same place for very long. OR No two species can occupy the same niche in the same community indefinitely. Gause’s Experiment Biomass Separately 200 150 100 50 Mixed population Biomass 4 8 12 16 20 24 Paramecium aurelia Separately 200 150 100 50 Mixed population 4 8 12 16 20 24 Paramecium caudatum Days Reduced niche overlap can allow coexistence • Behavioral Acclimation Green anole lizard forced to move to branches by introduction of Brown lizard. • Character Displacement Through evolution, 2 closely related species tend to be more distinct when they are sympatric than when they are allopatric. 1 Examples of Character Displacement Bill Length (mm) Species 2 Overlap 29 26 Species 1 23 Gr ee Ira ce Ch n in a Two species of Asian nuthatches of the same genera. Darwin’s Finches Bill Depth % in each class Geospiza fulginosa Geospiza fortis 40 20 same island 40 20 alone 40 20 alone 8 10 12 14 Bill Depth (mm) Communities Can Recover From Disturbance Succession The sequential change in the species composition of a community over (ecological) time - 100s to 1000s of years. The change in plant species is most often considered 2 There are 2 types of succession • Primary (1o) - Community development in an essentially lifeless area with no soil remaining from a previous community. Example - rock exposed by glaciers => lichens => mosses => grasses => shrubs => trees • Secondary (2o) - Community development following the disturbance of a previously existing community. Remnant soil exists. Example - abandoned farm => crabgrass => horseweed => perennial weeds => pine trees => hardwoods. Primary Succession Secondary Succession 0 years 30 years 7 years 95 years 15 years 150 years 3 Communities Don’t Always Recover From Disturbance Clear Cuts That Have Not Been Successfully Reforested 4