Download Morphological remarks The pollen Geographical remarks

A revision of the Ochnaceae
of
the Indo-Pacific area
Kanis
A.
Contents
Summary
1
Introduction
Subdivision
2
and interrelations
Historical
3
3
survey
subdivision
Proposed
5
remarks
Morphological
8
The
vegetative parts
The
inflorescence
8
8
The flower
Fruit
10
and seed
The
12
characters
Anatomical
12
pollen
Geographical
13
remarks
15
of data
15
Summary
Discussion
Remarks
on
17
the
presentation
18
of data
Ochnaceae
Family
20
Subfamily
1.
Ochnoideae
21
Tribe
1.
Ochneae
22
Subfamily
2.
Tribe
2.
Tribe
Excluded
3.
61
Sauvagesoideae
62
Euthemideae
66
Sauvagesieae
80
taxa
80
Index
Summary
This taxonomic
the Pacific
10
genera
been
of
some
of
a
A short
being
the
distinct section
history
Rchb.,
Lophireae
and
Rchb. The
names
is
taxonomy
the
Ochneae
are
with
Schreb.
for the
given
Sauvagesoideae
Asia, Malesia, Australia,
synonyms
Brackenridgea palustris
works:
previous
most
are
ssp.
listed.
A
kjellbergii
20
few
and
in
species
have
taxa
Kanis is
new.
have been overlooked in the past and that the traditional concepts
(v. Tiegh.)
Two subfamilies
specific
199
Southeast
than in
applied
whereas
accordance
Gomphia
correct
is
among which
names
Notochnella
of the
subsequently proposed.
Elvasieae
in
not been
from South and
concept
treated,
level
specific
of Ochna L. and
Kanis
(Gaertn.)
old
species
area
specific
some
have
species
species
wider
for the
infra
at
includes all Ochnaceae
much
accepted
shown that
Asiatic
type
are
accepted
It is
revision
Islands. A
are
and
original
respective
type
Kanis in the
a
partly
species.
genus
names,
currently
new
suprageneric
the tribes
in the
O.
used
L. and G.
Blanco is made
fascicularis
for
serrata
the
Brackenridgea A. Gray.
the Ochnoideae
including
subdivided
All
concepts.
longer accepted here, O. jabotapita
recognised:
Lindl.
newly
the
are no
subdivision
comprising
Sauvagesieae,
subtribes
Ochninae
of the
Euthemideae
and
Ochnaceae is
the tribes Ochneae
Planch.,
Ouratinae
and
and
(v. Tiegh.)
*
Kanis,
and the
Some
Sauvagesieae
general
characters
of the
remarks
pollen.
in the
are
An
subtribes
made
attempt
about
is made towards
It is assumed that the genera Ochna and
east
or
1
Asia, Malesia, Australia,
less relict-like
nature
and
of their
Sauvagesinae
a
Luxemburginae (Planch.)
characters in
better
are
the
understanding
Gomphia migrated from
the Pacific Islands
areas.
and
morphological
regarded
Kanis.
family, including
some
new
of the inflorescence types.
Africa into Asia. Other genera in Southas
long established,
because
of the
more
BLUMEA
2
VOL.
No.
XVI,
1968
i,
Introduction
The revision presented here
started
was
as
it
proved
soon
the Malesian
Certain
of the latter
genera
Malesia, which made comparitive studies
in
the
on
subfamily
in related
contribution
a
to
subfamily Sauvagesoideae,
extend the studies in the Ochnoideae
to
necessary
area.
for
preliminary study
a
the Flora Malesiana. After completion of a first manuscript
over
the boundaries of
only poorly represented
are
species from adjacent
areas
highly
desirable.
but
second,
A
less
not
motive for this revision
important
the situation that
was
critical monographic work including the Ochnaceae of the Indo-Pacific
after those
by
De Candolle
merit in
has
(1902 —07)
accepted because
the Pacific
The
area
of his
(1811) andPlanchon (1846 —47). The work of
certain aspects,
unusually
could only give
treated
area
covers
names
his
but
narrow
taxonomic
to
Hainan)
remote
most
of this
genera
The
island
local traditions.
family do
western
not occur
Old World
genera
are
on
the
means
that the natural
Kwangsi, and
boundary,
eastern
as
it is the
whereas neotropical
found,
is
practical
one
Mascarenes
African and
mainly
genus
chosen for
are
Africa, Madagascar, and the
treated here
This
in the Pacific Islands.
limits of this revision
between the species in India and Ceylon
Peninsula,
Islands.
(NE. Queensland). Fiji marks the
an
There is
reasons.
a
wide
easily have doubled the
African
on
the other. However,
there is
apparently
time
material,
a
rather close
two
they
are
Some
added
the subject of
brief, general
as
an
attempt
with special reference
completely
from
on
chapters
to
the
literature.
I
often than their
a
on
taxonomy,
the
taxa
treated. For
hope that
this
data
will
geography have been
and
morphology,
of the family
better understanding
on
other
facilitate
as
whole, though
a
I had
taxa
comparisons
to
rely
almost
with Ochnaceae
the African region and the Neotropics.
The completion of this paper
a
could
relatives and presently
eastern
was
made
possible
by
Netherlands
grant from the
a
Foundation for the Advancement of Tropical Research (WOTRO) and by
for
I have
Besides,
paper.
the
less parallel studies by other botanists.
more or
towards
more
of
relationship
studies
thorough comparitive
for the preparation of this
necessary
African Ochnaceae have been revised
since
gap
hand and those of the SW. Arabian
between certain species in continental Asia and others in Africa and Madagascar.
refrained from analysing
been
Australia, and
in Asia,
reached in the North (Himalayan Mts.,
where
group
Tieghem
not
and subtropical parts of South and Southeast
continental Asia, Malesia, Australia, and the Pacific
are
Van
no
written
was
has
treatment
concept. Floras
species
according
the tropical
boundaries of the Ochnaceae
and the South
area
study
tour
to
an
herbaria in London and Paris. This substantial support is
allowance
gratefully
acknowledged here.
I
am very
grateful
who encouraged
my
studies
to
Dr. C. G. G.
assistant, later
the
to
my
Professor Dr. H. J. Lam, formerly director of the Rijksherbarium,
first steps
on
the path of plant taxonomy and advised
this regional revision. I
J.
van
as
a
am very
Steenis, who made it possible
a
for
worker. He kindly provided
guest
Rijksherbarium
and
me to
me
to
his
thank both scientists also
me
successor,
to
extend
Professor
realize this plan, first
as
his
with all possible facilities from
the Flora Malesiana Foundation and he spent
valuablt time in reading and discussing
to
much indebted
much of his
the final draft of the manuscript. I should like
for their major contributions
to
my
general education
as
botanist.
Dr.
R.
Dr. P. W.
C. Bakhuizen
van
Leenhouts, and
den Brink
Dr. H.
Jr.,
Sleumer,
Dr.
W.
A.
van
Heel,
staff members of the
Dr. C.
Kalkman,
Rijksherbarium,
gave
A.
Kanis:
miscellaneous advices
1
am
during
greatly indebted
who made
revision
A
to
my
of
Ochnaceae
for which I
work,
of
assistance
Gratefully acknowledged
typed
typed
of the final
most
want
the
types in
pollen
treated here, kindly putting his results
taxa
Indo-Pacific
the
of
the
manuscript and compiled
for this
is
the institutes marked with
asterisk*.
Arnold
BKF*
The Forest
BM*
British Museum
BO
Herbarium
BRI
Botanic Museum and
C
Botanical
CAL
Central National
DD
Forest
E
Herbarium of the
FI
Herbarium Universitatis
Herbarium, Bangkok,
Herbarium, Copenhagen,
Botanic
GL
Department
K*
Herbarium
KEP*
Forest
KYO
Department of Botany, Kyoto University, Japan.
L*
Rijksherbarium, Leiden,
P*
Museum National
S
Naturhistoriska
SAN*
Forest
SAR*
Sarawak Museum
SING*
Herbarium
U*
Botanical Museum and
Jardin Botaniques, Geneva,
of Harvard
Botany, University
Botanic
of Glasgow,
first
Scotland,
U.S.A.
U.K.
Gardens, Kew, England, U.K.
Netherlands.
d'Histoire
Naturelle, Paris,
France.
Riksmuseum, Stockholm, Sweden.
Department, Sandakan, Sabah, Malaysia.
and Forest
of the Botanic
Department, Kuching, Sarawak, Malaysia.
Gardens, Singapore.
Netherlands.
Herbarium, Utrecht,
AND
INTERRELATIONS
survey
The Ochnaceae
systems.
U.K.
Institute, Kepong, Malaya, Malaysia.
Research
were
lasted until 1895 that
not
Australia.
Denmark.
University, Cambridge, Mass.,
SUBDIVISION
It is
personally visited
Switzerland.
Gray
Historical
I have
U.K.
Garden, Edinburgh, Scotland,
Conservatoire
Royal
examination of their
permitted
Florentinae, Florence, Italy.
G
of the
Vijsma, who
India.
GH
of
wife, who also
Marks and Mr. E.
India.
Herbarium, Howrah, Calcutta,
et
Nieuwkoop, who
my
Indonesia.
Institute, Dehra Dun,
Herbarium
a
first described
more
as
family by
a
A.P.
de Candolle in
unanimously accepted circumscription
surprising, however, that this family
The androecium and
especially
sight remarkable differences rather
the
than
was
not
recognized
gynoecium
a
in the
as
was
fundamentally, whereas anatomical
and
\ but it
1811
given by Gilg.
such in early natural
respective tribes
general resemblance. Nonetheless,
be demonstrated that the flowers of the Ochnaceae have much in
eventually
of the
U.S.A.
Herbarium, Brisbane, Qld.,
Royal
especially
Thailand.
(Natural History), London, England,
Bogoriense, Bogor,
Research
van
from
gratefully acknowledged here.
Arboretum, Cambridge, Mass.,
Museum and
Ochnaceae,
index,
The directors of the institutes mentioned below kindly
A
much.
paper.
collections. Their cooperation
an
very
disposal.
at my
of it in different stages, and from Mr. C. L.
part
prepared the drawings
3
the Rijksherbarium,
at
received from Miss E.
was
Area
thank them
to
J. Muller, presently palynologist
Mr.
preliminary inventory
a
the
palynological evidence
also indicates
show
at
it could
common
true
inter-
relationships.
In De Candolle's concept, the
the subfamily
as
distinguished later
andSimaroubaceae
accepted
l
)
on as a
was
under
to
the
separate
as
publications in
family,
family
Initially
a
after
subtribe of
this
sometimes
Ochnaceae
(1811)
chapter
are
comprised
he added
family (1824).
accepted long
the Ochnoideae
References
mostly
Ochnoideae.
A close
a
relationship between
Ochnaceae
by several authors. Reichenbach (1841)
a
tribe Simaroubeae in
given more extensively
under relevant
about what is treated here
tribe Simaroubeae which he
lower
taxa.
in
the family
the taxonomic
even
Rutaceae.
enumeration,
BLUMEA
4
According
later opinions,
to
this
VOL.
based
was
No.
XVI,
1968
i,
rather than
analogy
on
on
of
homology
certain floral characters.
The
L.
genus Sauvagesia
tribe of the
valved capsule
with
De Candolle
regarded by
was
of its
Violaceae, because
(1824)
1-celled
3-carpelled,
view
several authors
supported by
was
to
developing
seeds. However, in Sauvagesia the capsules
many
the margins of the carpels, whereas in the Violaceae the fruits
medians. His
belonging
as
ovary,
are
like Meisner
a
distinct
into
the
tribe Sauvagesiae
sometimes also
including
(1836),
Bentham
St. Hil. and
Luxemburgia
the
dehiscing along
Hooker (1862), Le Maout &Decaisne (1868), and Baillon (1873). They added other
to
3-
a
opening along
are
&
genera
allies.
its
Other authors like Bartling (i83o),Martius (1835), and Endlicher (1840) rather followed
Dumortier
several
later
a
as
The
described a separate family
(1829), who
of
versions
subfamily
in the Violaceae
be related
to
to
(1836), and finally
the Ochnaceae
to
the Ochnaceae
(1846). Lindley
referred
Sauvagesiaceae
(1853).
(1846 —47)
He
named Gomphieae.
tribe
it
instituted
two
at
Bentham
a
a
right (1846).
affinities. It
was
and others.
(1845), but transferred
it later
later
to
to
the
world revision of the Ochnaceae. He
as
belonging
the Luxemburgieae
to
one
tribe, incorrectly
including
neotropical
some
included in the latter tribe, although
were not
the
accepted Planchon's
Tetramerista
(1862), incorporating
tribe in
alleged
its
tribe (1830),
a
resemblance with Luxemburgia.
Hooker largely
&
as
own
other tribes in the family, viz. the monogeneric
and
Asia
in its
the Ochnaceae first (1836),
to
write
to
Sauvagesia and its closest relatives
he pointed
a
the Sauvagesiaceae
the first
Euthemideae from SE.
genera.
as
was
Sauvagesieae
family
by Endlicher (1840)
DC.
sensu
Gomphia (Ouratea), and Elvasia
regarded Ochna,
a
as
noteworthy history for
a
Planchon referred it primarily
Planchon
Sauvagesiaceae. Lindley, who published
first treated the
at
system,
Euthemis Jack has also
genus
considered
natural
a
Ochnaceae,
Miq.
whereas
Violaceae. Their system has had
a
concept in
in the Ochneae.
their Genera Plantarum
They accepted the Luxemburgieae
the Sauvagesieae
s.s.
considerable influence
treated
were
on many
as
part
of the
Floras in the late
19th
and early 20th century.
Engler published
in
general
a
outline of the family
and he
separate tribe
a
them
keeping
presence
or
these formal rank,
and
not
modern
1874, in which the
as
a
called them 'series'.
family
They
from
the Ochnaceae,
Several
Van
dealing
papers
and 1907.
Tieghem
both primarily
a
x
)
in
this
It is remarkable
resemblance
of the
"
in the
a
group.
that
two
structure
“Albuminosae
anatomical
on
1
”; later
was
also
the first
Unfortunately
years
earlier
of the ovary
he did
not
the
were
He
grounds.
published by
most recent
ex
Gaertner
and he excluded Tetra-
single tribe, incorrectly applying
with the Ochnaceae
He
on
(1895), accepted
Lophira Banks
Van
united the
the latter name.
Tieghem
between
world revision of the family.
keen observer and he made important discoveries, especially
field of embryology.
blastogeny
into
Together these comprise the
was
groups,
accordingly named Albuminosae’
his 'series'. He transferred
including
system,
two
based
associate this criterion with other characters.
Sauvagesieae and the Luxemburgieae
1901
were
'
were
from the Dipterocarpaceae into the ‘Exalbuminosae’ sensu Engler
merista
into
treated the Ochnaceae in 'Die natiirhchenPflanzenfamilien'
Engler's
Ochnaceae, still
tribe of the
He divided the
absence of albumen in the seed and
Gilg, who
of
the Ochnaceae in
less reflected. He united Elvasia and Tetramerista
the Sauvagesieae
recognized
'
and Exalbuminosae’.
He did
most
on
more or
from the Luxemburgieae.
apart
without giving
treatise
already
was
(Ber.
of
refer
publish
an
observation
in the
concerning
the
he caused much confusion by his unorthodox
Deutsch.
Lophira
to
to
this
Bot.
with the
Ges.
11, 1893,
25)
he had
Sauvagesieae (named by
early opinion which
is here
him
revived.
pointed
to
the
Luxemburgieae)
A. Kanis
ranks. Besides, his
papers
to
Ouratea
neotropical
and
11
as
5
as
genera
deserve
a
There is
taxonomic
Brackenridgea
to
were
5
A.
Tieghem's
sub-
are
and
palaeotropical
into another
as
subtribes
two
forms
mere
tribe,
considered best
are
genera
of his species
many
to
Tieghem
Van
Gray.
13
and the former
genera
Most of Van
raised
the Ochneae, viz. Ochna
with
subtribes
two
shortly
Euthemideae, Luxemburgieae,
genera in
all. Later authors followed
tendency, however,
a
Engler's
of all
taxa
Gilg rather than
re-evaluate Van Tieghem's observations
as
do
or
Van
not
Tieghem.
basis for
a
moderate subdivision of the African Ochneae s.l.
a more
subdivision
Proposed
in this revision,
Though,
five
at
and
into
species, whereas
status
and
subtribe with
one
or
5
small concept of
He united Ochna and Brackenridgea
respectively.
the level of section
at
Area
Indo-Pacific
unusually
Gilg had recognized three
s.s.
respectively.
genera
the
of
whereas the Ochneae and Elvasieae
tribe, subdivided
a
the latter
considering
with
and his
Aubl. (inch Gomphia Schreb.),
Ouratea
22
Ochnaceae
Gilg's Lophireae
family rank,
families of theOchnaceae
treated
the
of
rather scattered and he revised several of his ideas
are
He raised
publication.
and Sauvagesieae
L.,
revision
of nomenclatural problems
treatment
after
A
:
I have been
tribes,
I have studied
tempted
only
to propose
ten
belonging
genera
three of the
to
of the subdivision of the
improvement
an
Ochnaceae.
the first
In
place
a
few nomenclatural
changes
appeared
bring the
to
necessary
of subfamilies, tribes, and subtribes in accordance with the present
Code
names
of Nomen-
clature.
I have
(1874)
into
Exalbuminosae’ and ‘Albuminosae’
distinct subfamilies, named Ochnoideae and Sauvagesoideae respectively,
two
as
'
main division
accepted Engler's
Tetramerista which
was
The monogeneric,
Ochnaceae,
soon
after recognised
as
African tribe Lophireae which
'
referred by
was
albumen. However,
a
Gilg (1895)
to
the
was
a
2-carpelled,
included
not
'
Exalbuminosae
position in the Sauvagesoideae
non-distichous leaves and
omitting
Theaceous.
seems
more
in the
by Engler
because of its lack of
since it shares
logical,
I-celled, many-ovuled
ovary
with
that
sub-
family.
The
system is
proposed
not
distinct from that by
very
in the position of the tribe Lophireae.
The
Ochna L.
genus
Soc. Brot. II, 36,
here,
it
can
affect the
The
be
was
recently subdivided
of the Asiatic
Ouratea Aubl.
genus
Gilg (1925), mainly differing
ij.
into
three sections
possible other delimitation
a
was
taxa
are
subdivided
by Gilg (1895)
identical with Van Tieghem's
sperminae’ respectively.
the
same
taxa
a
Farron
distinction
(Bull.
at
of this
area
will
genus
into
by
me
(1967),
for the
species
should have
(I.e., 1963; Bull. Jard.
Bot.
of the
priority
Brux.
The genusBrackenridgea A. Gray
and
even
as
congeneric
by
subgenera (1925).
Suisse
the
Campylospermum
1965, 393)
was
for the
regarded by
(1875)
and
many
In
my
Tiegh.
same
proposed
for
Ouratea
for
name
typified
name as
as
name
used by Farron
concept.
authors
Ridley (1922).
'subtribe' ‘Plicoseminae’ (Brackenridgea s.l.) first in his
his Ochneae.
v.
These
and ‘Campylo-
1963, 208—212)
73,
correctly reserving
Old World treated here. This
over
33,
Bennett
not
sections, Neoouratea
two
'subtribes' ‘Orthosperminae’
Soc. Bot.
generic level,
as
the species of the New World. I have accepted this opinion, but I have used the
Gomphia Schreb.
(Bol.
treated
in the future.
species
(New World) andPalaeoouratea (Old World), treated later
two
by Robson
1962, 12). Since only the section Ochna is represented in the
safely assumed that
names
(Table
as
Van
related
to
Gomphia
Tieghem placed
his
tribe Ourateae but shortly after in
opinion Brackenridgea is indeedrelated
to
Ochna rather than
to
Ouratea
BLUMEA
6
VOL.
XVI,
No.
I,
1968
in
treated
Hal.f
Gray
Boerl.
A,
Taxa
Schreb.
Aubl.
L.
subtries.
Genus Ochna
1.
the
Jack
Brackenidga Ouratea Gomphia
Euthemis
2.
4.
3.
Vidal
Ii.
Diels
Sauvgesia Sinia
Korth.
sinia
Indo
Neckia
5. 6. 7.
Hil.
Bl.
Gaertn.
ex
St.
Banks
Indovethia Schurmansiel Schurmansi Luxemburgia fc-Lophira
8.
9.
\
as
them.
far to
as
Subtribe
)Kanis
Ouratine(Tiegh.)Kanis
given
complet numbers
Vi
r»Vi
"R
Luxembrginae(Planch.
-
Ochnace ,with
of
subdivson
■RlvaoQA
Tribe
provide
in
v.
Planch,
Engl.)
Luxembrgiea
tribus
tribus
on
on
based
based
PTflnoh.
Rchb.
flpaft
the
Tieghem,
Ourate
'
..
and
Hook.
(non
JO.
'
including
in
Lophirea
"RitVPmi
sta.nov.,
sta. ov.,
Kanis,
Kanis
33.
P.
(v,Tiegh.)(1902)
».
(Planch.)
592.
•_
been
Lindl
..
have
Prop sed
revison
1.
TABLE this
(1846)
Ouratine Bot. Luxembrginae Bot.
16
5
de
J.
Subfamily
Ochnoidea
1.
J.
Subtrius Morot, Subtri us Lond.
1)
2)
A.
and
more
tribes
(see
of
revision
been
have, therefore,
Other
the
here
accepted
evidence
Notochnella
sect.
and
sustaining
is
view
7
Tiegh.)
(v.
Ochna.
Van
Kanis
Tiegbem's
named Ochninae and Ouratinae
subtribes,
as
this
Area
Indo-Pacific
derived from palynological
data
13).
p.
at
are
first
sight rather
which is probably caused by their adaptation
ones,
the
of
Brackenridgea
The African species ofBrackenridgea
distinct by persistent stipules and
are
Ochnaceae
intermediate between Brackenridgea
less
or
respectively.
A
the newly introduced
Gomphia,
being
Kanis :
Possibly they
treated best
are
as
a
yellow pigment below the bark (cf. Robson,
a
The latter
with
genera
subtribe
1-celled
ovary,
Six genera of the
regarded
by only
all
genera
and African
St.
Hil.
usually
with
genera
In
my
whereas the
species,
the
opinion,
species
many
Sauvagesinae and Luxem-
subtribes,
5
the other neotropical
and
3 —5-celled
Sauva-
—3-carpelled,
2
a
American
and the remaining
genus
The
ovary.
and
stamens
ones.
of which have
always been
Neckia Korth. is also
represented
Sauvagesinae have been treated here, four
monospecific.
as
one
3 —5-carpelled,
and
all Asiatic
viz.
a
L.
gesinae comprise Sauvagesia
two
includes Luxemburgia
and
stamens
5—c\3
1962).
distinct section.
TheSauvagesieae have been subdivided into
burginae.
different from the Asiatic
seasonal drought conditions. They
to
ofSchuurmansia Blume have been reduced
three.
to
The Luxemburginae
are
identical with the Luxemburgieae in Planchon's original concept.
It should be noted that the subtribal subdivision
of
ment
Engler
genera
should
1
be interpreted
not
arranged according
the system
are
proposed
more
or
is
as
a
phylogenetical
similarities in their
to
respective evolutionary
ships
levels in certain
was
meant
less
levels
as a
or
sensu
tree,
since the
organs
are
designed by Jane
supposed
actual
on
and the results of her
tree
Most
more
the
an
on
seem
be
to
evolutionary
towards
attempt
system will be highly
the taxonomic
Univ. N.Y.,
morphological and anatomical data presently known. However,
lacking,
to
especially of the Sauvagesieae
(Thesis Fordham
anatomical studies. It is
taxa
on
(1966). She based her assumptions mainly
each phylogenetical
merely
morphology.
on
of theOchnaceae,
AmericanOchnaceae
own
are
the other hand,
On
relationships.
the subdivision presented.
made in the chapter
Decker
taxa
natural system, since it is assumed that natural relation-
reflected by
hypothetical evolutionary
opinions of Dwyer
are
arrange-
morphological characters, without regard
advanced in certain characters and primitive in others. Some remarks
A
an
(1874).
Table
their
will result in
proposed here
different from that in the tribes Sauvagesieae and Luxemburgieae
a
long
as
1941,
vidi)
non
synthesis of all
as
fossil records
speculative and, therefore,
not
of great value.
Finally,
a
remark should be made
although nothing
new
is added here.
on
suborder Theineae, and this position
They
to
usually
raised Engler's
the relations of the Ochnaceae with other
Engler (1909) placed the family
was
suborder,
not
in his order
fundamentally changed
with about
the
same
by
families,
Parietales,
later authors.
combination of families,
the rank of order, naming it either Guttales (Hallier /, 1912), Guttiferales (Wettstein,
1935;
Pulle, Compendium,
1952),
or
Theales
1938; Melchior 1964), Clusiales
(Takhtajan,
1959).
primarily referred the Ochnaceae
(1926).
Theales
to
his
Later he referred the Ochnaceae
(1959).
Guttiferales.
Most authors
agree
was
given by
exception
Decker
was
Theales which he
to
his
on
kept
among
the hypothetical
(1966).
(Pulle, Compendium, ed.
made
Ochnales, again
that the family is
A discussion of the views
of theOchnaceae
An
by
apart
a
Hutchinson,
3,
who
from his Guttiferales
finer segregate from his
the
more
primitive
in the
phylogenetical relationships
8
BLUMEA
VOL.
MORPHOLOGICAL
The
vegetative
in the
to
trees
usually adapted
sparsely branched shrubs
a
The
united,
entire
or
like the stipules,
of
Ochna spp.
is distichous in the
habitats
to
large
in
are
stiff, glossy,
some
Apart
parallel secondary and
primary
curved upward.
parallel
shows
to
v.
in
In
over
more
the
a
85
to
up
of the Ochnoideae
the
in
only
occur
long
usually
linear-
or
is sub-
texture
habitats. The margin
exposed
more
cm
are
obovate-lanceolate
are
which,
American
the
curved
or
less
nerves
is characteristic
distinct
spp.
for certain groups.
is found in the Sauvagesoideae;
nerves
tertiary
the
nerves
nerves
are
intercalated
are
remote
more
strongly curved upward and
are
and
some are
considerable length. Finally, the Asiatic species of Gomphia
connected by
nerves,
intra-petiolarly
Tiegh.
In the Asiatic Ochna
Brackenridgea
the margin
straight
sometimes
ones.
or
be small in certain
may
distinctly darker;
leaves
Compound
parallel, only slightly
many
between the
and above
shape and size, the venation of the leaves
from
advanced
various types
leaves of
Sauvagesoideae:
coriaceous
truly
denticulate.
or
Rhytidanthera splendida (Planch.)
A pattern of
some-
in the Sauvage-
in the Sauvagesoideae.
to
They
groups.
henningsii. The leaves
Schuurmansia
chartaceous
or
serrulate
often
often
they
as
are more
caducous, free
or
(ovate-)oblong, whereas those of the Sauvagesoideae
coriaceous
drought, but
to
Ochnoideae, non-distichous
eitherpersistent
are
often characteristic for certain
are
lanceolate. All leaves
one or
distinct intra-marginal
two
nerves.
inflorescence
Several kinds of inflorescences
simple
or
compound, lateral
times
even at
occur
in the Ochnaceae
;
terminal, with
or
always articulate. The mode
of
branching
is
or
cymose,
without
often
a
a
flowers
useful taxonomic
character,
are
some-
inflorescences
lose their leaf-like
are
appearance,
become compound
from
a
leafy branch,
(fig. la). By branching from the axils of the bracteoles, the
flowered inflorescences could develop into simple
cymose
thyrsoid,
specific level.
bearing lateral
If the
or
racemose,
terminal flower. Flowers
Theoretically, all different forms ofinflorescences could be derived
can
Sauvagesia L. It is
family,
ecological conditions like long dry periods,
extreme
stipules which
have been measured in
The
genus
in the
laciniate, and usually small. The leaves belong
dry
bushy shrubs.
as
in the tribes Sauvagesieae and
(see below).
phyllotaxis
AllOchnaceae have
of which have
Small shrubs in the subfamily Ochnoideae
state.
soideae generally do not show such striking adaptations
in floral characters
considerable
a
some
only known
are
occur
primitive life form
more
less advanced
rather
to
a
to
grow
spp.,
they develop typically pyromorphic forms. Undershrubs and herbs
times
is
species
only found in the amphi-Atlantic
are
represent
show other characters in
height. Others
metres
30
over
more
Euthemideae. Real herbs
assumed that taller
Some
rain forest like the MalesianBrackenridgea
attain
Smaller treelets and
are
REMARKS
woody family.
typically
a
are
tropical
been reported
1968
I,
parts
The Ochnaceae
height
No.
XVI,
monochasia
cymes,
or
one-
dichasia (fig. ib).
conferted towards the end of the branch and the bracts
the inflorescence becomes
thyrsoid (fig. ic).
by incorporating branches
once more
at
Such
the base that
are
a
thyrse
likewise
thyrsoid (fig. id).
Thyrses of
florescences,
type
c
in others
are
as
found in
lateral
Brackenridgea,
ones as
the terminal bud of the shoot is often capable
The
cymose
branches with
Brackenridgea the flowers
three
seem
to
or
in
some
species
well. A terminal flower is
more
to
grow
flowers
be arranged
are
evenly
on
a
as
terminal in-
developed
here and
vegetatively after flowering.
more
in
only
not
or
less
group
shortened. In
of conferted
sect.
umbels,
A.
Fig.
1.
Concept
rather than in
a
sometimes show
more
between b and
stage
Thyrses of
not
ones.
thyrse.
type
In
sect.
It is
As
inflorescence
flowers
or
stems
from
simple
branches
Indo-Pacific
The
the Ochnaceae.
arrows
but
they
indicate
are more
the
be reduced
into a
a
theoretical
develop-
indicating
not
a
gone
so
far and the
somewhat intermediate
to
one
main shoot of the
mention here the
yearly,
old
As
wood.
and without
type d
a
a
same
pyromorphic
it
is
result,
forms
impossible
a
leafy
occur
to
are
branches
develop
a
short
lower
obtusata
a
formed
ib,
on
ic,
upper
var.
specialised
with
axile
part).
Gomphia serrata, although
terminal
less shortened and they usually have
terminal
(compare
of Ochna
shoot is
in the Asiatic
The inflorescences
are
season
terminal flower (compare
frequently
too.
cymose
flower, especially the higher
flower, making monopodia! growth impossible.
burned off
found here
or
9
sense.
leaf-like nature,
may
developed
to
are
cymes
are
Area
also found in the Asiatic species of Ochna. The
Compound thyrses of
simple thyrses
the
compensated by the fact that the inflorescences
interesting
all
of
Notochnella the reductions have
side-branches, below the vegetative
part).
in
structures
Ochnaceae
c.
c are
The terminal bud is
pumila.
the
evolutionary
clearly their
much shortened,
This situation is
an
of
revision
A
of inflorescence
interpreted in
ment, not to be
are
Kanis:
or
lateral. The
cymose
reduced number of flowers,
BLUMEA
10
especially the higher
of the
A
(fig. re) could
raceme
less
or
(fig. if)
can
also be explained
reduced
to one
as
be
interpreted
be understood
compound
a
as
another specialisation
several
composed of
as
simple
a
more
compound
A
but it could
racemes,
cymose
viz.
a,
branches have been
of
a
to
in
for
nature.
racemose
grow
and Euthemideae have been described
Sauvagesieae
pairs
practical
There is
tendency
a
triplets, which
or
specialised study
reveal their
to
true
nature.
although
reasons,
is
dislocations,
or
With the
the flowers
among
exception
by
I doubt that these
indication of a thyreoid
an
this tendency is obscured by reductions
as
of type
lateral flowers and reduced leaves.
panicles
or
racemes
ultimate branches
However,
growth
flower.
completely
are
developed, making sympodial
are
compound thyrse (fig. id) of which all
a
The inflorescences in the tribes
me as
1968
I,
necessary.
shortened axis with solitary,
raceme
No.
XVI,
Terminal flowers
ones.
branches
vegetative
VOL.
of the
nature.
it would
require
of Neckia, all
genera
treated have compound, many-flowered, terminal inflorescences, which make sympodial
growth of the vegetative
term
‘Rispe’;
panicle
is
suggests
The
a
more
neutral
The
are
is
classified best by the German
probably lacking. The
treated here.
most genera
very
term
the mode of branching, but it
to
profusely flowering panicles.
Indosinia, Indovethia
Sinia,
genera
florescences which
organised branch
fading
are
two
to
is concealed here
branching
out
The flowers
are
much obscured
by the
the
genus
in
E.
and
apex
are
very
may
of somewhat
by concaules-
have
raceme-like
in-
occasional, similarly
primarily
are
The ultimate mode of
age.
strong reductions. This type is considered
by
me as
one.
Euthemis both of the forms just mentioned
whereas E.
leucocarpa,
have
an
much shortened and
flowers of successive
many
advanced than the previous
occur
and Schuurmansiella
,
towards the
base. Other branches
at
bundles of
resembling
In
in relation
raceme
The ultimate mode of branching
age.
are
and recaulescences.
cences
more
than
one
Schuurmansia has
genus
different
Possibly they
pyramidal form than those found in
more
a
parts necessary.
should be noted that terminal flowers
it
are
found. Small panicles
raceme-like
slender,
minor has
inflorescences with
shortened branches.
The
few
to
Neckia has lateral inflorescences that
genus
plants, but
often
the old wood in
it
with
thyreoid
distichously
can
be stated that
branches
cymose
Those
arranged.
but their ultimate mode of
The
larger
ones.
of
found above the leaves in smaller
are
They
empty bracts and
several, spirally arranged,
Summarising,
or
on
the inflorescences
which,
is
not
yet
composed
of
rachis with
a
of the Ochnoideae
with
accordance
in
the Sauvagesoideae
branching
are
a
articulate, terminal (?) flower.
one
raceme-like
are
are
cymose
the phyllotaxis,
are
paniculate,
or
completely understood.
flower
AllOchnaceae studied have
there is
tendency
a
to
regular
flowers. In
some
American
zygomorphy of the androecium which
genera
ofthe Sauvagesieae
is manifest in Luxemburgia
St. Hil.
The
calyx
is pentamerous,
Planch, with
very
shortly
ten,
quincuncial,
connate
at
the base.
caducous in several American
the
outer
forming
a
two
or
kind
Sauvagesoideae
except in the American
and Elvasia DC. with three
three
sepals
They
genera.
are
of wings. Ciliate
(Euthemis,
are
to
six
often persistent and
In the
African
considerably,
margins
Sinia, Indosinia, Neckia,
genus
though
occur
genera
lobes. The sepals
in
accrescent,
free,
unequally
genera
The
or
only
but they
Lophira Banks
several
Indovethia).
Blastemanthus
are
ex
enlarged
are
Gaertn.,
in
fruit,
of the subfamily
outer
two
and
a
half
A.
Kanis
sepals that primarily
than the inner
one
six
to
five
petals
petals
revision
of
the
Ochnaceae
the flower bud,
cover
the
of
Indo-Pacific
Area
11
usually much thicker and
are
caducous, contorted, and usually
is often found in certain
petals
in the American
occur
pentamerous. An
Elvasia DC. It is assumed that five
genus
original organisation of the Ochnaceous
represent the
inner whorl
extra
species of Ochna andBrackenridgea.
higher level.
apparently
Ten stamens,
in
whorl,
one
found
sect.
in
Notochnella and Ochna; I have counted
Brackenridgea
O.
sect.
jabotapita, but
up
to
Brackenridgea. Many
120
stamens, in
flower in O. obtusata
per
than
more
few
as
generic
at
Gomphia, Ouratea,
in
whorl,
one
twelve
as
occur
in
stamens
pumila. Staminodes
var.
to
flower.
are
and in Brackenridgea
Three
sepals and
The androecium shows various patterns, especially important taxonomically
or
fleshy
more
ones.
The corolla is
of
A
:
are
never
found in the subfamily Ochnoideae.
More variation
occurs
found in flowers with
antesepalous
stamens
a
in the
are
there
are
neotropical subtribe Luxemburginae
no
staminodes,
staminodes in
The
except
usually has
and
stamens
staminodes,
which
are
staminodes that alternate with the
staminodes in the
Neckia),
five distinct
there
whorl
forming
nerves,
one
are
same
or
The
eight, and
or
and
stamens
more
more
a
less
or
kind of
supplementary
connate
usually have
stamens
small
many
stamens
or
a
as
the base.
at
distinct
one
there
are
always
five linear
multiple of
Here
the five
The other
nerve.
about free ( Schuurmansia, Schuurmansiella,
smaller and
are
homogeneous,
more
whorl of five,
opposite
organs,
corona
whorls
outer
the
to
stamens,
with
(. Sinia, Indosinia, Indovethia).
of
gland-like staminodes (Schuurmansia, Schuurmansiella,
the number of fertile
ten
subtribes.
five
rarely
stamens,
which has
an outer
fused into five larger, petaloid
or
fertile,
alternating staminodes.
many
is much
pantropical subtribe Sauvagesinae
spatulate
often
are
five
Euthemideae,
whorl.
five fertile, antesepalous
or
In the tribe
important differences between the
Blastemanthus Planch,
outer
an
stamens.
sometimes with five
found,
In the tribe
Sauvagesieae
where staminodes
subfamily Sauvagesoideae,
reduced number of
many
Sinia,
and the fusion of staminodal
two
smaller, filamentous
or
Neckia). The reduction
structures
to
Sometimes
of
should be considered
advanced characters.
The
carpels
Ochneae,
found in Ouratea,
whorl
occur
occasionally
of three
to
are
which represents
taxonomic
fused into
a
in
Brackenridgea
single
The Ochnoideae have
ovary
in the tribeElvasieae,
and in
carpels
many
stamens
is
a
per
and
more
primitive character
of Ochna is
flower in species
a
partial
complete
or
than
a
carpels
to ten
Ochna, although
been counted in flowers of O. integerrima. It is assumed that
seven
Five
structure.
Brackenridgea; three
sect.
Notochnella
sect.
characters.
a
primitive
more
Gomphia, and Brackenridgea
large number of carpels
relatively
yields important
ovule each. The carpels
one
but free in the
one
also
gynoecium
with
a
up
are
carpels
to
15
in
have
variable number
fixed number of five.
usually found together
A
with
duplication of the corolla. Moreover,
such flowers usually have larger flower parts than others. This combination of characters
frequently
occurs
in O. obtusata and O.
whether polyploidy could be the
The Sauvagesoideae
have
two
cause
to
integerrima.
five
found in the tribes Sauvagesieae and
found in the subtribe
the
apex.
Two-
or
with
a
reduced
two
ovules
Lophireae.
per
three-carpelled, one-celled
are
ovaries
usually intruding
number of carpels
probably
are
one
to
ovary.
cell. Carpels with
Three-
Luxemburginae, although the septs
sinae, but the parietal placentas
interesting
fused into
carpels, always
Euthemideae has five-celled ovaries with
are
It would be
investigate
of these deviations of the normal pattern.
are
are
to
The tribe
many
five-celled ovaries
ovules
are
also
sometimes incomplete towards
found in the subtribe
Sauvage-
towards the base. One-celled ovaries
more
specialised
than others.
The
BLUMEA
12
in the tribe Sauvagesieae is
placentation
and it is
ovary
VOL.
completely basal
No.
XVI,
more
less restricted
or
in the African tribe
Lophireae.
of ovules in the Euthemideae
probably indicates another
The
has
torus
floral
spacing
no
disk-like appendages.
whorls,
sometimes
is found in the Ochneae:
to
the lower half of the
A reduction of the number
specialisation.
Extensions of the floral axis,
the Sauvagesieae,
in
occur
Indosinia and the gynophore in several American
fruit,
1968
i,
genera.
about hemispherical
like
swollen
A
in the
the
in Ochna and
torus,
or
subglobular
Ouratea and
in
in
suk-
Brackenridgea,
O
of
of
especially
-
cylindrical
sense
anthophore
■
Gomphia.
Fruit and seed
Ochnoideae. In the tribe Elvasieae each flower develops
x.
one-celled, one-seeded, non-dehiscent fruit.
one
or
as
of these
some
subtribe Ochninae they
remarkable for the
and
spaces
Ochnoideae
a
are
nuts,
a
is
contrast
bluish
are
swollen
torus
adaptation
an
inward projections
two
number of
black when ripe. In the
or
and
an
to bird
of the endocarp,
enlarged calyx,
both
dispersal. Brackenridgea
forming
connection, around which the seed is curved.
The
Euthemideae have red
probably
are
air-filled
two
The seeds of all
or
each
white berries,
dispersed by birds. The Lophireae
with five
one-
have one-seeded,
unevenly winged by the persistent calyx. The Sauvagesieae have coriaceous,
septicidal capsules with
The seeds
genera.
flower produces
without albumen.
seeded pyrenes. They
woody
contrasting with
this colour
transverse
Sauvagesoideae.
2.
aborted. The drupes
are
are
purplish red. Evidently,
is
single, stellate, coriaceous,
a
the tribe Ochneae each
one-celled, one-seeded drupes, although usually less than the
more,
carpels,
In
many
of the
seeds which
Lophireae
winged
are
contain
in Schuurmansia and
oil, those
of the other
some American
Sauvagesoideae
are
albuminous.
Embryological
(Bull.
characters
Mus. Hist. Nat.
of embryos and added
8,
proved
1902,
some
more
interpreted and evaluated these
here,
far
as
as
'isocotyled,
relating
taxa
to
endocarp
types,
these
adding schematic drawings.
descriptions,
Ochna species of
taxa.
incumbent' embryos.
and also
short descriptions of
are
Brackenridgea
sect.
species
strongly curved
of
it
as
studied. However, Van Tieghem's
Notochnella have seeds which
and
gave
ten
types
later. Farron (Bull. Soc. Bot. Suisse 73,1963,197—203)
of these authors in the
specialised terminology
distinction between the
be important in the tribe Ochneae. Van Tieghem
to
208—218) originally
I have
was
terms
Ochna have
necessary
erect
the
for
seeds and
Brackenridgea and
sect.
Gomphia
used
not
should be mentioned
around the inward
'isocotyled, incumbent' embryos.
not
sect.
projections of the
serrata
has curved seeds
'isocotyled, accumbent' embryos.
Anatomical characters
A short
much
to
paragraph
anatomy should be
on
the concept of the Ochnaceae
as
added,
as
studies in this field have contributed
natural family.
a
I
have
not
carried
out
any
anatomical research myself.
Gilg (Ber. Deutsch.
family
canals
in
from
are
an
Bot. Ges.
was
of Tetramerista
included in the
Van
Tieghem
especially
1893,
—25) demonstrated the homogeneity of the
20
characteristic for the Ochnaceae.
the Dipterocarpaceae.)
exception
11,
anatomical point of view. He found that cortical bundles without resin
in the
accepted
Miq. which
(Cortical bundles with
the circumscription
was
of
resin canals
Engler
(1874),
excluded, and Lophira Banks
ex
are
found
with the
Gaertn. which
family.
paid
taxa
He
much attention
that
are
to
referrred here
anatomical characters
to
in
the Ochnaceae s.l.,
the subfamily Ochnoideae. In
one
publication
A.
Mus.
(Bull.
Hist.
Kanis
Nat.
characteristic for this
Metcalfe and
:
A
8,
Chalk
types of wood
(1950)
me
pointed
also
some
Jane Decker (1966)
in the
as
representing
vascular
anatomical
an
buminosae’
vascular
the
the
rays,
a
the
be
other,
do
usually
more
The
pollen
Up
wood
simple
till the present,
the
on
not
pollen
conclusions
are
Investigations
at
generic
a
1.
1 —3
made of
Lophira
If
one
have
objections
anatomical
often tall
and
family
subfamily
Lophireae
vasicentric
vascular
heterogeneous
to
subdivision of the
a
Lophira
the
from the ‘Exal-
homogeneity of
will be
grounds. However,
whereas the other
trees,
size. Smaller sizes
(cf. Decker,
its
of
plants
are
it should
Sauvagesoideae
possibly correlated
1966).
a
limited
investigations
amount
in this
published separately,
24
of all
species
in
tribes. It
12 genera,
was
but
of information
field
some
recently
were
of his
The
type:
following
types
Lophira, Neckia,
mainly
tentative
from Southeast Asia,
demonstrated that the pollen
were
interesting differences
of all
were
but
taxa
found
distinguished:
Schuurmansiella.
Euthemis, Indosinia.
Schuurmansia type: Schuurmansia.
Ochna type: Ochna, Brackenridgea.
Elvasia type: Elvasia.
Ouratea type: Ouratea,
or
common
pores
are
and
Gomphia.
that colpi and
pores
bridged colpi. Type
mainly based
on
7
are
equally well developed. Types
has reduced colpi.
Other distinctions
the thickness of the wall, the development in the
less distinct ektexine and endexine, and the development of
layer of pillars under
compartd
in her
handand the Sauvagesieae and Euthemideae
on
5.
more
‘Luxem-
Indovethia type: Indovethia.
4—6 have in
distinct
the
on
shrub
His results
were
between the types
less
its
briefly rendered below.
have reduced
wall of
Lophira.
correct,
the tribe
parenchyma,
'cristarque' cells,
6.
7.
Types
its lack of
are
structures
3. Euthemis type:
4.
main
Ochnoideae,
genus
be
to
Lophira
The removal of
general resemblance, but taxonomically
2.
certain
vasicentric tracheids.
no
no
are
(see table 1).
Lophireae
higher levels.
or
and
only Erdtman (1952) has published
including representatives
studied has
of the subfamily
genera
and Elvasieae
of the Ochnaceae. Some further
made by J. Muller.
for
They distinguished three
axial
metatracheal
clearly indicated
beyond
not grow
with
characteristic
as
cortical
of vasicentric tracheids, and the type of the axial parenchyma.
presence
in mind that the
kept
cells
she accepted
(1925),
of view, there
point
proposed
one
perhaps
is
are
the sub-
in
distinct tribe. However, she found that the
rays,
relationship between the Lophireae
on
united here
amongst others again Tetramerista from the
Gilg
fully justified by
appears
that
rather isolated.
diffuse and vasicentric axial parenchyma,
From
cells
of the
importance
genera
considered by them
was
tracheids, whereas the ‘Ourateae’ (Ochneae)
Ochnaceae like
13
studied the anatomy of the wood, especially in
homogeneous
rays,
A
taxonomic
cells in the
Sauvagesoideae.
the Ochnaceae
anatomical grounds. Following
have
the
to
apotracheal
(Sauvagesieae). She excluded
‘Exalbuminosoideae’
Area
Indo-Pacific
of the tribe Sauvagesieae, metatracheal in the
in
position, however, being
on
the
mentioned mucilage
viz.
genera
The inclusion of Lophira
burgieae
of
he described the 'cristarque'
'cristarque'
the subfamily
to
parenchyma,
in
paratracheal
Ochnaceae
266—273)
1902,
Ochnoideae. They
referred by
genera
the
subfamily.
bundles in the Ochnaceae and the
family
of
revision
with the
a
tectum
a
more or
in the ektexine.
proposed subdivision
of the Ochnaceae
(table 1),
it will be
seen
14
BLUMEA
VOL.
XVI,
No.
I,
1968
gen ralised.
less
or
more
are
areas
Outlined
area.
Indo-Pacif
the
in
Ochnoidea
subfamily
the
of
genera
the
of
map
Distrbuional
2.
Fig.
A.
that types
1—4
Reduction of
Kanis:
and
and
In the
Elvasia
also
is
similar
to two
rather
being
of these four.
pollen
far
as
related
type
more
the
in
other.
Sauvagesoideae,
well
developed
the rather primitive,
less
or
In the
types.
of
distinct pillars
is
studied,
as
represented
by
types does
Lophira
genus
four
different
and Euthemideae
the coherence of the
the position of the
be
can
the situation
Sauvagesoideae
Although the presentation of pollen
not
subfamily
in this
are
support
appears
subfamily
one.
of Tetramerista
those in
to
in the
of the tribes Lophireae
types
respective pollen
natural
a
pollen
occur
5 —7
tendency
easy.
their respective
well demonstrated. This confirms
The
15
the Ochnoideae. Equally
the proposed subdivision of the Sauvagesoideae,
as
a
and the types with
type
is
different. The tribe Sauvagesieae,
types,
be
to
Area
Indo-Pacific
whereas types
appears
found in
only
the
of
the tribe Elvasieae and the subtribes Ochninae and Ouratinae
by
whereas the
Ochnaceae
subfamilies, but distinction between
(4)
(5, 6)
Ochnoideae,
distinguished
the
subfamily,
one
is
colpi
thin-walled Schuurmansia
Ochna and
of
bridging of colpi
in both
occur
pores
revision
found in
are
pores
whereas reduction of
colpi
A
the
is
Miq.
not
referable
to
but is
the Ochnaceae,
certainly
Theaceae.
GEOGRAPHICAL REMARKS
of data
Summary
The Ochnaceae form
subtropics.
In Asia
whereas the
in northern
In
to
the subfamily
and the
Atlantic
subtribe
do
c.
tribe Ochneae.
and
Ouratinae
not
is
also
v.
by
partly
c.
in
representatives
0
at
24
cross
the
extra-tropical,
N. is
the Tropic of Cancer
the neotropical,
are
a
mainly amphi-
Ochninae.
Amphi-Pacific
subfamily.
neotropical
and Idertia Farron,
one
subtribe
palaeotropical
a
tribe Elvasieae
monogeneric
The latter tribe is subdivided in
within the
occur
Tiegh.
represented,
few
a
30° N.
The subtribe Ouratinae comprises the
Rhabdophyllum
only
Sima from Kwangsi
genus
Ochnoideae there
pantropical
relationships
the
Ochna obtusata and O. integerrima
species
India,
family with
pantropical
a
only
variable species,
the African
genus Ouratea,
and the mainly
in parts
African Gomphia
of continental Asia and
genera
which
western
Malesia.
The subtribe Ochninae comprises the palaeotropical genera Ochna and
Brackenridgea.
Ochna
is
Ochna
also
five
species in
the
in
occurring
two
species, probably
In
by several species
represented
a
distinct
are
African
Sauvagesieae. The latter tribe is
in
six
the amphi-Atlantic
two
genera
genera
genera
Africa and
to
respectively
the
Fiji,
and
sect.
comprises
some
other
Madagascar.
the monogeneric
are
Malesian
subdivided into
tribes
and
Lophireae
the pantropical
subtribe
neotropical
and species of the Sauvagesinae
in
with similar
restricted
from
seeds
to
are
are
Southeast Asia and the
and
tribe
Luxemburginae
found in tropical America,
western
is
especially
Borneo and Luzon
found
continental
in
Pacific,
and only
Vausagesia Baill.) is found in Africa with
of the Sauvagesinae in Malesia s.l.
rather than via Africa. This
are
eastern
Sauvagesia L. (inch
genus
winged seeds, occurring
treated
there
and
occur
species. The relationships
Pacific
in
only
species. Brackenridgea
subtribe Sauvagesinae.
The majority of the
eight species
section,
Sauvagesoideae
subfamily
and the pantropical
three, mainly African sections,
with four
sections, ranging from the Andaman Islands
in
Euthemideae which
in
continental Asia
true
to
are,
therefore, amphi-
for Schuurmansia,
the Solomon
South America only.
Southeast Asia and
a
genus
Islands,
with
since other
The other Sauvagesinae
western
Malesia.
Sinia
and
BLUMEA
16
VOL.
XVI,
No.
I,
1968
gen ralised.
less
or
more
are
areas
Outlined
area.
Indo-Pacif
the
in
Sauvgesoidae
subfamily
the
of
genera
the
of
map
Distrbuional
3.
Fig.
Kanis :
A.
Indosinia
seem to
A
be limited
revision
small
to
Neckia is found in Sumatra, the
apparently
occurs
Schuurmansiella
central
in
Distributional maps of the
subfamilies
(figs.
their localities
by
delimited by
included
Less
as
generalised
less
or
whole
a
in
areas
certain localities
to
of relatively
if only
aieas
or
a
whereas
distributed
indicated
genera
are
groups
are
known from such island.
aie
6,
island
or
respective
are
genera
larger islands
4,
for the
given
widely
few collections
Indovethia
in Sarawak.
the distributional maps of
in
given
are
one
respectively.
only,
Borneo
area are
more
since
17
Philippines.
monospecific
rare,
Brackenridgea, Euthemis, and Schuurmansia (figs.
Ochna,
and the
western
in the Indo-Pacific
genera
areas
and
Sumatra
eastern
Area
Indo-Pacific
and South Vietnam
Kwangsi
generalised outlines,
even
the
of
presently known. Those of
as
more
The
3).
2,
Ochnaccae
Malay Peninsula, Borneo,
be restricted
to
appears
the
of
under the
species
genera
8).
7,
Discussion
The present
distribution of
and
geological
distribution of
most
It is
in
in those of
Distributional
due
The
genus
the
exploration
species
are
Malay
Ochna jabotapita is confined
to
the
closely related
are
Africa, of which
neighbouring
dry
a
O.
islands
well,
as
is almost absent.
season
spread
never
the Mascarenes, such
are more or
less adapted
O. lanceolata from Malabar and
to
Schweinf.
(Forsk.)
inermis
genus
collection from Samar.
probably likewise
are
being
Ceylon,
in
beyond
into Malesia
south-western part of Ceylon. Its probably
everwet
Madagascar and
in
are
that the
Peninsula.
species occurring in continental Asia
that
one
its Asiatic subcenter
origin,
and present
areas.
found of the four treated. It
the northern half of the
closest relatives
in
arch, where
of these
past
imperfectly known. The
to occur
prove
Ochna is doubtless of African
genus
where three
will
easternmost
by
mind, however,
in
in North and South Sumatra and South Borneo
gaps
insufficient botanical
to
kept
only known from the Philippines by
is
that this
probable
very
particular
be explained
course
treated is still
and species
genera
Neckia, for example,
should of
taxa
circumstances. It should be
ecological
also
is
as
O.
The
mauritiana DC.
seasonal drought. Species
to
Ceylon,
are
found in
found in
eastern
the mountains of
Yemen.
Ochna obtusata and O.
from
more
East
a
related
and
genus
a
upward
at
not
Malesia.
It does
Celebes,
which
not
to
too
O. lanceolata.
rise
to
not
the valley
in
adapted
in
Java,
might indicate
to
to
in Asia is
its
more
Climatic
relatively
at
caused
specific
They
probably
are
each other in Assam and
the Chittagong
eastward
or
less
Hill Tracts and
the
down along
coast
of
recent
by
have
arrival in
more
level,
western
of Sumatra and
the
Malesian
east
area.
ecological changes.
recent
although
some
local
races
it
possibly
Birma into Further India and
west
and
climate, although
everwet
conditions in the past
the islands north and
certainly
differentiation
time.
a
from where it reached Ceylon
Africa,
a
at
meet
of the Brahmaputra.
spread through India and
occur
They
westward into
dry period.
severe
closely related: possibly they developed
became disjunct
obtusata occurring
through
time. It is
a
disjunction
given
O.
rather
area
Gomphia also originated
permitted Gomphia
present
are
integerrima spreading
O.
southern India
stand
than
jabotapita
and Garo Hills,
Chittagong
can
O.
to
Pakistan,
the Khasia
The
integerrima
single species, of which the
of
The
It has
can
be
distinguished.
The
genus
Brackenridgea
Gomphia. Evidence for
tion.
Although occurring
Malesia s.l.
17
on
the
was
probably
in Malesia
before
this assumption is found in its present
other,
in
it
eastern
appears
Africa and
to
be
the arrival of Ochna and
wide, but disjunct
Madagascar
completely absent
on
the
one
distribu-
hand and in
from India and
Ceylon.
BLUMEA
18
In
western
Malesia it
coastal
area
Southeast
it does
as
It is
the
other, form
with
areas
relict-like
short dry
a
Only Schuurmansia
area.
the
Moluccas
represented
is
elegans
is
more
common
on
and the Pacific
It
remarkable,
is
distinct section.
a
Asia and
the
Pacific
western
Indosinia, Indovethia,
monospecific
common,
pioneer-like
geologically
in
nature
of its species.
of Makassar
through
disturbed
more
It is assumed that
than
Strait,
general
same
from Borneo
species, reaching
occurrence
Neckia
genus
Samar, which is within the
three
Its
both sides
on
Malesia
western
section.
one
The fact that Sinia,
more
to
by
in
with restricted distribution probably indicates
The
Solomon Islands.
is understandable from the
regions
S.
the
to
ecological conditions,
recent
palustris
B.
in Southeast
genera
of the subtribe.
from continental
season.
within
groups
distributed from the Mentawai Islands
is
to
northeastern,
wet,
genus
Malesia, Queensland,
eastern
affinities with those in Africa.
nature
the
Guinea,
from the Philippines represents
andSchuurmansiella are monospecific
a
Gomphia, but including the Andaman
area as
probably due
very
even
the Sauvagesinae
concluded,
close
1968
i,
found in New
distinct
slightly
two
that B. fascicularis
no
No.
surprising that Brackenridgea hookeri and
already
As
in
XVI,
and in Fiji. The apparent absence of the
and the Moluccas is
occur
same
it is
side and B. forbesii and B. nitida in
however,
have
Eastward
Queensland,
Asia
not
not
one
the
on
of
about the
covers
and Mentawai Islands.
VOL.
be concluded
can
from present collections.
The
monogeneric tribe Euthemideae
Cambodia.
in
area
and it
may
centre
of that tribe. Apparently it
the
genus
or
dry
It
is
a
of West
cross
different
such
genera
that
matter
The
Karimundjawa.
as
genera,
as
they
are
Literature concerning
to
are
is
to
drier and
a
east
more
usually found
taxa
ON
age
of
poorer
type
of Sumatra.
been collected in Java and the Lesser
ever
makes
subjected
is
area
occurrence
to
of most Malesian
the barrier
crossed
never
late in the
too
since it is found
difficult
to
THE PRESENTATION
as
as
province
be able
to
to
the Lampong
complete
in
use
OF
as
in West
occur
of Java
poor sandstone soils
treated is given
far south
as
not
volcanic soils
young
everwet
area
understand, why widely distributed
and Neckia should
on
young
which is probably
leucocarpa,
adapted
of Sumatra,
west
relatively
a
Ochnaceae have been found in the
that the
importance
or
are
not
in peat
Java.
The
suitable for
swamp
forests.
DATA
possible.
Enumerations and
for reference in
certain
areas,
in this respect. A selection of general works of reference
under the family.
cited literature is
taxa,
than E.
is
species
given only if being generally
if being of historic
given
Borneo which is still the Malesian
Malesia.
Java Sea,
It is
REMARKS
lists, however,
no
Brackenridgea, Euthemis,
these
The
area
small
a
the tribe Sauvagesieae
in
species of Ochna apparently
western
young
probable explanation
is
smaller
Gomphia probably arrived
genus
most
or
stock
in Borneo and the islands
The deciduous
Sunda Strait and the
Districts and
a
Malesia and in
western
reached the islands north and
Java eastward, which
lowland rainforest in
Java.
western
near
Ochnaceae have
from Central
improbable.
everwet
old
an
However, it should be noted in this respect that the
monsoon
genera
of
no
found in
which probably indicates
covers
common
remarkable that
Sunda Islands.
a
only
the fact that the former
kerangas, especially
It is
or
never
the Philippines,
Euthemis. E. minor
explained by
best
of
well have originated in
Celebes,
Java,
is
it descended from
Possibly
secondly
synonyms,
also
arranged according
to synonymous
chronologically.
basionyms
in
to
the homotypical
method, that
chronological order, finally
to
is
first
homotypic
A.
Kanis
:
A
The synonymy of the respective
nuda,
ttomina
far
as
subfamilies,
tribes,
taxa
and lower ranking
key
under the
given
the
Inio-Pacific
names
Area
19
and effectively published
treated. Full references
area
given under family,
are
whereas only relevant basionyms
species,
repeated
are
and varieties.
of the
has been taken
care
rank and
same
facilitate comparison
to
avoid duplication with those of higher
to
taxa.
the subfamilies, tribes, and
to
of
includes all valid
taxa
the
Ochnaceae
drafting the descriptions, all possible
In
between those of
are
to
and
genera,
subspecies,
under sections,
A
relevant
as
the
of
revision
respective
is
genera
those
genera,
given under
subspecies
to
or
the
family. Keys
species
to
varieties under the respective
species.
The distribution of each
specimens
The collections
and
areas
on
cited
are
The
spelling
A
as
of
specimens without
with
or
left
is
generally running
within
that
area
aire
that of the Times
to
given
at
certain
a
the end.
Atlas, midcentury
names
on
was
chosen instead of
and numbers. Advantages
the distribution of
and the
taxa
identification
an
of the accepted
possibility
method
mention old
to
collector's number. Only few collections without collector's
a
geographical
ecological
annotations
annotations
no
than
precise
more
notes
given
are
emended from observations
names
and local
'
India
Orientalis',
name,
have been
of species
labels,
botanists.
guarantee the
I
can
names
not
have been
since
more
or
and Indonesia respectively. Some
these
are
known
Genera and
each
to
be
or
lower
usually
from
generalised
species in
taxa
these might be of
correct
spelling
less adapted
Malay
currently
species excluded
taxon,
on some
western
scant
Malesia could
the field.
in
use
from literature and
Malay
for
labels and in local floras. Notes
on
Vernacular
as
similar
sequence
treated after the corresponding
are
out.
Short
be
is
names
enumeration of specimens
by collectors'
detailed information
are
varieties.
or
possible.
geographical
list, arranged
enumerations of
of unknown localities within
Specimens
of untraced localities
geographical
as
south. Islands
to
larger island.
first, those
far
Geographical
distinguished, species, subspecies,
known and in this order. The
as
and from north
the mainland or
area
edition,
far
as
west to east
taxa
arranged by geographical units, political units, administrative units,
are
by localities,
from
briefly indicated.
is
taxon
given under the lowest
are
names
used
to
the
are
some
or
mentioned
use
to
as
far
foresters
of the
application
as
known
and ethno-
names.
Only
spellings, presently used in Malaysia
mentioned by others have been discarded,
for non-Ochnaceous plants.
for taxonomical
or
geographical
reasons,
are
discussed
the end.
at
An index
For
practical
respective
to
effectively published
reasons,
taxa
as
no
references
accepted
here.
names
are
of
given
genera
to
pages,
and lower
but only
taxa
to
has been added.
the numbers of the
BLUMEA
20
VOL.
No.
XVI,
1968
i,
OCHNACEAE
Ann. Mus.
DC.,
Plant.
Paris
Intr. Nat. Syst.
Meisn., Pi. Vase. Gen.
129;
(1846)
Walp.,
175;
Ann.
Engl.,
357;
diagr.
1
Le
371,
(1903)
&
170;
(1902)
16
D.-T.
60;
1 —4,
Tiegh.,
v.
in E.
213 —215;
Ann.
Sc.
f., Arch. Neerl.
Fam., Nachtr.
4
(1915)
Siph.
3
Ex.
Nat.
ibid. ed.
2,
f.
Pollen
Chalk,
&
Mete.
141;
Erdtman,
Morph.
Anat.
282; Takhf., Evol. Angiosp. (1959)
Melch. in Engl.,
(1966)
illeg.
39—55.
Type
—
Syll. Pfl.
(1904)
19
93
(1846)
—96.
5
B,
12,
Pi.
(1829)
(1840)
Ann.
2
Ann.
68.
Nat.
Type
—
Bot.
to
racemose,
(1870)
Tiegh.,
P., Nat. Pfl.
&
—338,
2
(1926)
1
f. 76,
340,
ed.
(i960)
77;
(1959)
1
2,
1195 —1198;
248, apud DC.,
t.
Ann.
Nat. Plant.
genus:
VIII,
fused with
stigmas free
sometimes
or
at
i—2
16
nom.
(1835)
19
50;
Sc. Nat.
(1830) 289.
II,
VIII,
‘Ordo
—
(1845)
4
Lindl.,
Bot.
t.
711;
Sauvagesia L.
(1904)
±
96.
—
with
countries
Remarks:
The
Carpels
or
Type
genus:
—
10
or
±
or
5 —10,
free,
to
Sepals
5.
caducous.
dehiscing length-
free with
ovule,
1
epigyn respectively;
or
capsules. Seeds
Leaves
many-
both; pedicels
contort,
latrorse and
basigyn
I-
polygamous).
(—15), superior,
fused,
I—oo,
small
or
large,
30 genera
and
possibly
c.
250
species through the tropical, rarely
(S. Africa).
The following
is
compiled
Compound leaves
in
flowers
Godoya splendida
in
from
my
deviations of the characters
neo tropics:
(b) Zygomorphic
Petals
branches.
without albumen.
description presented
taxa.
lateral, terminal,
basifix,
2 —5
carpel; styles
per
terete
Inflorescences
bisexual (rarely functionally
fused. Fruits: drupes, berries,
winged,
Indo-Pacific
thyreoid, bracteate;
pores.
ovules
2—c-o
either side.
base, quincuncial, persistent.
apical
Distribution: About
subtropical
on
Stamens 5 —io—oo; anthers
o—oo.
with
or
actinomorphic,
little connate
Staminodes
(a)
4
tall trees, with ±
cymose,
or
333
Fam. Fl. Pi.
Planch, in Hook., Ic. Pi.
912;
(1852)
Sc.
v.
343;
flowered,
or
E.
in
in
(1909)
6
161; Decker, Phytomorphology
Bartl., Ord.
small undershrubs
very
articulate. Flowers
wise,
Fam. ed.
49; Mart., Conspectus
(1853)
simple, glabrous; midrib prominent
a
18
(1904)
Jack.
Woody plants,
or
Aschers.
Fam. Fl. Pi.
Traite Bot. Syst.
(1964)
2
108,
104,
Hutch.,
290;
(1902)
8
Vester, Areale Angiosp. Fam. (1940)
(1950)
1
Hutch.,
53—67;
Soler.,
Tiegh.
178—189; Gilg
Pfl.
(1912) 166; Gilg
1
745;
stipulate,
free
v.
416; ibid.
405,
157—161,
Engl., Syll.
Schnitzl., Iconographia
Fam.
Gen. Pi.
Tiegh.,
v.
131 —138;
149—153;
161—204,
(1907)
(1925)
(1935)
‘Ordo Sauvagesiae’
(1846) 60;
Euthemidaceae
Euthemis
(1901)
Eichl., Bliiten-
13;
Gilg, Festschr.
315;
5
Emberger,
200;
ibid. ed. 3
343;
—
Endl.,
Sauvagesieae’
Walp., Rep.
12,
(1895)
I
(1873)
4
Ochna L.
genus:
Sauvagesiaceae Dum., Anal.
Veget. Kingd.
Ill
21
(1952)
ed.
Fam.
Gomphiaceae
—
(1903)
219 —226;
Dicot.
Pi. Tax.
&
2
(1902)
16
IX,
(1908)
178; Wettst., Handb. Syst. Bot. ed. 4,
73,
VIII,
Nat. Bot.
Sc.
203;
3,
15
t.
6
1141;
591 —595;
H., Gen. Pi.
&
B.
474;
1—-28,
Bartell., Malpighia
Gen.
H.,
&
(1874)
2
37,
P., Nat. Pfl. Fam.
&
Nat. Bot.
P., Nat. Pfl. Fam., Nachtr.
Hall,
(1853)
3
(1840)
(1846)
5
Nat.
(1836)
2
Bull. Mus. Hist. Nat. Paris
113, 181, 208—212;
33,
548, 549; Ann. Sc.
97—117;
E.
Lindl., Veget. Kingd. ed.
Gilg
Syst. Anat. Dicot. (1899)
381,
Gen. Pi.
Endl.,
47;
Ord.
Bot. cd.
Decne., Traite Gen. Bot. (1868) 369; Baill., Hist. Pi.
257 —262;
in Morot, J. Bot.
(1837)
2
Bartl.,
735;
Nat. Syst.
Planch, in Hook., Lond. J. Bot.
474;
Leop.-Carol. Akad.
Nova Acta
(1878)
2
(1849)
Maout &
(1824)
1
(1830) 136;
Bot.
(1837) 66; ibid.
1
Lindl., Veget. Kingd.
(1862) 316;
398 —410; Prod.
(1811)
17
(1830) 383; Lindl.,
Luxemburgia
Planch.
St.
Hil.
own
observations
mentioned
occur
on
the
in the
Kanis :
A.
(c)
Sepals
io
Blastemanthus Planch.,
in
Ochnaceae
the
of
Area
Iniio-Pacific
sepals and petals
3 —6
in several American
(d) Caducous sepals
21
in Elvasia
DC.
genera.
Herbs in Sauvagesia L.
(e)
KEY
I.
the
of
revision
A
Stamens
Leaves
with
apical
2
Fruits
i-seeded
1—J,
Stipules intrapetiolary
pores.
flowers, usually thyrsoid,
Stamens
3.
GENERA
drupes
on
swollen
a
ovaries
12—00;
simple
sometimes
united.
Inflorescences
with
(1 —)j—00,
cymes.
Leaves
(3 —)5—io(—15); embryo straight.
without
intra-marginal
an
2.
1.
Stamens
10; ovaries
opening
with
flowers in
Stamens
Carpels
5.
in fruit. Leaves
longitudinal
2
conferted,
2 —5,
alternate,
5.
Ovary 5-carpelled,
S.
Ovary
curved. Leaves with
embryo
5;
with
Anthers
cvd
Ochna
I.
nerve
3.
torus.
( Ochnoideae-Ochneae).
opening
remote
±
THE
free.
Carpels (3 —)j—io(—15),
10
—cv>.
distichous
Anthers
2.
TO
fused.
not
Fruit
Stipules
a
distinct
free.
or
3
intra-marginalnerve
berry
torus
appearance
Brackenridgea
2.
capsule;
Gomphia
3.
of umbelloid
Inflorescences
more
or
.
distinctly enlarged
not
(Sauvagesoideae).
Anthers
berry.
a
of
more-seeded
distichous
5-celled. Fruit
slits.
clusters
cymose
a
opening by
1
apical
(Euthemideae).
pore
Euthemis
4.
i-celled.
2—3-carpelled,
Fruit
Anthers
capsule.
a
opening
by
longitudinal
2
slits
(Sauvagesieae).
6.
Ovary 2-carpelled.
6.
Ovary
Seeds
7.
all
with
distinct
without
Fruit
opening
branches
6.
anthophore
Indosinia
anthophore.
with
valves.
3
Inflorescences
simple,
compound
or
the rachis
bearing
a
varying
number
of
but
bracts,
only
Inflorescences
terminal
Staminodes
9.
10,
in
Staminodes 00, in
9.
Staminodes
10.
Neckia
(or pseudo-axillary by sympodial growth), many-flowered.
whorl.
1
more
Petals
Fruit
than
of inner
respectively.
whorl
as
long
8.
subglobose
whorl,
1
as
those
of the inner
spatulate,
10,
the
whorl
alternatingly
with
Seeds
winged.
Fruit
much-branched
opening
with 3
linear, equal.
Petals 3 —4
X
v.
Tiegh., Bull.
204,
405;
18
Leop.-Carol. Akad.
6
(1895) 138;
(1925)
21
illeg.
—
Trees
to
63.
shrubs.
strongly
i
on
a
1
10—00
Remarks:
with
DC.
Inflorescences
Schuurmansia
Exalbuminosae’ Engl.,
‘Exalbuminosae’ Gilg in E.
cymose
in
or
&
(1902)
16
Nova Acta
P., Nat. Pfl. Fam. 3,
P., Nat. Pfl. Fam. ed.
&
Phytomorphology
16
(1966)
2,
twin.
45,
1
or
or
but
distinctly petioled;
thyrsoid.
more
The African
treated
nerves
Flowers bisexual.
whorls,
free.
Carpels
even
±
straight
Petals
5
—10.
(3 ——io(—15)
beyond fruitfall.
Fruits:
gynophore. Seed exalbuminous.
The tribe Ochneae
were
—
Sinia
sepals.
the
Schuurmansiella
persistent style.
Ann. Sc. Nat. Bot. VIII,
ovule; style gynobasic, persistent
torus
the tribe Elvasieae is restricted
Elvasia
(1902) 548;
Ochnaceae ‘ser.
distichous, shortly
Inflorescences
with
swollen
Distribution:
20.
8
as
Ochna L.
Stamens
whorl, free,
drupes
(1874)
—
H., Gen. Syph. (1903) 315; Gilg in E.
Leaves
curved.
o.
60.
nerves
OCHNOIDEAE
Exalbuminosoideae Decker,
Type genus:
or
Staminodes
in
—
&
(1903)
2
37,
D.-T.
the
long
as
ro.
Mus. Hist. Nat. Paris
ibid.
under
panicles
I.
fusiform.
3 —5
5.
longitudinal slits
SUBFAM.
188,
and
1
9.
7.
Indovethia
Fruit
larger.
sepals
Staminodes of inner whorl 25 —30,
10.
flower.
1
7.
8.
and
shortened.
axillary;
Inflorescences
a
Flowers
winged.
not
(nearly)
8.
Flowers
3-carpellcd.
to
genus
the
occupies approximately
same
area
as
the
family,
the neotropics.
Lophira
by Gilg (1895,
Banks
1925)
ex
as
Gaertn. and
representing
the Ochneae because of their exalbuminous seeds.
I
the
American
separate
would rather
tribes,
genus
related
place Lophira
BLUMEA
22
in
(a)
than
more
The
Stamens 8—2c.
(c)
Carpels
(d)
differs from
and petals
(b)
style,
fused into
4 —5,
in
fruit
tribe Ochneae in the following characters:
the
into
a
2
—5-celled
coriaceous,
a
Gen. Pi.
Endl.,
(1840)
(1902)
16
VIII,
189,
343,
195,
Nomenclator
543;
Nat. Paris
ibid.
405;
(1846)
593,
Nat. Plant.
648,
Akad.
Leop.-Carol.
(1895)
139;
D.-T.
‘Sect. Ochnaceae
DC.
47,
Ourateae
—
181—183,
Bot. VIII,
genus:
are
20,
illeg.,
189,
204,
343;
ibid.
Pi.
Linné, Spec.
(1760)
50;
mainly
ƒ, Fl.
Fl.
African
1
in South
Ann.
(1901)
&
Ind.
Br.
52,
B.
Ind.
II
in Morot,
Tiegh.
5,
P.,
Type
—
Fam.
Pi.
Mus.
Hist.
Bot. VIII,
2
Engl.,
231;
17
(1811)
410;
A., Prod. Fl.
&
523;
(1862)
5
PR
Fam.
3,
Nat. Pfl. Fam. ed.
Ochna
genus:
407,
excl.
414,
Ann.
(1907)
163.
6
2,
L.
Elvasia
(1902)
16
547—549;
39; ibid. IX, 5
i
21
—
v.
in Morot,
Ridl., Fl.
705;
(1925)
For.
67,
71;
p.p.
(1902) 196,
Wendl.
f.
quoad
(1902)
350;
Prod.
33 —36,
Sc.
—
Nat.
Type
syn.
214;
ibid.
in Bartl.
18
2
a
other
third
genera
Fl.
Br.
J.
(1824)
1
Am.
Linn.,
42,
Wendl.,
(1873) 368;
232;
6
(1895)
139;
119,
(1922)
illeg.
J.
Bot.
nom.
illeg.
Beitr.
—
—
Bot.
J. As.
Soc.
emend.; Lecomte,
v.
125;
Taxon
syn.;
11
Adans.,
Tiegh.,
Ann.
Fl.
P., Nat.
&
11;
147, in
(1902)
Pi.
Bartell., Malpighia
Discladium
16
Ind. ed.
Gen.
Boerl., Handl.
King,
36 (1962)
(1758)
7
2
(1799)
Benn. in Hook.
364; Gilg in E.
Soc. Broter. II,
nom.
err.
Bot.
2
4,
Endl.,
204;
(1893)
(1902)
1
152;
(1877)
1
Symb.
Roxb., Fl.
735;
1
ed. Burm.,
in Morot,
(1903)
&
16
4
Vahl,
Spec. Pi. ed.
(1834)
1
Burma
Ceyl.
Bot.
354;
Linn.
Hist. Pi.
Mal. Pen.
Boehm. in Ludw., Dcf.
229;
(1789)
Ind. Or.
Baill.,
Robs., Bol.
Jabotapita Plum., Pi.
8
1
Willd.,
p.p.;
Pen.
317;
8,
inE. & P., Nat. Pfl. Fam. 3,
Tiegh.
(1911)
(1754)
5
Handb. Fl.
Trim.,
173;
Gilg
ed.
509,
Kurz,
Nat. Paris
Diporidium
Bot.
Nova Acta
OCHNA
(1797)
(1875)
(1763) 364,
16
—
America.
4, 2
1
non
2,
48 —51.
Bot.
172.
also in the Indo-Malesian
area;
occur
genera
Linn. Gen. Pl. ed.
H., Gen. Pi.
(1893)
Pfl. Fam. ed.
—119,
‘Genera Ochnea’
—
J. Bot.
(1902)
8
(1907)
5
Nat.
P.,
&
&
(1811)
17
113
Sc. Nat.
Hook., Lond. J.
in
E.
45.
Paris
(1903)
Gen. Pi.
513;
Paris
W.
(1890)
Gen. Indo-Ch.
(1962)
Mus.
1
1
153;
(1753)
Schreb.,
Ene. Méth.
(1832) 643;
1142;
Ned.
Beng.
15
2
I
315;
Lamk.,
1158; DC.,
(1840)
v.
18
I.
Carey,
(1966)
16
in
410.
Ourateae
—
Veg. Kingd.
Ann.
found in Malesia and the Pacific and perhaps also in East Africa;
Gen. Pl.
epiginous
(1902)
16
ibid. IX,
in E.
Gilg
315;
Mus.
p.p.;
179.
illeg.; Gilg
nom.
Lindh,
547—549;
(1811) 406,
17
(1849)
1
(1903)
Ann.
DC.,
197;
60,
39,
Bull. Mus. Hist. Nat. Paris
found in Africa and
(1791)
and ±
cell
Aubl.
Ouratea
is
(1874)
nom.
204;
Distribution: Two
genus
per
J. Bot.
Gomphieae Planch,
—
Ann.
Phytomorphology
Engl.,
194,
384.
H., Gen. Syph.
oligandrae’
(1902)
16
2
37,
&
(1925) 63; Decker,
21
(1830)
illeg.; Walp.,
notn.
(1902)
8
(1903)
18
(1841)
in Morot,
Tiegh.
v.
‘Sect. Ochnaceae polyandrae’ DC., Ann. Mus. Paris
Bartl., Ord.
ovule
I
OCHNEAE
I.
Rchb.,
(1868)
7
Bull. Mus. Hist.
128, 181—183;
127,
1142;
Ann.
Walp.,
475;
with
ovary
i-celled, i-seeded fruit.
distinctly enlarged.
not
Tribe
(1846)
1968
I,
3 —6.
developing
Torus
No.
ovule.
i
tribe Elvasieae
Sepals
XVI,
tribe in the Sauvagesoideae, because of its non-distichous leaves and carpels
distinct
a
with
VOL.
Sc.
Bull.
Nat.
Type species: O. jabotapita L.
2
(1825)
24;
Kuntze,
Rev.
A.
Gen. Pi.
(1902)
(1891)
i
Kanis
emend., incl.
104,
in Morot,
214;
353; ibid. 18
(1903)
A revision
:
Bot.
J.
51.
of
the Ochnaceae
Ochna;
typ.
(1902)
16
the
D.
23
Mus. Hist.
Tiegh., Bull.
v.
Area
Indo-Pacific
126; Ann. Sc.
Lectotype species:
—
of
Nat. Bot.
VIII,
Nat. Paris
(1902)
16
atropurpureum (DC.) Wendl./
8
196,
Ochna
=
atropurpurea DC.
Polythecium
53;
Tiegh.,
v.
ibid. IX,
5
(1907)
Ann.
174.
Sc.
Nat.
Bot.
(1902)
VIII, 16
P. ciliatum
Lectotype species:
—
ibid.
196, 366;
(Lamk.)
(1903)
18
Tiegh.
v.
Ochna
=
ciliata Lamk.
Pleopetalum
(1903)
17
(Lamk.)
Tiegh., Bull.
v.
Ann.
97;
Tiegh.
v.
Polythecanthum
species:
Tiegh.,
Inflorescences
flower;
Sc.
=
not
turning red in fruit. Sepals
whorls, yellow.
I —2
opening with
as
many
apical
2
ovaries,
as
Lectotype
—
5,
a
(1907)
5
sometimes
Distribution:
northern
S.W.
160,
±
at
cvMii
2
bearing
base
at
±
hemispherical
or
4
in
species
Confined
to
continental
the
the
soils.
obovoid;
majority
tropical
S.
Remarks:
and S.E.
of the
distichously
with small axillary buds,
new
inflorescences;
distinctly
in fruit.
tumid and
Petals
5
ovule
(—10)
stigmas
atropous;
—3(—5), greenish,
turning
I
below
or
less
1500
to
1930,
dry
years.
of the problem, applying
The
t.
Ochna L.
genus
jabotapita
L.
(1753)
black
s.l.,
a
(1753)
is elucidated in the
genus
was
defined
the sole species
it should be
genus
protologue:
The
as
on
red
torus
and
having
as
Ochna L. and its
are
numerous
aspects
accepted here.
stamens
per
flower.
described originally and only 'Burm. Zeyl.
as
having
(L., Fl.
10
Zeyl.,
to
only collection from Ceylon
designated
as
and dry habitats
variety. The specimen depicted, however, belongs
defined
specimen of Hermann certainly belongs
it is the
far northward
analysis of the nomenclatural
an
stamens
1747,
per
a
the type collection of O.
by
same
specimen
Ochna, having
seen
flower.
93). The
cited there, together with another phrase of Burman and
Apparently
the
the
and the heterogeneity
genus
the modern rules, and his conclusions
is cited under the type
56.'
Ouratea Aubl.
fruits
of O. jabotapita
circumscription
Robson (1962) made
was
as
seasons
of this concept have caused controversial interpretations of the
in later
and
419).
Linnaeus'changing
species
of the
Mascarene Is.
altitude,
m
adapted
S.
Sahara,
Hainan,
Ceylon,
in Africa
and the
Madagascar,
areas
Asia,
species
Dispersal by birds because of conspicuous
calyx (Ridley, Disp. Pi.,
The
nerves
the margin.
filaments subterete; anthers
± united. Fruits
or
the
to
near
small,
many
torus,
whorls;
more
small,
curved
nerves
terminal thyrses with
or
scars,
Stipules
right angles
flowering and developing
5 —!o( —15),
Ovaries
short branches
of the Arabian Peninsula,
part
on poor
are
Lectotypc
—
Merr.
transverse
±
nerves,
distinct annulus of
the foothills of the Himalayan Mts. More
123.
175.
undershrubs.
greenish, enlarging and turning red
Stamens
Malay Peninsula;
Ecology:
O.
Bot.
P.
when ripe.
black
type
J.
lucidum
species:
subcoriaceous,
or
mostly lateral
cymes,
twice after
pores.
on
IX,
joining, veinlets
persistent,
±
once or
Bot.
chartaceous
pedicels filiform, articulate. Flowers with
in
163; in Morot,
(1903)
9
45.
Ochna integerrima (Lour.)
branches,
lateral, simple
peduncle
branching
Nat.
Leaves
conferted, caducous bracts, leaving
sometimes
(1903)
irregular secondary
in
joining
sometimes
terminal
Nat. Paris
18
Tiegh.
v.
the margin,
near
the midrib and
Hist.
VIII,
Ann.
Tiegh.)
united, caducous.
upward, especially
a
Mus.
Bot.
treelets with spreading
or
intrapetiolarly
near
Nat.
Ochna obtusata DC.
=
v.
thorelii (v.
P.
Shrubs
Sc.
to
This discrepancy
phrase
and plate
in Hb. Hermami.
many stamens
per
flower.
Linnaeus personally and therefore
jabotapita
L.: the type
species of the
Ochna L.
name
known
as
the species
Ochna
was
incorrectly used by
Ouratea Aubl.
previously
Consequently
Kuntze
(1891)
to
he renamed under
described in Ochna L., including
designate
the
genus
generally
Diporidium Wendl. f (1891)
its type
species.
BLUMEA
24
The
the latter
The
superfluous
a
Tiegh.
v.
nomenclatural
respective
of
synonyms
Diporidium Wendl. f.
Tiegh.
dehiscing
the receptacle.
by
Tiegh.
v.
terminal
2
(Cf. Robson,
v.
1.
Petals
5—10
by
2.
Inflorescences
2.
Inflorescences
Petals
1
many-flowered,
4.
filaments.
Anthers
mm,
group
Tiegh. (1902)
(1907)
in
are
the
all taxonomic
are
of the
species
in
Sepals
long
TO
as
anthesis
the
v.
Tiegh.
globose,
to
Polythecanthum
—
L.
jabotapita
basally inserted
±
SPECIES
sepals.
3 —7
Stamens
Stamens
than
the
on
....
to
up
1
Anthers
O.
lanceolata
2.
O.
jabotapita
at
1
Anthers
anthesis
during
ij
as
X
long
as
filaments
Sepals
more.
less.
or
Ochna lanceolata
Spreng.,
Syst. Veg.
Sepals
(1825) 596.
2
x
long
as
var.
as
pumila
long peduncles.
cm
or
most
O. obtusata
3.
g
1.
sepals.
long peduncles.
cm
filaments
as
25—50
12—25
inflexed
or
long
as
X
O.
cylindric
branched.
larger
on
spreading
2
THE
much branched.
many-flowered thyrses
during
Pleopetalum
—
Fruits
pores.
cymes
fruit
Ochna
fruit
in
spreading
in
fruit
Type:
—
or
O.
O.
Heyne
inflexed.
obtusata
usually distinctly
4.
I.
making
Tieghem
African
study the
not
Tiegh.
van
3.
4.
of African
1962a).
somewhat
2—3-flowered
Inflorescences
3.
as
A.
Tiegh.
—3(—s)-flowered, hardly
by y\— 15
15—25
the
mm, about
2—5
Inflorescences
3.
a
Discladium,
v.
placed by
v.
Type species:
—
KEY
I.
Polythecanthum
Ochna spp. I did
Discladium
—
Polythecium
—
Anthers
on
for
genus
genera.
Section
v.
and
(1903)
Diporidium Wendl. f. (1825) and Polythecium
genera
Ochna L.
name
included in his
was
The Asiatic species
of Ochna L.
synonyms
1968
I,
name.
Pleopetalum
genera
taxonomic
of Ochna L.
species
type
No.
XVI,
Tieghem (1902) incorrectly reserved the
Van
species.
or
VOL.
reflexed.
integerrima
in Herb.
s.n.
~
Rottier
O.
wightiana
Ind. Bot.
1
Thw.,
En.
(1869)
51;
233;
Pen.
(B holof?, K)
Wall,
(1840)
Ceyl. Pi.
Ic. Pi.
(1858)
I
Benn. in Hook,
Brand., Ind.
Nat. Bot. VIII,
(GL
Pen. Ind.
O.
528;
K
heyneana
W.
Fl.
Gamb.,
in
O.
W.
&
Hb.
moonii
A.
(E,
Thw.
var.
var.
lenticels.
or
2554
Stipules
K
(ovate-) oblong,
or
Fl.
—
up
Ann.
by
Wight,
166.
Wight
111.
(1842) 528;
Bedd., Fl. Sylv.
in Morot,
Type: Hb.
—
221;
(1915)
I
Tiegh.
—
=
3
(1893)
Diporidium
16
J. Bot.
470
1
(1902)
Wallich
2808
old fl.
1
Type:
(1834)
Wallich
Bot.
En.
152;
Walp., Repert.
2807A
ex
1224
1 —2
)
(1846) 653; Walp.,
Hb. Heyne
I
(1842)
(BM, E,
ex
m
mm.
18
(1903)
Ann.
52.
1
—
(1849)
Type:
180.
—
Walker
fr.
Ceyl. Pl.
Handb. Fl.
i\ (?)
5
Sc. Nat. Bot. VIII,
holo) Ceylon,
C.P.
to
(1864)
I
1
Trim., Handb. Fl. Ceyl.
Madr.
v.
152;
fl. 23-VII-1818.
/3 Thw.,
(err.
4—8
166.
Fl.
(1834)
1
Walp., Repert.
223;
524;
105;
359.
(Travancore),
Hook., Lond. J.
moonii Trim.,
treelet,
(1891)
(1902)
16
(1915)
Tiegh.,
Hook.
Thwaites C.P.■
Shrub
v.
1
old
holo, P) Cuttalam,
O. walkeri Planch, in
2g$
Or.
t.
(1875)
Gamb.,
129;
1
1
IX-1818.
Ind. Or.
Pen.
(1840)
A., Prod. Fl. Pen. Ind. Or.
&
Madr.
Diporidium walkeri
Br. Ind.
f, Fl.
Gen. Pi.
Rev.
1
Drury, Handb. Ind.
70;
127; Ann. Sc.
holo, K)
Ind. Or.
(1906)
Trees
wightianum Kuntze,
Cuttalam, fr.
A., Prod. Fl.
&
W.
ex
172;
Ind. Or.,
Ceyl.
1
1
(1858)
(1893)
70.
—
234,
O.
p.p.
wightiana Wall,
excl.
typus.
—
ex
Type:
Hb. Gardner (K, P, PDA holo?) Ceylon (Jaffna), fl.
high,
Leaves
much branched.
with
1 —3
(ovate-) lanceolate, i|—7 by f—2-|-
mm
cm,
Branchlets
covered with
long petiole;
lamina ovate,
rounded to acute, sometimes
A.
Fig.
Distributional
4.
obtusata
var.
acuminate
slightly
I—4
simple, with
Torus
persistent.
3—5
(—7)
by
f
to
up
mm.
mm
by
8
2807A
Wight
Distr.,
Kattalaimalai:
Barber
Ceylon.
(err.
C.
Trimen
488
P.
(K),
(1893).
ex
—
(K);
(E);
East.
(K);
Hills,
Wight 184
Hb.
(BM,
Heyne
Wallich: 28o8
K.D. 395
319
s.n.
ex
Hb.
=
Palni
Hb.
Brodie
1224)
up
5,
1
mm
rachis
together;
the lower
high,
mm
5
2—8
0.
mm
obovate-oblong,
style
0.5 —0.7 mm;
to
cm,
base,
at
subcoriaceous.
or
close
2-J
including
7!—TO
long,
5—mm
long branches.
5,
by
2—5
in fruit
mostly
Fruits
mm
Sepals
filaments; anthers straight,
long
mm
2—4
on
up to 2
are
cited localities.
slightly emarginate
to
up
Petals
mm.
(3—)5 —7,0.7 —I by
Wight 470
Syamalay:
Hb. Rottier
with
fruit
in
,
3—5
South Peninsular India,
Madurai
(K);
25 —50,
in fruit
from
chartaceous
base,
The westward
area.
gamblei and pumila
I —2,
mm.
Wallich
Travancore : Hb.
Distr.,
Stamens
0
in the Indo-Malesian
of varieties
flowers, standing
long,
25
slightly generalised
towards
cm
Area
Indo-Pacific
sometimes
acute,
separate
j\ —10 by
areas
areas are
to
mm
long; stigmas mostly
6
Distribution:
INDIA.
—3(—5)
I
high, r§
mm
Ovaries
mm.
10
to
up
1
c.
ovate-oblong,
to
rounded
apex,
long; pedicels filiform, f—2
mm
ovate
at
the
of
(sect. Ochna)
the outlined
denticulate, subentire
margin crenately
Inflorescences
Ochnaceae
of Ochna
species
uncertain;
partly
the
of
revision
their border districts. Other
in
only
A
map of the
of O. obtusata is
extension
:
Kanis
Ceylon.
E,
—
2162
Mundanthoru: Barber
Walker
Gardner
K.D.
295
394
(E, GL,
(E, K),
(K, P);
Prov., Trincomalee:
2836 (DD);
(K);
3175
K,
fide
Twhaites
Hb.
ex
Madras
(CAL),
—
North.
Trimen
C. P.
Dimbham
Tirunclvely
3222
L), Wight K.
(E).
1347
Mannar:
2iojB
Madras, Nilgiris Distr.: Wight
Nelhimalai: Fischer
Tope: Auglade
=
Wallich
K, P),
(GL, K).
D.
(K);
393
—
Ghat:
Kerala,
—
; Coimbatorc
Lushington
Beddome
Kuttalam:
s.n.
(BM);
ioSj
Heyne
s.n.
Ill
(K).
Prov., Jaffna:
(1893).
1221
Distr.:
(K).
(K)
s.n.
N.W.
(BM, CAL, K).
Thwaites
C.
P.
2554
Prov., Puttalam:
—
?,
Nangala:
fide
Alston
(K).
Ecology:
In the drier lowlands and in low
jungle
on
ridges
up
to
1200
(?)
m
altitude.
Flowers in May and June.
Vernacular
names:
Ceylon:
bo-kéra
(Singh.), katharai (Tamil), fide
Trimen
(1893).
BLUMEA
26
Remarks: In O. lanceolata
different
as
there
Possibly
I had
since
to
species
base
flowering
conclusions
material
merged
to propose
some
entities that have been described
old
same
be extremely
to
populations.
collections
I
rare.
did
lanceolate leaves and slender branches.
mediate. Plants from Ceylon have
rather sclerophytic:
are
predecessors,
enough
see
the
good
plants
with small,
thick,
narrowly
have rather
collection from Travancore is about inter-
A
more
less
or
oblong leaves and comparatively long
anthers. Future studies should be made on ample material in order
of the different characters
Unfortunately
my
not
branches. Plants from Tirunelvely Distr.
stout
as
infraspecific division. Generally speaking,
any
from the Coimbatore and Nilgiris Distrs.
bluntish leaves and
1968
i,
between regional
about the
on
material appears
recent
more
No.
XVI,
time.
minor differences
are
my
I have
Spreng.
some
at
VOL.
in relation with
to
check the constancy
differences in habitat and distribution.
The original description of O.
lanceolata Spreng. (1825) is rather short and incomplete.
1
It
described from
was
Malabar
wightiana Wall,
'foliis
ex
lanceolatis
description
are
W.
pedunculis
Sprengel's
the Indian collection of Ochna
heyneana
O.
material
from K, I found
spp.
very
and there is
correct
for the
mark
to
the
&
species
dispersed
Cuttalam in
It is
name.
a
of Forskaol
whether that species
of O. lanceolata
of this
material
that fits
1818
among
Sprengel's
material
type
Spreng. have been named
under
no.
a
is
true
Ochna: O.
(Fl. Aeg. Arab.,
(Forsk.) Schweinf.
inermis
204)
1775,
conspecific with the
certain resemblance and it is
one
probable that
Africa. Planchon
spp. in East
walkeri Planch, from
as
the
O.
Ceylon,
(1846)
parvifolia
and Vahl
to
treated here. The
both
17
(1811)
species
also pointed
at
a
I
1,
is
30,
original
the
Bot.
t.
an
with the
agree
on
same
414,
have
question
a
described from
decide
(Symb.
parvifolia
O.
2807A and added
later synonym for Euonymus inermis Forsk.,
Candolle's description and plate in Ann. Mus. Par.
Ochna
Sprengel's
the collector
as
duplicate of the
a
conclusion of Wight &Arnott (1834), that it is hardly possible
a
characters
of
subsolitariis'
treated here.
part
Yemen, which is indeed
descriptions
conspecific with
be
(1834). The
A.
sheet that originates from Rottler's
a
near
O. lanceolata
burned in Berlin, but
was
quite probable that this is
Originally Heyne's collections
Vahl. Wallich
only
can
W.
Lamk.,
lucida
O.
conspecific. If
doubt about its identity. Therefore I accept O. lanceolata Spreng.
no
name
well. It is
as
He mentioned Rottler
too.
type
herbarium. It bears material collected by Heyne
description
Indian species
(= pedicellis!)
this direction
in
material.
(1834) and
A.
&
crenulatis,
pointing
donor of his
or
the
DC., that I regard
ex
different from O. obtusata DC., than it
Spreng, is really
O.
from
distinct
as
O. obtusata DC., and O. nitida Thunb.
1790,
f.
2.
33)
for De
true
There is
affinity with
some
resemblance between O.
Vahl from Yemen, and O. atropurpurea DC.
from South Africa.
There has been confusion in literature about the
as
3
different elements
Beddome
(1869) pointed
and Trimen (1893)
the latter
are
seems to
combined under this
at
a
aware
by
me to
represent
is
error
1224.
caused
fi
by
a
a
is
very
1224
probably
Wall,
ex
cited under the species (=
W.
a
form of O.
var.
(1858),
O. obtusata
Bennett
&
DC.).
(1875)
A., although
two
typ.)
varieties of
is considered
hand, the collection
C.P. 2554
lanceolata Spreng. Possibly Bennett's
specimen of Thwaites in K, labelled O. moonii and numbered C.P.
is
very
different from the collections undei the
and K, but similar
very
wightiana
form of O. obtusata DC. On the other
is
This specimen
BM, BO,
It
var.
O. moonii Thw.
also undei
of the considerable differences between the
O. mooniiThw. The collection C.P.
cited under
(see
relationship with ‘O. squarrosa’. Contrarily,
cited this species under O.
be
identity of
name
probable that
the
to
one
two
in
same
number in
P.
specimens
in K and P
were
wrongly numbered and
A.
in fact represent
Kanis:
A
I have
Colebrookii’
named O. stricta
Wallich
( nom
under
nud.).
.
the
to
suggested by
as
obtusata
of O.
Ochna
2.
sheet in K
same
2,
Pi. ed.
(1799)
For.
2
4,
(1762)
1
Selsk.
Mcth.
(1854)
collections
definitely from
(1762)
Pi.
510,
=
(1891)
{non Thunb.
- Discladium planchonii
105.
quoad
L.; Trim.,
16
J. Bot.
(1902)
Thw.,
(1869)
Hermann
(Hinidoon
Ceyl. Pi.
En.
(1902)
127.
—
VIII,
by
4 —5
oblong,
or
18
koenigii
i^—2
2
i
25
mm
pedicel;
c.
(obovate-)oblong,
by
3
—6 mm,
3—6 by
fruit
up
±
$—I
to
10
Distribution:
3455
O.
non
Kurz,
lucida Lamk.,
Not. Pi.
As.
Morot, J. Bot.
(1846) 650,
5
4
16
(1868)
7
523,
Ann.
K,
in
err.
rufescens
BO,
351.
err.
emend.
Lectotype:
Bedd., Fl.
543;
O.
syn.
Tiegh.
v.
Sc.
Nat.
PDA
P,
—
(E, K) Ceylon, old fl.
1015
Ann.
Tiegh.,
v.
(BM,
in
(1902)
16
(1875)
1
Diporidium
—
cm,
\
z\—5
to
s.n.
mm
7 —10
tapering
long,
high, i|
by
at
3
ex
long.
Ceylon.
in
fruit
squarrosa
in Morot,
Bot.
VIII,
holo?)
16
Ceylon
Fruits
0.5
up
mostly
1—2
(C,
5,
by
per
up
cm
to 2
acute at
long,
up
to
to
3
1768.
Stipules
obovate-
main axis
reduced
1—-6
mm 0.
Sepals
-lanceolate,
to
mm
5,
7 —10
long filaments; anthers
mm
mm;
or
the lower
high,
fl.
base, margin
many-flowered;
mm
I—2
0.4—0.5
flower,
holo) India,
lamina mostly
to
holo)
Sc. Nat.
with lenticcls.
obovate-oblong
with
P
P
162; Ann.
times branched
3
to
12—25,
—0.6
Vahl
petioles;
to
161; Ann.
(1903)
9
(1903)
9
compound,
up
Petals
Paris
quoad distr. (E p.p.,
apex, obtuse
at
0, in fruit
mm.
base. Stamens
Ovaries 6—io,
Nat.
p.p.
Hb.
long
mm
dichasial,
mm
—4!
471
blackish, covered
acuminate
or
mono-
Hist.
Wight
Mus. Hist. Nat. Paris
Koenig
with
acute
cm
mm.
mm
Mus.
Hb.
Type:
—
chartaceous. Inflorescences
pedicels
persistent. Torus
234.
C.P.
Type:
—
branches
long;
44.
Leaves
—4
spinulose-denticulate,
6—
(1893)
Bot.
VIII,
Walp.,
70;
Linn. Spec.
Rottb., Dansk.
non
—
Linne,
squarrosa
fr.
Walker
Paratype:
(1858)
1
Tiegh., Bull.
45.
mm.
by
—
Bot.
(1962) 48—51;
11
Griff.,
non
Tiegh.
v.
Ceylon,
Nat.
much branched. Branchlets
treelet,
5 —io
suggestive
Diporidium squarrosum Kuntze,
Hook., Lond. J.
in
Tiegh., Bull.
v.
(1903)
v.
(1903)
18
Griffith, affixed
obtusata DC.;
Merr.
illeg.;
—
squarrosum
Polythecium rufescens
Discladium microphyllum
Shrub
O. wallichii
fl.
Corle)
Sc. Nat. Bot. VIII,
Bot.
1
Thwaites
Type:
—
Discladium
sp.
are
similar
very
to
Willd.,
41;
O.
=
(Lour.)
nom.
Hook. /, Fl. Br. Ind.
Handb. Fl. Ceyl.
369.
and
K)
more
O.
—
120;
(1824)
I
mult.
Merr.
(BM)
Ann. Sc.
Tiegh.,
Benn. in
51;
(Lour.)
DC.) Planch,
ex
v.
Ouratea spec.
auct.
Linn.,
syn.
Walker 67 in Hb. Hook. (K) Ceylon, fl.
3
belong
Taxon
Robs.,
513;
=
integerrima
Discladium
—
Hb.
&
545,
O.
=
205
p.p.
Lectotype:
—
rufescens
(E,
from Hb.
illeg.; Burm., Fl. Ind. (1768)
O. integerrima
1
O.
Griffith
not
separate fruits
div.
auct.
(1783)
2
(1877)
I
(1902)
Sylv.
Hb.
different collection, being
a
Moon, Cat. Pi. Ceyl.
p.p.;
Rev. Gen. Pi.
O. nitida
from
are
(1753)
I
732, &
nom.
731,
N.S.
illeg.
nom.
125.
was
stipulacea
O.
name
is of unknown origin, but it is all
The
(1875).
Bennett
1158,
Skrift.
(1797)
4
464,
in
Wallich is numbered 802 (BM) and named ‘O.
Some
lanceolata Spreng. It does certainly
are
I
2,
Fl. Br. Burma
Enc.
27
ft being clearly
var.
(BM, E, K) bearing the
2806
no.
jabotapita Linné, Spec.
Spec. Pi. ed.
Vidensk.
of
Area
DC.
Linné, ihid. ed.
non
Indo-Pacific
2554, the type collection
(nom . nud.). This material
and closely resembles O.
Planch,
the
description.
Colebr. (ttom. nud.). One sheet from
nov.
of
herbarium material from the Botanic Garden in Calcutta that
some
seen
dispersed by
partly
C.P.
duplicates of
accordance with its original
the Ochnaceae
of
revision
style
8
by
5 —7
6
mm
mm.
long,
in
28
BLUMEA
Fig.
c.
X
5. Ochna
jabotapitaL.
inflorescence,
3;
g.
stamen,
x 1½; d.
x
3;
h.
—
a.
flower:
Branchlet
1
fruiting
sepal,
2
flower:
VOL.
XVI,
with inflorescence
petals,
2
and
No.
and
I,
young
some stamens
sepals detached,
x
1968
2½
leaves,
detached,
(
b.
x½;
X
Walker 75,
3;
e.
E).
mature
sepal,
X
leaf,
3; f.
x
½;
petal,
A.
CEYLON.Fraser
in Hb. Vahl
(E, P),
(K).
s.n.
—
Prov., Hiniduma:
Walker
67 (K),
C.P.
part of
everwet
correctly indicated
a
(K);
Ochnaceae
(E, K),
Macrea
(E, U),
(K);
Pelawatta:
P).
?
—
on
as
for
West Indian
a
the Asiatic and African varieties of O. jabotapita L.
in
commonly
from
that certainly
Ceylon,
The
Herb.
Wight
obtusata DC. and does
fit
not
belonging
to
Ochna
Tiegh.
v.
obtusata DC.,
J. Bot.
O.
Ann.
99;
squarrosa
(tton L.) Rottb.,
Fl. Ind. ed. Carey,
(1834)
1
8, f.
t.
syn.
Beng. Pi.
i
212,
Suppl.
3;
1;
non
jabotapita
411,
(1846)
Nat.
(1903)
specimen
belongs
was
found
mixture of material
Prod.
n;
51.
Paris
—
L.
O.
to
must
species
Van
L.
pi.
50,
45.
a
jabotapita
problem
a
—
(1824)
(1902)
16
(1903)
9
1
735;
Discladium obtusatum
Sc. Nat. Bot. VIII,
O.
&
Ceyl.
(1903)
Brand.,
f.
129;
309;
Or.
2,
x
2
351.
—
164; in Morot,
Type: in Hb. DC. (G holo)
Ann. Mus. Paris
597;
Griff,
Not. Pi. As.
v.
Tiegh.
4
196.
Corom.
x
(1797)
(1811)
(1854) 464,
in Morot,
1
Benn. in
J.
1
(1858)
(1864)
220;
(1906)
6;
(1840)
1
Dalz.
70;
&
Bedd., Fl.
Hook./, Fl.
Gibs.,
Sylv.
&
1
(1875)
Diet. Econ. Prod. Ind.
5
(1891)
237;
I
For.
(1902)
196; Prain,
Fl. Bomb.
&
Gamb., Fl. Madr.
(1795)
510,
411;
1
Sind
quoad descr.,
p.p.
illeg.
Bot.
16
1
(1936)
Cat. Pi.
I
(1824)
nom.
&
Say.-Din,
Ceyl.
1
(1824)
O. integerrima
(1902)
illeg.; Illustr.
t.
472,
735; Spreng., Syst. Veg.
(1846) 649; Walp.,
=
220;
1
(1915)
89.
t.
Prod.
J. Bot. 5
nom.
Das, Fl. Assam
grandiflora Moon,
O.
—
3
Br. Ind.
128; Talb.,
(1910)
t.
p.p.;
A., Prod. Fl. Pen.
111. Ind. Bot.
Wight,
Ceyl. Pi.
545,
(1799) 1158,
&
W.
Burnt.;
37;
(1783)
2
4, 2
169; Benthall, Trees Calc. (1933) 97, fig.; Kirt.
(1921)
Planch, in Hook., Lond.
dium lucidum
En.
Ch. Nagp.
(1938)
17
N.S.
Cooke, Fl. Bomb. I,
233;
Ind. Trees
4
Skrift.,
rufescens Thw.; Watt,
(1933) 518; Kanj.
40
syn.
(1839)
(1874) 60;
(1893)
I
Haines, For. Fl.
&
excl.
p.p.
Handb. Ind. Fl.
For. Fl.
Type: Roxburgh, Pi.
DC.,
(1825)
The
DC.
description concerned.
89; Willd., Linn. Spec. Pi. cd.
t.
Cat. Pi. Bomb.
lucida Lamk., Enc. Meth.
O.
18
been
collections
on
O.
GL
this
to
which bears
(1962)
11
Mus. Hist.
Bot. VIII,
based
was
affinity with
A clue
(1811)
Taxon
125; Ann.
17; Drury,
O. lucida Griff.
J. Bomb. Nat. Hist. Soc.
f.
17
O.
for
(1762)
It has
of Walker's
herbarium in
own
to
(1842) 528; Thw.,
1
Brand.,
165; Haines, Bot. Bih.
—
Paris
(1832) 643,
2
Grah.,
Basu, Ind. Med. Pi. ed.
41.
Tiegh.
v.
Dansk. Vidensk. Selsk.
62,
Handb. Fl.
Trim.,
(1909)
(1795)
152;
(1861)
50,
excl.
523,
439;
1
69; Walp., Repcrt.
Bomb. Fl.
(1869)
vernacular
a
L.
L. The material of the latter
belongs
(1902)
16
Sc. Nat.
Roxb.,
t.
is also
illegitimate.
is
(1962)
fl.
Pi. Corom.
172,
also
Tiegh., Bull.
v.
Roxb.,
Ind. Or.
and May.
April
squarrosa
on two
close
a
Greville)
jabotapita
Aim. Mus.
Bot.
J.
obtusatum
(1903)
17
O.
South.
—
L.
description.
Herb.
(ex
(1825) 596, Robs.,
2
in Morot,
Ind. Or.,
epithet that
it fits in much better with the
material in P
Spreng., Syst. Veg.
Pleopetalum
in E
O. obtusata DC. and O.
Ceylonese origin and
microphyllum
with the
in
corresponding sheet
3.
(L).
(DD, E).
remark under Ochna.
see
name
(1753)
jabotapita
sheet in Wight's
in another
Tieghem's original
O.
to
and suggests
47',
propr.
However, the corresponding
be of
an
description based
belong
of Discladium
original description
labelled
100
s.n.
471 PP-
discussed under O. obtusata DC. Planchon
DC. is
ex
emended De Candolle's
erroneously
Wight
for the species that should be correctly named O. obtusata
use
of O. nitida Thunb.
status
de Silva
Flowers in
lectotype:
of Ouratea. The later
species
Hb.
different entities. Robson
result this East Indian species of Ochna is named with
name
(BM); Koenig
s.n.
Worthington 6604
Hakkinda:
very
in Hb. Hermann
29
(CAL, L, P);
s.n.
low altitudes.
at
Area
Indo-Pacific
1278
139
based
was
the
of
‘zeyl. 9' (BM); Jonville]
Hermann
1015
Ceylon
(1753)
specimen
a
the
(BM, BO, K,
3455
L.
As
168
(E),
75
of
West Prov., Kalutara:
Thwaites
In the
Ecology:
A revision
:
(K); Gardner 167,
74
(C, P);
Kanis
125, Ann.
Ann.
(Lour.)
Sc.
x
(1849)
Merr.
—
Nat. Bot.
2
179;
Discla-
VIII,
16
(1902)
ibid,
351;
(1903) 164;
9
VOL.
BLUMEA
30
Sonnetat in Hb.
Type:
O.
Thunb.,
nitida
Ann. Mus. Paris
(1823)
L.
thecium nitidum
Type:
O.
224;
Tiegh.,
v.
Buch.-Ham.
pumila
Hook., Lond. J.
60; Benn. in
Beng. Pi.
237;
Bot.
(1903)
1
Gamb., Fl.
Trees N. Beng.
Fl. Assam
(1929)
(1936)
1
Kirt.
30;
—
nana
Buch.-Ham.
W.
ex
Ann.
Sc.
Hb.
Buch.-Ham.
Hb. Buch.-Ham.
Sannyashikata,
Gorakhpur,
Nat.
O. collina
Edgew.,
India
Thw.,
cordata
Brand.,
175;
Nep. (1825)
(1894)
50;
146; Prain,
(1903)
1
(1874)
For. Fl.
W. Ind.
&
Ber. For. Circ.
(1933)
2, I
(1916)
(1902)
16
VIII,
373.
221.
—
(1834)
—
Type:
—
Dalz.
152;
Wallich
Gibs.,
&
Wallich
Lectotype:
Paratype:
—
Das,
105.
fl. 30-III-1802.
Nepal, Terriany Forest,
i
&
(1891)
1
99;
Cow.,
&
Kanj.
517;
Rev. Gen. Pi.
Sakranda,
20,
fl.
3761/1
3761/2
ex
(1846)
1
Engl.,
non
43.
Jahrb.
Bot.
(1902)
30
Bhyrubpur.
Type: Edgeworth 336 (Kholo)
—
IV-1844.
(1858)
1
nud.;
nom.
Hb. Moorcroft (K),
ex
Ceyl. Pl.
En.
—
13-IV-1809.
Trans. Linn. Soc.
(Saharanpur)
Bot.
(1864)
1
54.
128; Haines, For. Fl. Ch. Nagp. (1910)
7-IV-1809.
Wall. Cat.,
ex
Wallich 3762
Specimen:
—
N.W.
O.
(K),
(K),
humilis Buch.-Ham.
O.
354.
(1849)
A., Prod. Fl. Pen. Ind. Or.
&
O.
Poly-
—
(1903)
18
sp.;
=
(1842) 528; Planch,
1
Nairne, Fl. Pi.
Basu, Ind. Med. Pi. ed.
&
Bomb. Fl. (1861) 46; Drury, Handb. Ind. Fl.
ex
125.
Don, Prod. Fl.
736;
Diporidium pumilum Kuntze,
Tiegh.,
v.
180
fl.
Descr. List N.
Buchanan Hamilton in Hb. Lambert (BM holo),
O.
(1902)
16
(1902) 368; ibid
16
1
(1849)
1
Swartz,
Ouratea
=
Fl. Kum. (1927) 81; Cow.
170; Osmast., For.
(1921)
2
220.
pumilum
Polythecium
524;
(1906)
(1915) 166; Witt,
1
Or.
&
Ind. Trees
Brand.,
309;
—
DC.,
ex
non
251;
513
Ann.
Walp.,
Morot, J. Bot.
Ann.
(1875)
1
nec
(1797)
4
(1902) 196; Duthie, Fl. Upp. Gang. Plain I,
I
Madr.
Haines, Bot. Bih.
Br. Ind.
Thunb. (1918)
Méth.
(1824)
1
45.
Thunb., Fl. Ceyl.
735;
165; Walp., Repert.
(1839)
1
(1824)
I
prob. Ceylon,
(1846) 652; Walp.,
5
Hook. /, Fl.
Bomb. I,
in
Prod.
DC.,
ex
Ene.
Sc. Nat. Bot. VIII,
111. Bot. Him. Mts.
Royle,
Cooke, Fl.
Ann.
Tiegh.
(G holo) Ind. Or.,
in Hb. Delessert
Juel, Pi.
597,
(1903)
18
nud.; Thunb.
nom.
132,
Prod.
\
(1846) 650;
5
v.
(1794)
illeg.
Lamk.,
nec
Mus. Hist. Nat.
Or.
16
now.
(1825)
2
(1788) 67;
Discladium nitidum
—
Ind.
Ups.
12,
Planch, in Hook., Lond. J. Bot.
jabotapita
in
Ac.
Tiegh., Bull.
v.
Ann. Sc. Nat. Bot. VIII,
99;
(P holo)
pi.
412,
Syst. Veg.
(1903)
17
1968
i,
Pleopetalum lucidum
—
Mus. Nat.
(1811)
17
Spreng.,
7;
43.
Lamarck
Veg. Ind. Occ.
Prod.
non
(1903)
18
in Morot, J. Bot.
No.
XVI,
70,
409.
—
O.
L.
squarrosa
cordata
var.
Benn.
in Hook./, Fl. Br. Ind. 1 (1875) 524; Trim., Handb. Fl. Ceyl. 1 (1893) 233.
Diporidium
—
cordatum
in Morot,
Tiegh.
v.
Ann. Sc. Nat. Bot.
O.
excl.
moonii
syn.
Bedd.,
Fl.
wightiana
Sylv.
1224
moonii
v.
A.
Polythecium pedunculatum
(L,
P
—
127.
Type:
—
Polythecium cordatum
Thwaites C.P.
1222
Tiegh.,
v.
Gardner
Hb.
ex
(1858)
1
=
70, p.p.
O.
excl.
moonii Trim.,
=
O. lanceolata Spreng.,
Ann.
Walp.,
Handb. Fl. Ceyl.
Handb. Fl. Ceyl.
Ann. Sc. Nat. Bot. VIII,
16
6
i
(1902) 369.
7
(1868)
(1931)
(1893)
—
42.
234,
Type:
543;
—
p.p.
O.
—
Thwaites
holo?) fl. 1833.
Tiegh.,
1223
v.
fj
var.
jabotapita L.;
ex
Ann. Sc. Nat. Bot. VIII,
Hb.
Tiegh.,
Gardner
Ann. Sc.
(BM, K,
16
P
(1902) 369,
nom.
nud.
—
holo) fr. 1853.
Nat. Bot. VIII,
16
(1902)
373.
—
Type:
holo, S) India, Gurhwal, fl.
Polythecium kingii
v.
Tiegh.,
(P holo) Saharumpore,
Polythecium
var.
PDA
v.
Thwaites C.P.
Falconer 333
(1902)
Alston in Trim.,
51;
Tiegh.,
thwaitesii
16
369.
Moon
sensu
&
(BM, BO, P,
Polythecium
Specimen:
L.
W.
ex
Bot.
(Jaffna) fr.
(1869)
3
J.
(1902)
Ceyl. Pi.
En.
squarrosa
Wall,
Polythecium
C.P.
Thw.,
O.
16
holo?)
PDA
(BM, K, P,
VIII,
discolor
v.
Bot.
Ann. Sc. Nat. Bot.
VIII,
16
(1902)
374.
—
Type: King s.n.
Gard.
Tiegh., Bull.
Mus. Hist. Nat. Paris 9
(1903) 158;
Ann. Sc.
Nat.
A.
Bot.
(1903)
18
VIII,
fr.
Quilon,
Kanis
Type:
—
54-
of
revision
the
Ochnaceae
Wight
162
the
of
Indo-Pacific
K.D.
=
Area
31
(E, GL, K, L,
392
holo,
P
S)
VI-1836.
Discladium dalzellii
(1903)
18
Bot. VIII,
A
:
Tiegh., Bull.
v.
44.
Mus. Hist. Nat. Paris
Type: Dalzell & Stocks
—
(1903)
9
(BM, K, L,
s.n.
161 ;
Ann. Sc. Nat.
holo, S) Canara,
P
fl.
Discladium leschenaultii v. Tiegh., Bull. Mus. Hist. Nat. Paris 9 (1903) 162.
Pleopetalum
—
leschenaultii
(1903)
v.
fl.
Ceylon,
&
Sc.
fr.
Nat. Bot.
(1903)
v.
Hort. Bot.
ex
O.
fl.
Calc,
Stipules
elliptic,
mostly
a
fr.
&
Cameron
by
(obovate-)oblong
or
finely
or
a
base. Torus
(3 —)5,
by
ovate
i\—15
0.6 —1.2
mostly
in
A.
in
lbs.
a
per
long
is
based
on a
a
by
20
with
style
by 71
by
8—12
India,
to
up
on
i—5
or
little
a
or
of O.
status
S.
c.
10
—
4
c. 0.5
by
—
descr.;
(K) Kurnool,
i\—8
mm.
5
Petals
Stamens
10 mm
or
many-
15 —25
filaments
30—'7o( —120);
mm.
Sepals
0.
obovate,
5 —io,
in fruit up
or
sometimes dichasial
high,
mm
more
or
cymes
apex
acute
emarginate, margin
—3-fl° werec l
up to
cm,
mostly
or
lamina
petiole;
Ovaries
to 20 mm
m,
new
to
5—10,
long.
Fruits
in rather
sandy
or
soils.
rocky
Ceylon.
dry jungle, scrub,
Flowering
and
shortly
leaves: in North & Central India from February
July.
as
used
an
antidote
as
(' sitka '),
an
to
snake-bite;
also for
a
decoction of the
consumption
and asthma
ornamental in tropical Asia and perhaps
1938).
squarrosa
an
States, East Pakistan,
generally
=
L.
moderately
± 0.82
kg/dm
hard
and
synonym
of O.
in
jabotapita
of O. nitida Thunb. has been uncertain
Thunberg's
originating
from
ever
concept of the
the Cape
weight
.
(1762) has been commonly
illegitimate
1794.
close-grained;
3
use
L.
for this species
(1753).
since this
species
Juel (1918) enumerated three collections under that
as
Upsala,
17
holo)
166.
1916, 34,
s.n.
base
obtuse,
2
mm,
Himalayan
1200
1890)
nudum in
nomen
(P
(1915)
1
long
mm
mostly monochasial,
poor, sometimes
(Watt,
it is
J. Bot.
210
mm.
Reddish-brown,
name
Pleopetalum
III-1891.
(3 —)5 —18
subunguiculate.
or
fr.
Bengalore,
0, in fruit
mm
Kew Buil.
nud.;
Lectotype: Beddome
Inflorescences
—
The plant is
wood:
mixture,
name was
17
holo)
(?) high, deciduous. Branchlets slender
obtuse,
used by the Santals
time, but
herbarium in
on
10
cubic foot
as
Gaudichaud
Gamble, Fl. Madr.
ttom.
—
Leaves
mm.
10
base
(Sayeedud-Din,
of
The taxonomie
published
The
mm;
Campbell).
Africa
(P
(not seen).
(obovate-)lanceolate,
Ceylon from May
Remarks: The
for
10 m
rounded,
at
—
in Morot,
164;
little flattened; anthers (3 —)6 —12 by 0.4—0.7
3 varieties in
root
166,
94.
given in certain menstrual complaints
Properties
51
2—3
during development of
or
is
(Rev.
128;
(1903) 163.
9
Type:
—
1881
(1926)
to
high, i\—3
From sea-level
Use: The
also
a
to
3 —5, up
April,
(1906)
—5-flowered branches,
grassland vegetations
before
root
I
tapering
0.4 —0.8
Distribution:
open
47.
(Mysore)
up
ovate-oblong,
long,
by
Ecology:
to
with
mm,
mm
ig—3
9
(1903)
(1915)
subcoriaceous.
—1 mm
to
(1903)
little acuminate, sometimes rounded
denticulate,
flowered thyrses
at
1
11
(K)
600
treelet,
or
little attenuate, sometimes
less
18
Trees
Fl. Madr.
3 —8(—16)
acute
J. Bot.
s.n.
1837.
Brandis, Ind.
ex
Undershrub, shrub,
stout.
in Morot,
164;
Type: Leschenault
—
Mus. Hist. Nat. Paris
Hist. Nat. Paris
Rec. Bot. Surv. Ind.
al,
Paratype:
—
Mus.
Nat. Bot. VIII,
beddomei Gamble,
O.
(1903)
9
(1903) 46.
Brandis, Madras, Kambakum Hill,
D.
Calder &
fl.
Sc.
gamblei King
Type:
Nat. Paris
18
Tiegh., Bull.
v.
Tiegh., Bull.
Ann.
99;
VIII,
1820.
Discladium gaudichaudii
gaudichaudii
Mus. Hist.
Tiegh., Bull.
99; Ann.
name
was
name
of Good Hope and from
was
probably
from his
Ceylon.
validated by De Candolle (1811), who based his description of the species
material of Indian
or
Ceylonese origin
in Hb.
Delessert, that might
have been named
BLUMEA
32
by Thunberg (not seen).
DC.,
O.
as
lanceolata
smaller leaves.
It is
It
based
not
collections of O.
on
make
to
jabotapita
later homonym
a
mentioned that Lamarck
book
and
of the
the
gave
Thunbergian
'Thunb., Prod. Pi.
same
DC.
ex
wrongly composed
Cap. 67'
as
in
synonym
nomenclatural
no
In this
(1788).
the author. As
citation
together
misinterpreted again
was
incorrectly cited
is
has
(1811)
nitida Swartz
giving Thunberg
This
has
much
the original
opinion.
my
description (1846),
as
it
was
consequences
from the
with his
correct
De Candolle
result,
a
should be
it
context
cited the latter West Indian species
(1797)
species.
doubt from
probable
O. obtusata
or
area,
L.
of O.
erroneously
page,
(1811, 1824)
most
Planchon's emendation of the
The position of O. nitida Thunb.
since it is
decision beyond
a
O. obtusata DC. is the
accept
1968
i,
the only other species of Ochna in the
possible
description and plate, but
Therefore I do
No.
XVI,
be conspecific with either O. jabotapita L.
must
Spreng.,
not
VOL.
description
own
in Index Kewensis, where
the original publication of O. nitida
as
Thunb.
The
Don
O.
name
(1825)
but
collection,
specimen
Lamarck's
is
(1854)
Griffith
without
name
is
considered
is
Finally,
O.
Burma
at
was
some
Inflorescences
Leaves
I2
2.
—
Leaves
2^ —5
1.
time
a.
var.
mm
from
as
an
v.
0
tuberous
v.
Tiegh.
Shrub
—
ex
or
—
Tiegh.
moonii Thw.
Type:
near
—
to
ij
of
chartaceous
belonging
is cited
obtusata DC.
lanceolata Spreng, (see
O.
as
O.
to
a
but this is
own
homonym.
species
s
synonym,
collection
The variety [3
under that species).
the
probably
by
and later authors.
(1877)
Kurz
not occur
in the wild
likely, however, that
jabotapita
O.
or
the
From
same as
L.
cm
THE
Shrubs
or
with
many
distichously conferted,
subcoriaceous,
mostly
acute
to
acuminate
at
obtuse
or
rounded
at
apex.
long peduncles,
O. cordata Thw.
obtusata DC.
P. thwaitesii
v.
var
without
v.
—
s.s.
—
Tiegh.
O.
—
D.
grandiflora
Polythecium
annulus.
Moon
var.
gamblei
v.
var.
pumila
Tiegh.
—
O. lucida! Lamk.
—
v.
succulent,
Undershrubs, sprouting
gaudichaudii
discolor
obtusata
-
Branchlets rather
c.
Discladium dalzellii
O.
*
b.
cm
Branchlets
apex.
tip
3—7
caducous
treelets.
stembase
—
introduced
was
VARIETIES
long peduncles,
scars.
or
TO
glaucous, coriaceous,
on
and the
ornamental species.
a
cymes
own
my
Burma
in
state
t^le
near
simple
up
annulus
whitish
obtusata
leschenaultii
O.
0
waxy,
on
broad
waxy,
mm
Inflorescences
yearly
a
not
i
2
latei
a
Merr.
as
with
conspecific
Moon
me
listed for Burma
thyrses
bracts, leaving
2.
is
name
of his
one
is made up of three different elements. The type
regarded by
KEY
i.
his
so
on
of O. jabotapita L., whereas Griffith
synonym
mountains of Assam. It is
adjacent
in
Lamarck and
to
several
accepted by
as
This
in BM.
preserved
Lamk.,
by
same
the holotype.
as
now
description, based
new
conclude that O. obtusata DC. does
I
designated
not
the
on
L.
‘O. squarrosa’
studies
(1858)
sensu
be
and
(1824)
possibly based
are
with O. lucida
completely
integerrima (Lour.)
being
squarrosa
jabotapita
O.
should
synonymous
a
illegitimate
an
1224 is
as
not
reference
a
O. moonii Thwaites
Thwaites C. P.
is
De Candolle
by
descriptions
the Lambert herbarium and is
in
gave
with O.
conspecific
Both
Candolle's specimen
De
O. lucida Griff.
later authors.
validated
was
several floras.
in
originally
was
collections,
Buch.-Ham.
pumila
stated
as
Tiegh.
—
P.
D.
—
kingii
Tiegh.
Hb. Linti. (G holo?)
Jnd. Or.
treelet with slender branchlets,
long. Leaf-blades variable, mostly
acute to
i|—2\
mm
0
near
acuminate at
apex,
the tip. Stipules 3 —8
mostly
mm
acute to attenuate
A.
at
Kanis
A
:
revision
the
of
Ochnaceae
base, margin finely denticulate, chartaceous
compound,
rachis
many-flowered,
in fruit
up
anthers
6 —12
to
long, the lower \—1
cm
4
long,
mm
than
more
2
x
long
33
Inflorescences
not waxy.
pedicels
Petals
persistent.
cm
as
long;
cm
Area
Indo-Pacific
subcoriaceous,
or
| —4(—6)
the
of
Stamens
5—10.
long,
cm
—
30 —6o;
the filaments.
as
Distribution: India, East Pakistan, Nepal, and Ceylon.
INDIA.
Naka:
S. Konkan:
Bombay,
Goa: Dalzell
s.n.
Talbot
K.D.
Res.:
Latham
17567 (BM);
Koenig
(BM).
s.n.
Bourne
3496 (K);
Barber
1678
(K).
Chuta?:
(K);
DD,
(BM,
Mooney
Andhra
—
2162
(K).
—
Parikud
Hamilton
Tripura,
Agartala:
Pakistan.
NEPAL.
(K).
—
Sober Isl. :
(DD).
—
(BM);
(K);
670
Hb.
Kahasah:
(DD);
Haines
Prov., Karaitivu:
Worthington
1202
(DD),
former
Hb.
Surv.
Haines
(BO, P);
Ecology:
From
and
Himalayan
sea
level
Clarke
Brodie 119
Thwaites
to
up
names:
India.
narole, ramatanachampaka
P.
yerra-juvi (Teligu).
—
champa-baha
Apart from data
the works of Watt
fide
1200
also
m
Trimen
Bombay:
(1893).
(BM);
—
7116
(CAL);
Chit-
—
(BM,
East.
Jaffna:
P), Dyke
K,
Prov.,
s.n.
Trincomalee,
Dpt. Agr.
Perad.
s.n.
Chempianpattu: Simpson 8004
434
(BM).
the
especially
country,
foothills
Orissa:
Rather
—
of
the
Central
too.
on
Andhra Pradesh: sunari,tammichetta,
chilanti,
herbarium sheets,
and Kirtikar
(1891)
variable
found in
of the respective localities.
in
sellendi
yerra-jammi,
(Uriya).
—
&
appearance,
shape and
explained
(Tamil).
those of local floras
Basu
(1933)
especially
that
were
size of the leaves
as
On the other
caused
hand,
by
India. I found it impossible,
the island.
Only
consulted.
and,
likely
to
can
a
some
conspecific.
lesser extent, of the
ecological
from the
to
where
as
that it is caused
however,
studies in the field
from
here
regarded
differences in
it is
used and especially
were
Ceylon,
in
are
by the separation of the marginal Ceylonese populations
on
muda (li),
(Santali).
It could be
in Peninsular
Mysore:
—
(Salem Dist.), panjaram,
buin-champa, kaniari, nobinisworo, pata-champa
is
variability
(Konkani).
Madras: padalakkonai
This
33
2380
(CAL, P).
s.n.
North. Prov.,
—
Batticaloa:
—?
southern
kanak-champa
—
have been described in the past
taxa
(P).
s.n.
Hb. Gardner
altitude in hilly
the
in
separate species
specific
Haines
Jenkins s.n. (BO);
(BM, E).
ex
Belihuloya: Worthington
about
(Kanarese).
Ceylon, mal-kéra (Singhalese),
pool
Gamble
(E).
1223
Worthington 875
Point:
and
shengodu, sherundi, shilandi (Tamil).
flower parts
(K);
5526
Mts.
Vernacular
Remarks:
17086
R.: Haines
Sonakalla:
Assam:
Clarke
(BM); Sylhet:
2421
(E);Leschenault
C.
Puttalam:
Simpson 9846;
Foul
6818
Cowan
Dhankuta: Stainton 77
(BM, K),
Ghats
Eastern
F.
Haines
Burhait:
(K);
—
Gamble
Chandka
State:
Panighat: King’s coll.
Sabaragamuwa Prov., Balangoda: Worthington 3766 (BM).
Western
a
(E).
Karaka:
Mooney 389
(CAL); Khandpara:
3
823 (DD);
2380
Watt 258
(BM),
ïoSi
(K);
5523
(K);
1535
Bailadilla:
Draper
Athmallik
Econ. PI.
in
Beddome
Coast:
Godavery, Gorge:
Barber
(K);
170
Puri:
R.: Haines
Burio:
(K);
Bassu
Chatterjee s.n.
(BM);
(BM);
3
(DD); Adijar:
s.n.
Coromandel
(K);
(CAL);
4129
(K);
s.n.
(DD); Salem, Ariyur
Barnes
s.n.
Lawson
J.:
2124
Travancore,
(CAL).
Nugattene Gap: Simpson 8466 (BM);
Bihar:
b
2380
(BM, BO, P);
Gardner
Fischer
(K);
14220
Barumi F.
(E); Silrope Jaldi:
4047
Shutu
Russellkonda:
Dhenkanal,
Bengal, Dacca, Kashimpore:
1222 ex
Madras:
Mooney
Dampier
(CAL);
5527
Gauhati:
(E);
Thwaites C. P. 1224
N.W.
Orissa, Ganjam,
(BM); Banbripore:
Debbarman
East
(K);
V.
19595
(E); Vishakhapatnam, Ragupaliem:
s.n.
3
Kaudahs
Narayamasamy
Fernandess
Koenig s.n. (S) ;
(DD, K); Chingleput:
s.n.
8850
Bastar, Chitrakot:
Surv.
Parganas:
s.n.
Bazar: Sinclair
C. P.
N.
Kuicholi:
Haines
Amlekhganj: Brough 564
Ceylon.
Thwaites
1243
—
(DD);
Pi
Econ.
Ranger
N.
Ish: Carter 517
Santal
Bihar,
Kamrup:
E.
(DD).
s.n.
(DD); Maharajpore:Kurz
tagong, Cox
Cleghorn
Kalapathar:
Mayurbhanj, Lulung:
9323
Hills:
Belta:
Karwar:
(DD);
140
48 (CAL).—
(13M, K,L, P, S);N. Kanara,
Malabar Coast:
Madras, Nilgiris,
—
Deccan: Cooke
1667 (P);
Stocks s.n.
Braganza
Kerala,
—
Ponnachi
For.
Hills:
Madhya Pradesh,
Krishnan
K);
(DD, K),
Pradesh, Nellore, Krishnapatam:
N. Circars:
—
(K);
209
(DD,
5525
Kolli
Dalzell
Kanara: Dalzell &
k
(DD);
s.n.
Kambakkam
Donald
K),
116
Talbot
Naidoo
(DD, K);
s.n.
Mysore,
(E, GL, K, L, P, S).
392
—
Coimbatore : Kistnasawmy
Gamble
(DD, K); Vuyala:
s.n.
—
(K); Chandwar:Ritchie 1667 (K).
162
Quilon: Wight
(K).
Anshi:
(DD);
s.n.
Talbot 116
(CAL),
Dalzell
Hb. Hook.
ex
at
centre
distinguish
elucidate the
circumstances
least partly
of the
gene
useful infra-
matter
further.
BLUMEA
34
this variety is
Perhaps
VOL.
found in the Upper
not
ornamental is has been collected from the botanic
dabad,
b.
gamblei (King
var.
King
Shrub
treelet of
or
crenate,
compound,
to
5
of
the northwest
to
stat. nov.
in
wild
a
As
state.
Calcutta, Lucknow,
an
Mora-
Lahore in West Pakistan.
as
O. beddomei Gamble
—
the lower
Orissa, Ganjam, Goomsur:
(K); Godavery:
Nallamalai Hills:
2
O.
—
a
gamblei
Beddome
1079
(BM).
Petals
5
8.
0
rounded
base, margin faintly
whitish.
pedicels
—
to
Inflorescences
2—4
long,
cm
Stamens 35 —7°; anthers
the filaments.
(DD);
Dhaba:
Andhra
Perakonda:
(BM);
1082
long;
cm
at
or
mm
—5
obtuse
India.
9556
—
glaucous
waxy,
as
Madhya Pradesh?,
—
Beddome
long
as
Duthie
Bjalputti:
little emarginate
persistent.
Peninsular
branchlets,
stout
(obovate-)oblong,
i|—2(—4)
cm
X
and NE.
(K).
2945
\—f
than
more
Chanda,
Haines
sometimes
racbis
with
appearance
subentire, coriaceous,
Central
Bombay,
Nandhigaw :
s.n.
Gangetic Plain
gardens
long. Leaf-blades
rounded,
or
or
cm,
long,
mm
Distribution:
INDIA.
succulent
mm
many-flowered;
up
5\—8-J
3 —4
mostly obtuse
denticulate,
fruit
Brandis) Kanis,
ex
slightly
Stipules
the tip.
at apex,
in
far
as
1968
I,
Brandis.
ex
near
Saharanpur, and
Dehra Dun,
No.
XVI,
58 (DD), Haines
10058
2945,
(DD, K);
15849 (K);
Ramaswami
Kurnool:
(K).
2944
Pradesh, Karimnagar, Mahadeopur,
Gamble
Hooper s.n. (K); Nellore, Murrenikinda:
Duthie
Wardaguran:
Donald
Beddome
-
Edwards
s.n.
(K);
1269 (CAL); Anantapur, Kotlakota:
Gamble
Brandis
15246 (DD, K), Bingham 15260 (DD); Cuddapah: Beddome 1085 (BM); Nallamalai Hills:
(K); Horsleykonda: Gamble
s.n.
Fischer
4376 (CAL),
Madras :
Gamble
Ecology
Thick
From
:
bark,
adaptation
are
to
found
low altitudes
each
Fischer
Kambakam
to
up
in the
Only
were
of these
Gamble (1915).
to
other collections
L and S
c.
are
much
was
of his
CAL
Cameron
s.n.
var.
very
(Tamil?).
or
as
as
the
type
—
Quilon
under
more
that
that
are
of
suggestive
a
-
better
Andhra Pradesh: kuka-movi
—
for
name
who
some
gave
waxy.
&
A.
—
I have
the type collection.
not seen
ex
and
this material,
DD.
description
collection. In
in K
was
by Beddome, Cameron, and himself
of his
K
only
new
two
species
O.
beddomei,
of the collections
the specimen
name.
me
without
mentioned
as
lectotype.
named O. gamblei King by Brandis according
concerned has
the
However,
Occurrence of the
O.
of Gamble's
several localities
collections is mentioned there, however,
suggestive of O. obtusata DC.
pumila (Buch.-Ham.
W.
(K).
Bombay: sonari-chattu (Marathi?).
by Brandis (1906)
probable, because of the rather
ex
600
(E).
obtusata. Stands of both varieties
var.
King's manuscript
own
with the
along
I admit
rather thin and not
not
Buch.-Ham.
v.
at
K.D. 392 from
to
are
Fischer
named by himself. One of these is bearing flowers and was chosen by
Wight
is
leaves
waxy
mentioned several collections
from different localities
any
one
be regarded
to
but probably it is kept
Gamble (1916)
(K),
inland stations of Peninsular India.
in
Madhya Pradesh: rakat-rohan (Hindi?).
species.
2561
other occasionnally.
consequently this has
indicating
Bourne
previous
India. Madras: koorykaly
names:
Remarks: Originally Ochna gamblei
new
m,
16545 (BM, K); Chittoor, Horsleykonda:
Mysore, Bangalore:
—
Hill:
1350
collections in CAL. It has been validated
for the
(K).
s.n.
and coriaceous,
branches,
dry habitats than
near
—
(DD); Chingleput,
stout
Vernacular
(Teligu).
s.n.
Gamble
(DD); Ballipalle:
15144
Palmaner:
(K);
s.n.
more
a
superficial resemblance
duplicates of this collection
var.
in
obtusata. Besides, all the leaves
succulent
var.
gamblei
in Travancore
moist climate there.
DC.) Kanis,
stat.
nov.
pumila Buch.-Ham.
—
ex
O. collina
DC.
—
Edgew.
—
O.
nana
Polythecium pedunculatum
Tiegh.
Undershrubs with slender stem, mostly
30—60
cm
high, only
branching
at
base,
A.
from
yearly
sprouting
Kanis
obovate-oblong,
A
:
revision
woody,
obtuse
mostly
distinctly denticulate, herbaceous
peduncles
not
long
as
(2 —)3 —y(—8|)
branching; pedicels
the filaments
as
or
Distribution: Central N.
INDIA.
Uttar
Pradesh,
tuberous
roots.
at
Choki:
23580
Inayat
21683 (DD).
Saharanpur,
Sakranda:
Bira: Hearle
(K); Panna,
s.n.
(K); UparGhatJashpur Ra.:
Pokhuria:
(CAL);
37
(E).
W.
1818A
(K), 1819B (DD),
Ramshaihat: Haines
s.n.
438
Satpura Hills:
Hooker
Ecology :
From
Vernacular
Remarks:
300
names:
O.
mm
long, about
States.
Garhwal: Falconer
Edgeworth
86 CP
Ranger
309 k
Tundi:
A.
s.n.
Harsukh
Haines 2942
Mooney
A.
Biswadih:
Campbell 8466
Gamble
653B (BM); Champasari?:
Assam:
—
754
Bihar, Dhanbad: J. Campbell
—
Parker s.n.
(DD);
(BO, DD),
Fischer
309K (DD); Jalpaiguri:
(DD).
927
Chakia:
Daibhar:
Bhumka:
(DD); Surguja,
S);
Bahraich,
(E);
1242
P,
21683b (DD);
(K); Chhindwara, Khapa:
Gamble
Lall
Hira
Mukerjee
(DD, K);
s.n.
Hamilton
CL,
333
Dudwa: Harsukh
Campbell 8409 (E);
(GAL); Darjeeling:
Hamilton
s.n.
to
up
900
it here
as a
easy.
Mts. in
Nepal
obtusata.
The
but their
flowers
Mann
31
(BM).
higher,
jungle, (stunted) Dipterocarp
in
in
hilly
country:
S. of the Upper Gangetic Plain. Coming
India. Bihar:
champa-baha
couple
specimens
foothills of the
yearly after the
up
two
years
that the
typical characters
measure
certain genotypes. Finally
are
successive
named
by
me
seasons
plants
O. obtusata
inside the
and
area
can
a
appearance
to
the cool
Certainly
not
with
be
there is
a
from
Lower
generative
for
hereditary,
a
burning
is
certainly
noticeable selection for
are
also better
in the northern hills.
for Peninsular India
Gamble (1915)
Burma is also
'O. integerrima (Lour.)
in the
thereof. It is also possible that
all dwarfed forms of O. obtusata DC. do belong
Kurz
by
hazards for
protected garden. Perhaps
possibility that the pyromorphic plants
winter climate
var.
pumila,
of the plants is completely
in India and it could have caused
(1894), Cooke (1902), and
fruticulosa
to
DC.
var.
only be given
from fire and other
consequences
will be found
of
their inflorescences and
obtusata. Proof
var.
pyromorphic
variety has been erroneously recorded
specific
that
the result of the morphogenetical
of the
that it
that has been found
Steenis, Fl. Mai. 1,4,1934, P- XXXVII).
and by growing from selected seeds in
eventually
species by
suspicion
distinction by inflorescence and androecium
natural stand of pyromomorphic
a
separate
strong
collections from the foothills of the Himalayan
those in
to
a
(Hindi).
a
phenomenon
concerned have been collected
much similar
old agricultural
Nairne
J?)
van
as
a
as
by ecological conditions and that the morphological characters
are
adapted
also
on two
Stainton
have grown during
very
of
564,
(see
simalkata
regarded
habit. I have
obtusata DC.,
however,
separate variety,
(Brough
it will be found
caused
(Santali),
DC. has been
ex
form of O.
My evidence is based
stems
are
protecting
some
Lambert
and possibly
m
pumila Buch.-Ham.
pyromorphic
a
is rather
parts
Hb.
grassland vegetation. Mainly
open
also in species of several other families
treat
in
authors because of its conspicuous
previous
only
O.
2-J —4
fires.
jungle
an
Sathia: Nat.
(K); Sarashwatipur:
Mts. and hills
Himalayan
a
cymes;
(K).
s.n.
(= sal) forests, and
I
but
faintly
( —4)-flowered
the longer, without scales,
(DD); Kunjara: Mooney809 (K).
(E);
7714
Khera:
Thompson
1819E (CAL); Kurseong:
Buchanan
Terriany:
SIKKIM.
is
margin
(BO).
NEPAL.
all
long. Leaf-blades
mm
—3
anthers
Inayat 21683b (DD, K);
Doutulpur
Bashi Ram 57
Campbell
A.
2
stem
21683a (DD); Gorakhpur:
(DD); Mandla,
Tarai: Modder
Bengal,
—
the
at
70 —120;
35
base,
at
Edgeworth 356 (K);
Kheri:
Kankraha: Harsukh
E.
8—16
acute
India, Himalayan
NE.
&
Madhya Pradesh,
—
Stamens
Area
Indo-Pacific
little longer.
a
Inayat 21683c (DD, K);
(DD, E);
the
Stipules
apex,
the lower
long.
Pilibhit, Bargad: Inayat 21683a (DD, K);
Chandan
acute
of
chartaceous. Inflorescences
or
cm
the Ochnaceae
to
long,
cm
1 —4
of
Merr.
as
by
already
different and is
to
var.
Dalzell
noted
&
pumila.
Gibson
This
(1861),
by Talbot (1909).
regarded by
me as
con-
BLUMEA
VOL.
No.
XVI,
Ochna
4.
ed.
(1790) 338; ibid.,
(L,
29
=
Willd.
O. wallichii Planch, in
(1793)
Beng. II,
Fl. Br. Burma
412.
(1901)
15
Diet. Econ. Prod.
Burk.,
Gagnep., Suppl. Fl.
(1891)
1
105,
(1902)
16
VIII,
(1935) 265,
2
24,
II-1919.
’,
439;
Cochinch.
Para-neotype:
—
incl.
Prel. Rep. Pegu
Ann.
(1849)
1
stipulacea Colebr.
(1875) App. A,
Col. Frang. (1886)
(1906)
179;
nud.;
nom.
App. B, 35;
30,
304,
Watt,
727;
Craib, Fl.
412;
&
(1963)
71.
Discladium wallichii
(1931)
Das, Fl. Assam
1
(1936)
(BM,
Trees &c. ed.
Tiegh.,
Ann.
Sc. Nat.
A.
holo) Burma, Amherst,
K
3
Rev.
wallichii Kuntze,
v.
—
Soc.
1
Diporidium
—
R. As.
Siam. En.
(1946) 674; Hundl. & Chit, List
Wallich 2804
Type:
128; Ridl., J. Str. Br.
(1911)
2
(1935) 1569; Kanj.
2
1
normale.
var.
Util.
Trees
Mai. Pen.
—
O.
syn.
King, J. As. Soc. Beng. 62, II (1893) 231; Bartell.,
Gen. Indo-Ch.
351.
(1846) 650; Walp.,
5
excl.
524,
140;
Dansk Bot. Ark. 23
Lars.,
Bot.
J.
(1875)
Brand., Ind.
155;
219;
Gen. Pi.
fl.
vang
mai
Pi.
205; Laness.,
(1891)
5
244;
42;
N.S.
Neotype: de Pirey in Hb. Chevalier 41165
—
Beih. Bot. Centralbl. 28,
(1911) 83; Hoss.,
(1961)
Soc.
integerrimus Lour., Fl.
Elaeocarpus
—
Lond.
1
(1875)
44
(1877)
1
Diet. Econ. Prod. Ind.
Malpighia
Hook,
Br. Ind.
f., Fl.
Soc.
Kurz, J. As.
Bot.
205.
Chevalier 41180 (P) Annam, Long Quang Tri, ‘bong mai do’, fr. V-1919.
Benn. in Hook,
59
(1943)
holo) Annam, Long Quang Tri, ‘bong
P
de Pirey
For.
Trans. Am. Phil.
(Lour.) Merr.,
integerrima
Masam., Fl. Kait.
emend.;
1968
i,
36
3-II-1827.
O.
Planch, in
brevipes
Benn. in Hook.
42.
(K
—
O.
Hook., Lond. J.
Br. Ind.
(1875)
1
Bot.
(1846) 652; Walp.,
5
Hundl.
52 5;
Pi.
Rev. Gen.
Kuntze,
Chit,
&
(1891)
1
Ann.
(1849)
1
ed.
List Trees &c.
105.
—
I 8o;
(1961)
3
Hb. Hook.
Type:
Or., fl.
As.
Not. Pi.
Griff,
Gomphia
syn.
As.
J.
Ind.
crocea
in
,
Diporidium brevipes
holo)
err.
f. Fl.
(1893)
II
Soc. Beng. 62,
1614; Kew Bull.
(1935)
(1854)
4
-
2 32
318,
quoad
Madamaca Pator:
1835, fructifer
O. lucida
(non Lamk.) Griff.,
Not. Pl. As.
in Hook,
f., Fl.
(1875) App. A,
30.
(1875)
1
typus.
tant'
Br. Ind.
in
1
(1871)
II
40,
(187s)
525;
Diet. Econ. Prod.
Burk.,
Griffith ‘Mergue.
Type:
—
49,
King,
(1935)
2
Ad littoram
(not seen).
(1854) 464, 111.
4
err.
syn.
O.
'ad litoram
Griffith,
Type:
—
523,
/, Fl.
Ouratea crocea
—
p.p.
maris Ins.
Br. Ind.
J. As. Soc. Beng.
463; Kurz,
Benn. in Hook.
Jack;
sumatrana
t.
605, f. 6,
maris
illeg.;
nom.
Benn.
L.; Kurz, Prel. Rep. Pegu
squarrosa
prope
Amherst: Feb.
1835'
(not
seen).
parviflora
O.
in Hook.
'in
syh'is
O.
140;
Griff,
Rep. Pegu
List Trees &c.
ed.
(1872)
295,
descr.; ibid.
For. Fl. Br. Burma
1
Hook,
ƒ,
Fl.
Fl. Br. Burma
44
1
205.
—
Type:
1
Diporidium
(1891)
105.
—
H°;
(1875)
non
&
(Pegu),
wallichii
parviflora
var.
155.
—
Bonn,
Type: Griffith,
As. Soc. Béng. II,
Burma
33,
nom.
II
in
Bot.
(1877)
1
Pi.
(1963)
1
nud.; J.
205;
As.
&
126.
Soc.
(1875) App. A,
(1875)
44
Hundl.
Beng. II,
30,
App. B,
41
35;
(CAL holo, K, P) South Andaman, fl.
Kurz
41,
Gilg,
s.n.
(1872)
syn.
Jahrb.
List Trees &c.
ibid.
295;
O.
pumila
33
ed.
44,
ex
(1904) 238,
3
(1961)
(1875)
II
Ham.
nom.
42.
140;
Benn.
DC.; Kurz, For.
—
illeg.; Brand.,
Type:
Scott
s.n.
fl.
Kuntze
(NY holo)
(Planch.)
Polythecium
(1870)
err.
524,
Chit,
J.
74;
Prei. Rep. Pegu
Type:
—
(1877) 206;
Kuntze
15
(1901)
Nair, Fam. Burm. Fl.
Soc. Beng.
Diporidium wallichii (Planch.)
105.
wallichii
O.
—
Fl. Br.
For.
35;
Andam. Is.
128; Hundl.
holo) Rangoon
464.
Bartcll., Malp.
Andam. Is. (1870)
42;
(1875)
As.
Br. Ind.
Ind. Trees (1906)
(CAL
3
(1961)
Rep.
(1877)
fruticulosa Kurz, J.
(1854)
524;
App. B,
(1875)
O. andamanica Kurz,
in
(1875)
4
(not seen).
squarrosa («0« L.) Kurz, Rep.
Prei.
Chit,
O.
i
Mouhnein
prope
Pi. As.
Not.
Fl. Br. Ind.
f„
helferi
var.
Kuntze
v.
brevifolium Kuntzc,
Cambodia,
Tiegh.,
var.
Rev.
Gen. Pi.
i
(1891)
N'Cor, fr. IV-1875.
longifolium
Ann.
Sc.
Kuntze,
Nat. Bot.
VIII,
Rev.
16
Gen. Pi.
(1902)
374.
1
—
A.
Type:
Kanis
A
:
the Ochnaceae
of
revision
the
Area
Indo-Pacific
between Salween
788 (K, P) Tenasserim, plain
Helfer
of
37
R. and
Gyaing R.,
fr.
III-1837.
Discladium harmandii
(v. Tiegh.) Lecomte,
f.
674,
82:
(1963)
23
Fl.
Craib,
129;
70.
Siam.
Hö
14;
&
En.
(1931)
1
Type: Harmand
—
Polythecium griffithii
Griffith
K.D.
785
Ann.
Tiegh.,
v.
(K,
P
Ann.
Sc.
—
O. harmandii
—
Fl.
Indo-Ch.
Gen.
D; Lars., Dansk
62
5
(1927)
1
(1946)
Bot. Ark.
holo) Laos, Attopeu, fl. 2-III-1877.
Sc.
Nat.
fl.
holo) Mergui,
P
351.
Merr., Lingn. Sc. J.
Nat.
Bot.
holo) Cochinchine, Poulo Condor,
762 (P
Harmand
(E, K,
1229
178, f.
(i960)
(1902)
16
75;
Suppl.
Gagnep.,
Vietn.
Tiegh.,
v.
(1911) 706, f.
I
244;
Fl.
Du'o'ng,
Diporidium poulocondorense
Type:
Ann. Sc. Nat. Bot. VIII,
Tiegh.,
v.
Fl. Gen. Indo-Ch.
Bol. VIII,
VIII,
fl.
(1902)
16
Griffith
Paratype:
(1902)
16
360.
—
1875.
374.
1088
Lectotype:
—
K.D.
=
787
p.p.
(K, P), Mergui, fr.
Polythecium pellucidum
King
s.n.
(P, U)
(L,
S.
P
holo)
Tiegh.,
v.
Ann. Sc. Nat. Bot. VIII,
Nabee
Andaman,
S.
Boh, fr. IV-1890.
(1902)
16
374. —Lectotype:
coll.
Paratype: King’s
—
s.n.
Andaman, Bajajag, fr. 29-IV-1893.
Polythecanthum
Polythecium thorelii v. Tiegh., Ann. Sc. Nat. Bot. VIII, 16 (1902) 375.
—
thorelii
fl.
v.
Tiegh., ibid. IX,
(1907)
5
—Type: Thorel 970 (C, E,
175.
holo) Cochinchinc,
P
1862—66.
Polythecium cochinchinense
thecanthum cochinchinense
K,
holo) Cochinchine,
P
Ann.
Tiegh.,
v.
Sc. Nat. Bot. VIII,
Tiegh., ibid. IX,
v.
fr.
(1907)
5
175.
(1902)
16
Type:
—
375.
Thorel
—
970
Poly-
(CAL,
1862—66.
Polythecium lefevrei v. Tiegh., Ann. Sc. Nat. Bot. VIII, 16 (1902) 375.
Polythecanthum
—
lefevrei
Tiegh., ibid. IX,
v.
Gen. Indo-Ch.
fl.
VIII-1864.
1
—
(1911)
707,
p.p.
Tiegh.,
v.
annamensis Lecomte,
Polythecium latifolium
—
harmandii
Pierre
(1911)
i
16
VIII,
707,
fr.
1875.
(1902)
p.p.
373.
O. harmandii
—
Type: d’Orléans
—
s.n.
fl. IV-1892.
Ann.
Sc. Nat. Bot.
Gen. Indo-Ch.
Type:
Bot.
Ann. Sc. Nat.
Fl.
annamensis Lecomte,
var.
578 (P holo) Poulo Condor,
Lectotype: Lefèvre
—
Tiegh.,
v.
latifolia Lecomte, Fl.
(1946) 675.
1
O.
—
Fl. Gén. Indo-Ch.
(P holo) Laos, Luang Prabang,
var.
175.
Paratype: Harmand 803 (P) Poulo Condor,
Polythecium inaequale
var.
(1907)
5
1
7036 (P
(1911)
707;
IX,
(1907)
5
174.
O. harmandii
—
Gagnep., Suppl. Fl. Gen. Indo-Ch.
holo) Cochinchine,
Tay-ninh,
Mt.
Deon-ba,
III-1866.
Polythecium pruinosum
Lecomte, Fl.
Baria,
retusa
Pierre
(C,
fl. II-1877.
var.
P
Pierre 7032
O.
O.
Ann. Sc.
(1911)
705.
Ann. Sc.
—
Paratypes:
cambodianum
holo) Cambodia,
(P)
Cambodia,
grandis Ridl.,
v.
Nat. Bot.
Type:
—
Keith
s.n.
IX,
5
(1907)
Pierre 33
—
174.
(P
O. pruinosa
—
holo) Cochinchinc,
Ann.
Sc.
5
(1907)
fr.
Soc.
1
ex
59
SING
(1911)
126.
&
harmandii
fr.
IX-1865
IX,
707.
5
(1907)
—
176.
—
O.
Lectotype: Pierre
fl. III-1870.
—
Paratype:
Ill—1870.
(1911) 83;
Bangtaphanoi,
O.
Chiang,
Fl. Mai. Pen.
1
holo) Pcrlis, Ginting Kabok,
DC.) Hundl.
(1963)
—
III-1867.
Nat. Bot.
1
175.
Lectotype: Pierre 1804 (P holo)
Bien Hoa, Bao
(P)
Dinh,
Samrong Tong, fr.
(BM, CAL, K,
Pi.
Mt.
707.
Gen. Indo-Ch.
(SING) Thailand,
Fl.
Bot. IX,
Prov. Samrong Tong, Pen Lover,
Prov.
pumila (non Buch.-Ham.
Nair, Fam. Burm.
Nat.
(1911)
Pierre 1804
Tiegh.,
J. Str. Br. R. As.
13037
1
P) Baria,
cambodiana Lecomte, Fl.
Lectotype: Ridley
Paratype :
Tiegh.,
v.
1804 (BM, CAL, K,
Polythecanthum
harmandii
7033
1
Lecomte, Fl. Gen. Indo-Ch.
Phu Quoc,
&
Tiegh.,
Cap Tiwan, fl. VIII-1868.
Polythecium pierrei
var.
v.
Gen. Indo-Ch.
(1922) 365.
fr. III-1910.
—
—
fr. 27-III-1890.
Chit,
List Trees &c.
ed.
3
(1961)
42;
BLUMEA
VOL.
Undershrub, shrub,
slender. Stipules
obovate-oblong
by
2—7
times obtuse
2—4
rachis
0, in fruit
mm
ovate-oblong,
to
obovate,
by
15 —25
by
10—16
by
4 —6
\
c.
0.4
—0.8
mostly
Fruits
ASSAM. Simons
Thakur
E.
Hill
Chand
Rup
Distr.,
Lace
238
—
(BO),
5,
J—6(—lo),
(25—)30—60
by
0.7—1.1
the longest;
outermost
0.5 —0.7
sometimes
on up
mm;
to
1
style
anthers
by
10—15
long branches.
mm
N.
Malaya, Laos, Cambodia,
Wallich
Res.:
Native coll.
Ran
R.:
(CAL).
339
Henzada
16 KL
Distr.:
Scott
Lace
Amherst: Falconer
314
311
Res.:
Hamilton
s.n.
(K),
s.n.
(K), Helfer
802
Shaik
K.D.
786,
DD,
K.D.
Wallich
L),
Distr., Bassein
789
Prov.:
Pegu
—
2803A
(K); Toungoo
Res.: Po
towards
Khant
(K),
(K,
790
P);
Amherst
Parish
16896 (S),
2804 (BM, K); Tavoy
s.n.
Tenasscrim
—
(K);
172
Distr.: Meebold
428 (CAL, L, P, U), 383 (BO, DD); Kaleinaung Res.:
Mokim
Pyu:
95» 221,
294/2 (K),McLelland
1163 (DD).
Meebold
(K),
Bassein
(CAL, K);
429
Khant
—
(E, K).
2980
Wallich
(K, P),
787
s.n.
-
Oliver
(K),
Po
(DD).
173
Distr., Pyinchaung:
Lace
Kurz
Rogers
Prov.,
(DD);
53
(BM),Maingay
Lobb
(DD),
313
(K);
7123
Mandalay
(CAL); Myankhlaing
14080, 14388 (DD); Myangone:
(BO, DD),
K).
Rogers
Labutta:
80
R.:
E,
Pakokku
Magwe Prov.,
Parkinson
Ta Ywe
(CAL, DD,
(E); Toukyeghat valley:
31
(BO, DD, SING), 316 (BO,
237
(DD);
3310
Sebjauk
Clarke
GL, K, P, SING),
E,
Distr.: McLelland
(CAL),
s.n.
Parkinson
24890a (L), 24906 (L),
(DD).
Rogers 164
—
Distr.,
Prome
Kurz
Pagodas:
Koeltz
(BO, DD, L); Cherrapunjee: Koeltz
s.n.
(BM,
2805B
Range Off.
Sampanego:
(SING);
1920
7
Parish
Kanjilal 5308 (CAL),
Khasi Hills:
(L).
Distr.,
(L), 600, 796,
490
Tura:
Hills,
3683
Prov.,
Moulmein: Falconer
14988 (CAL),
unequal, the
10316 (DD); Myaungmya Distr.,
Estate:
Gallatly
Distr.,
Stamens
mm
Sepals
mm.
Distr., Wagyaung Fts.:
(DD); Rangoon
(K); University
Prov. :
8
Akyab Distr.,
Mines
Kurz
Insein
(E);
f—I
red.
-
(K); Sylhet:
s.n.
Irrawadi
Toukyeghat,
s.n.
Garo
Tracts, Kassalong
Alejyami: Lovy
s.n.
monochasial;
in fruit. Petals
subunguiculate.
or
some-
compound,
Torus
turning dark
turning dark red
base
at
Inflorescences
persistent.
(—25)
acute,
—3-flowered,
0,
mm
10
1
mm
Pakistan, Burma, Thailand,
Distr., Pyinmana: Rodger
A67 (DD, K).
Eyre
by
11
E.
(BO, L);
Ruby
Prov.,
Yemethin
Ra.,
to
up
BURMA. Arakan Prov.,
Sagaing
4—9 mm,
long; stigmas
mm
India,
Hooker
Pakistan.
Chittagong
high,
6—io(—15),
2976 (L), 3018 (L);
Thakur
(L),
29979
to 20
up
s.n.
Chand
Rup
mm
2—8
mostly
apex,
mostly
6—20
Hainan.
Vietnam,
S.
6
filaments,
Ovaries
mm.
N.E.
:
long
mm
2—3,
Distribution
branches
-
z\ —7
in fruit
mm,
cm,
at
chartaceous.
the basal
5
tapering
8— 15 mm,
...
with
(—75)
lamina
long petiole;
mm
obtuse
acute or
long;
cm
to
up
deciduous. Branchlets
up to 45 cm,
2—5
denticulate,
to
1968
I,
rarely obovate or linear-lanceolate,
sometimes
up
dbh.
high,
m
12
Leaves with
—1|(—4)
in fruit
long,
cm
to
up
mm.
margin finely
base,
at
—z\
ovate
2—3
(obovate-)lanceolate,
or
many-flowered;
high,
by
mostly acuminate,
cm,
pedicels
treelet
or
5—8
No.
XVI,
38
Ba Pe
Badul
880 (DD); Mergui Distr.: Griffith K.D. 783 (K, P), 936 (K), 1088
K.D. 787 (K, P); Tenasserim:
=
KhanI 81
Bhamo
(CAL); Mergui
Headwaters:
State, Menyaw
E,
K);
6851
between
(E,
Salween
ANDAMAN IS.
King
Middle
(L, P);
s.n.
NicoBAB
coll.
Port
Payap
(Q; Chiang Mai,
(K, L, P),
Kasin
Sör
ensen,
Circle,
io8
343
Bunchwai
Shan
—
to
Khant
State:
Paing Kyun,
E.
11351
Abdul
(DD).
Huk
—
Lace
R.:
Beddome
Kachin
(CAL).
82
of Salween
(K, P); Thaungyin:
K.D. 788
Mouat:
Kurz
Doi
(BO, K, L,
=
—
Larsen &
Phrae:
King
RFD
P,
25965 (BO, K);
Hosseus
Larsen
& Hansen
Winit
Gram
&
(C);
199
State,
Karin
—
4586 (CAL,
1084 (BM),
Dickason
(K, L);
near
K,
P),
Fang:
Tambalu:
Mae
23
10469 (K);
King’s
coll.
coll.
K,
80
(E),
Nabee
P);
(DD, K,U);
s.n.
Nakhon
(C).
Sai
Prayoon Bunkhruy
—
(L),
Sörensen
Chom
of Li:
N.N.
s.n.
=
Sawan
Prov.,
Rachaburi
=
RFD
9217
26226
Tak:
Put
100
=
1346
(K, L) ;
Doi
5398 (BKF);
3978 (K).-
Gram &
Syrach-
Tripagodas
(C); Pong
(L);
Hansen
Garrett
5447
RFD
Circle,
&
Smitinand
Thong:
King
605
(P, U).
Larsen
,
(K); Muang Payao:
Yok: Larsen
81
coll.
(CAL, K, P),
544
km N.
Garrett 18
121
(CAL,
s.n.
King’s
Taeng:
Kerr
Bunnak 50
—
Khari:
2583, 3398 (C);
1566 (K);
(BKF, K).
Kerr
Sankra:
E,
HuaiTak:
Mae
Dhani
Prain’s
(CAL, DD),
Kurz
(P).
Nicobar,
(K),
Syrach-Larsen
SING); Wangka:
s.n.
2547—2555,
Mae Li:
7051
91
(BM,
445
RFD9812 (BKF);
Hansen
coll.
Bara
Winit
Sutep:
Lee:
26227
King’s
1182
S. Andaman:
(DD);
34
(CAL, P, SING);
s.n.
Brookesabad:
Dao:
Parkinson
(CAL, K),
Parkinson
Vanpruk 176 (K),
(C); Ampur
87
1
286
(BM, K, L),
(K); Lamphun,
Kerr 5352
Luang:
Larsen
87
Andaman:
(BKF), Sdrensen,
4827
Maharat
(E).
342
6763 (E); Melaung
Circle, Chiang
Mae Hot : Kulchat Nitikul
Bo
Po
(BM, E); Thatay Kyun:
Cubbitt
Gyaing: Helfer
(P, U);
s.n.
Is. Kamorta:
THAILAND.
RFD
Chin
and
coll.
King’s
(P);
s.n.
Bajajag: King’s
1339
8
L).
84 (BM),
Boh:
I.:Proudlock
Mezaligön (Kaukwe):
Distr.,
Nam
Kanchana
Pass:
Ron:
Buri:
A.
82
Boomkrong
=
(K);
Marcan
2443
(K),
SF
653
Kruat, Kao
Hansen
harsen
&
Laos.
Hansen
de Choam
Mong:
Irai:
Béjeaud
18
14398 (P);
Chevalier
F.R.:
Bien
Khet
Pierre
322
Poilane
Mt. Bao
Harmand
Talmy
Schmid
Robinson
3084 (P);
HAINAN.
Nui
S,
3318
Khanh
(P);
Nui
(K, P);
1033
SING);
Alleizette
ChunI 70300
Distr.,
P.
(K);
Ka Chik
Poilane
(E,P);Kg.
3
Ahnot:
Chevalier
Pierre
(P);
7032
(C, P);
403
(P),
307
(P),
Poilane
6754 (K, P);
Thien,
(P);
Mt. Dinh:
(P);
33
Poulo
(=
Ca
Thuan,
30328,
6863 (P);
Thua
Muller
Byh:
(P); Long Khanh, Giaray:
C6n Son
Annam, Ninh
30433,
Tha
Point A: Lefèvre 328
Ti Wan: Pierre
36692 (P);
—
Contest Lacour
r
30329
Hon
Tre
I.:
Condor):
Na:
Poilane
Poilane
(P),
2804
Poilane »931,
Hue: Lecomte & Finet
(P).
41163 (de Pirey s.n.), 41180 (de Pirey 29)
Shan:
Lau
Shan: Lau
—
Tsang
K,
238
LU
=
17007
1273,
Tonkin,
(BM, BO, CAL,
1641 (BM, P); Chung Ngo Chang:
3633 (P, S);
3390,
38 (BM, E,
Lam
Puket
Ko
Yai
P); Wan-ning
Distr.:
How
Lau
70318 (K, P),
Distr.: Fan-ta: McClure
Lung:
Circle, Ranong: Haniff 338 (SING);
Yao
Yai: Kerr
Vanpruk 603
15746 (BM,
Nakon Si Thamarat
—
Annandale
—
6640 (K);
Butang: Ridley
2488 (C, SING).
Sonkhla :
(P).
(P).
Shan: Law
Thailand.
Kloss
(K),
(K, SING); Trang?,
14033
s.n.
3352
How &
CCC
9138
P).
MALAY PENINSULA.
16628
Quoc:
Pierre 7035 P-P-
Trang: Chevalier
Peninsula:
Heo
al.
(P); Kampot: Geoffray
Phu
Tuy, Cape
Mt. Zuin:
Lin-kao Distr., Lin Fa
Fung Ling:
(P); Yeung Ling
(BM, E, K,
Kerr
Chim
Nha
Tri: Chevalier
266
3,
Liang 63433 (K, P);
Kan-en
Han
Phuoc
(Lemesle) (P).
de l'Enfant: Poilane
et
(P); Quang
K, L, P, SING); Ch'ang-kiang Distr.,
(P, S);
Hoa,
Thorel
Preah Vihear:
Magnen &
(K, P); Tiy ninh,
30327
Vo Dat: Chevalier
378
Xoan:
between
4028 (NY).
Thu Due: Pierre 7034
(BM, CAL, K, P);
p.p.
Ban
Reap?, Pagode
Siem
Samrong Tong:
s.n.
(C, CAL, E, K, P);
970
(K, P);
p.p.
(K, P), Lefèvre
s.n.
s.n.
Deo:
1804
Srok
(P), Gourgand
7
Can Tho: Chevalier
7036 (P); Long An,
40831 (P,
Massif de la Mère
(P).Eberhardt
Haiphong,
Phnom:
and L.
(BO, P);
Nun:
Bang
(C).
Talbot de Malahido
13239
(BM);
19773
Trat,
Sarensen,
(P); Vientiane,
Xingh
A
(P); Kg. Thom,
31999
Santuk, Prey
Kuntze
R.:
Thorel
(P),
74
Pierre
P), 1804
762, 803 (P),
(BO, P),
?
(E, K, P);
Hance
Hb.
31984,
Srok
(P);
N'Cor
(P); Phong Dinh,
Chiang:
Chevalier
5639 (BO, P),
1274
K,
(BM,
in Hb.
in
L.
Ban
Larsen &
(C, K),
Schmidt 335
A Loi: Poilane
( Fleury ) (P); Kg. Speu,
31741
Mondolkiri,
Mt. Deon Ba: Pierre
Hoa,
Chevalier
61
and L.
(K);
132
(C, SING);
2777
20472
between
(P);
11919
Ubon
—
Marcan
Sörensen,
W. of Paksane:
(SING);
Phu
Circle, Rayong,
627b, 863
Poilane
Dee
Dang:
Kaen,
(K).
2827
Plain:
Seidenfaden
Schmidt
Din
(K),
889
Buri
Ko Saket:
Harmand 1229
Pierre
(P),
s.n.
Sabab:
Pac Bac:
137
P),
Phnom Penh: Alleizette
(C, P);
7033
1804 p.p., 7035 p.p.
996 (P);
K,
(BO,
Pursat:
(P);
s.n.
(P);
Kanai:
Wang
Khon
(K);
Put
Seng:
Collins
Makham
(K) ;
Sook
Par
7231
Chanta
—
698g (C, S);
s.n.
between L. Sa Roi
Lecomte & Finet
Sap:
Vietnam. Cochinchina:
Pierre
Schmidt
d’Orléans
11743
Couderc
(P),
Khao
Talbot de Malahido
Saravane,
Chum
(K).
318
U);
Kao Tao: Kerr 16162
(K);
(P); Kg. Chhnang, Prey Anghop: Chevalierr36189 (Duquesnoy) (P); Kdey
Kralanh
Penh Lover : Pierre
Poilane
Noi:
RFD
Si Racha:
234
(BO, L,
Put 2307
—
6837 (K, SING),
Kerr
13738 (P); Pakse, Attopeu:
317
Couderc
Tonlé
98,
39
Circle, Chumphon (Sapli): Haniff &
Ban
Circle,
17647 (K); Pliu,
Chang
Poilane
SING);
P,
Collins
6047
Area
Circle, Loei,
1196
Plaing:Lakshnakara
19963 (L),
E. of Vientiane:
Béjeaud
14908 (K, P);
36999 (P);
Ko
(C)'>
Surat
Udon
Circle, Korat,
Loy:
(P); Luang Prabang:
L. Loi and L. Pata: Poilane
Cambodia.
Kerr
RFD
=
Savannakhet:
13688 (K,
Ko
H.B.
—
—
Smitinand
Prachinburi
—
Ban
Buri,
Chumpon:
324
(P);
98
(SING);
Poilane
Chanta
7114
185
Met: Dussaud
124
Dan
Bunnak
Spire
Sitarn:
Indo-Pacific
Kan Kradai:
(K).
12721
the
of
Teysmann
(SING).
s.n.
Kerr
Rachasima
—
(C, E, K);
971
(E, K);
(Q;
193
Chang:
(K).
1324
(K),
(K); Prachuap,
Keith
Ko Tao:
Ochnaceae
the
10378
Saming: Put 607 (K).
Collins
Phe: Put 2723
Kerr
(L); Wang Saphung,
10114
Khao Chalat:
Ko
(K),
(K); Bang Saphan:
Wieng: Lakshnakara
Circle,
of
A revision
Hua Hin: Marcan 2470
4248 (BO, K, SING);
RFD
=
2623s
RFD
Marcan 2160
Nur
Kanis:
K,
=
17310
RFD
SING).
Circle,
Ko
3398 (BKF,
—
Surat
K);
Batong
Ban Don,
Circle,
Samui: Kerr
Phayam (Delisle I.):
Ko
Tunkah: Goldham
(K);
(K);
12543
in
Curtis
3033
(L)adang:
Ta:
Kong
Ta Samet:
Kabok:
Hb.
Is., P.
Kerr
Seidenfaden
Kerr
14297
(K);
15037 (BM,
CAL,
Batten
K, SING).
Kedah, Koh Mai F. R.: Kiah S.F. 35131 (BO, K, L, S, SING); Langkawi Is.:
Pooll
s.n.
(CAL,
K,
SING).
—
Malaya.
—
Perlis, Ginting
Ridley
—
s.n.
(SING);
P.
Vernacular
Langkawi:
names:
Corner
Assam:
(S. Burma), mok-song-hu
tan-luang
S.
(N.
Vietnam:
xo’-lai
Lao),
S.F.
37888 (A, SING),
khimdabeng
Burma:
(N.E.
(cây-bông-)mai-do,
kra-chè
Lao),
mai-nui
Thailand:
(N.
mai-vàng,
or
F.
N.
71204
(K).
indaing-seni (N. Burma), yodaya
(Shan), myauk-min-thwege (?).
chang-nao
K.
t’a-chi-bang (Karen),
Thai), champanam
huŷnh-mai
(S.
(Annam),
Thai).
câm-lai,
(Moi).
Ecology:
From
sea
level
up
to
1200 m
forests, often of a mixed Dipterocarp
are
(Garo).
Wyatt Smith
found
near
river
banks,
type,
small shrubs
in
on
hilly
country,
loamy, sandy,
near
sea
shores,
in
or
moist
or
dry,
deciduous
rocky soils. Tall specimens
and pyromorphic undershrubs
BLUMEA
40
dry hill forest.
stunted,
in
generally
part
little
a
Use: The bark
appreciated
are
cubic
in the
recorded
used for huts
as
Light-brown, hard,
(Watt,
1890)
± 0,87
=
new
in the southern
Malay
(Lanessan,
tonic
decorative by the Vietnamese, especially
wood is
foot
especially
digestive
a
of
during development
or
February and March,
in
restricted,
1968
I,
before
mainly
yields
No.
XVI,
shortly
area
but less
bitter and
of wood:
Properties
per
earlier,
tastes
as
(tet). The
Year
Flowering
in the northern part of the
leaves:
VOL.
Peninsula.
1886).
The flowers
during the Buddhist
New
in the Andamans.
and
close-grained
brittle; weight
lbs.
54
3
kg/dm
.
Remarks: All Ochna specimens found between the hills of Assam, the Malay Peninsula,
and Flainan
in
considered
are
by
me
this part of the world,
in
use
of the variability of this species.
remarkable,
sometimes
are
infraspecific
dominant
do
come
taxa
the herbarium
among
that
Merrill
description of Elaeocarpus
(Lour.)
Merr.
I have
stems,
as
seen
the
correct
from tuberous
sprouting
O. obtusata DC.,
that
integerrima (Lour.)
O.
Griffith (1854)
described three
new
1 have
Ochna
O.
Griff,
crocea
integerrima (Lour.)
described from
was
distinctly filamented
subsessile
10
are
also
(1871)
his
more
spp.
it
were
by
not
and leaves.
a
is considered
obtusata DC.
Although
given
considered
was
as
name
is
a
synonymous
referred Griffith's
parviflora
plant
to
Griff, is
smaller flowers and leaves,
already
from
.'
indicating
are
.
to
15
stated above,
Bennett
ever
species
He did
a
as
probably
as
such
was
a
Kurz
for
any arguments
(1893), and
mistake,
from
depicted
made
no
other authors.
other records
a
flowering
of
specimen
O. lucida Lamk. and
to
by later authors and consequently
=
considered by
give
only
since.
No reference is
‘squarrosa’
in
that all three
and the embryo
torus
was
not
later homonym and illegitimate. Both
O.
seen
position in Ochna with
a
of the
(1875), King
described and
newly
have
holotype specimens
ovaries rather than Gomphia with
sumatrana Jack.
Burma
I
by
Merr.
that Kurz made
however,
me,
of the
some
fruiting specimen. The characters: ‘filamentis
Besides, the characters
accepted by
was
of
The
by
name
Griffith's
ovaries.
up
seen
with O.
from Tenasserim.
to trace
and
Griffith in Amherst.
Griffith's
Ochna
5
probable,
Ochna lucida GrifF.
collected
as
spp.
This
Kurz.
Merr. and
in accordance with those of Ochna. This
more
Gomphia
quently
specimens
different from
Originally
nature.
stamens
conspecific with Gomphia
as
the
and
stamens
decision, but
It is
fruticulosa
(Lour.)
Ochna
not very
flowers, inflorescences,
been able
not
are
reflexed in fruiting specimens
fundamental
very
pluribus persistent’ and ' ‘ovaria plura abortiva
many
variety, these
of
not
named O.
Burma and N. Thailand with low
the Indian pyromorphic
elsewhere. From the descriptions it is highly probable, however,
or
conspecific with
are
a
is
treated here.
from
separate
with O. integerrima
synonymous
several of his collections,
CAL
of
partly caused by ecological
in the flowers.
interpreted and emended the
correctly
Unlike
characters
not
specimens concerned have been
as
a
calyx, which
but this character is probably
me
roots.
Merr. in
is in the
only exception
has
are
Thailand, but they
Lour, and I accept the combination O. integerrima
specimens
united in
are
at least
parts
and useful
Small-leaved forms
parts of
striking similarity
for the species
name
be
vegetative
distinguish clear
to
adjacent
might
also because
but
of the
studied.
specimens
(1935)
integerrimus
pyromorphic
some
a
traditions,
and size
shape
impossible
and this
too
On the whole there is
doubt
no
found it
I
from Burma and Assam
have
Differences in
have been commonly
names
of local
only because
materials from Indo-China and
among
circumstances.
I
but
Several
conspecific.
as
not
names
Bennett
were
conse-
incorrectly
(1875)
wrongly
O. obtusata DC.
form of O. integerrima (Lour.) Merr. with slightly
already
local forms
noticed
do
not
by
Bennett
deserve
any
(1875).
In
my
taxonomical
opinion,
status.
A.
Kanis
A
:
revision
Gen. PL
Gray, New
U.S.
Bot.
Expl. Exp.
1
(1853)
359;
(1895)
6
Gilg
in Morot,
in E.
&
Nov.
Campylopora
Pfl. Fam. ed.
Indo-Pacific
Am.
(1857)
4
(1880)
App.
(1899)
1
19
Area
41
181.
in E.
Nat.
A. C.
74;
77;
Baill.,
51;
(1862)
1
Hist. Pi.
15
4
(1901)
160;
(1902)
393;
16
Am. Arb. 36
Smith, J.
B. nitida A.
(1902)
8
(1857)
3
P., Nat. Pfl. Fam. 3,
&
Bot. VIII,
Type species:
—
Sc.
&
H., Gen. Pi.
&
(1871)
1
Ann. Sc.
(1925)
21
2,
(1962)
Arts
B.
Bartell., Malpighia
221;
202;
Ac.
421;
Gilg
39;
Bull. Mus. Hist. Nat. Paris
Tiegh.,
v.
Proc.
Ann.
(1902) 46,
16
284; Furt., Gard. Bull. Sing.
the
of
Oliv. in Hook., Ic. Pi. 3,
34;
Filip.
Bot.
J.
P., Nat.
offprint of
5,
Queensl. Fl.
Bailey,
142;
Tiegh.
v.
Fl.
F.-Vill.,
Ochnaceae
(1854) 361; Walp.,
318; Seem., Fl. Vit. (1865)
(1873)
the
BRACKENRIDGEA
2.
A.
of
Ann. Sc.
547;
(1955)
Gray.
Nat. Bot.
16 (1902) 404.
Brackenridgea
VIII,
Type species: C. australiana (F. v. M.) v. Tiegh.
=
—-
nitida A.
Gray.
Notochnella
Bull. Mus. Hist.
Tiegh.,
v.
Nat. Paris
(1902)
8
Ann. Sc.
549;
Nat. Bot.
16 (1902)
Brackenridgea
VIII,
Type species: N. fascicularis (Blanco) v. Tiegh.
403.
=
—
fascicularis
Trees
treelets
often
appearance,
rachis
often
at
of
up
often
many
chartaceous,
ones
veinlets branching,
made
branches.
spreading
Leaves
of the lower
some
successively,
the
with
caducous.
laciniate,
apex,
F.-Vill.
(Blanco)
or
simple
growing
partly parallel
±
or
often
small, free,
nerves
higher
small,
flowering; bracts
which is
Petals
5( —10),
the
from
distinctly tumid and red
endocarp;
when
yellow.
downwards
apex
Distribution:
Obs. Nat.
annularly
Fruits
I
Pacif.
to
the
of
two
description
genus
combination: ‘? B.
I
anthers
5(—io),
black
turning
ovule
intrusions
connecting
2
dehiscing
obovoid;
almost
or
of
so
presume
the
by
name
Exp.
on
1930,
and
1000 m
red
265).
a
3,
new
Dispersal
The fruits
calyx
(Guppy,
also capable
are
and endocarp;
this
and from the
was
New
1885, 289, pi. 54).
the
name
solely
Asa
from
species
is based
Fiji.
altitude.
and
torus
exocarp
published by
Since
generic description
KEY
Petals
Petals
fruits
c.
Wand. Borneo, 1904, 187)
was
of
to
up
between
spaces
hookeri (Planch.)'.
the
areas,
Ridley, Disp. PL,
air-filled
Sumatra, the Malay Peninsula,
Queensland,
NE.
on
in 1853. The
Gray
Fiji:
B.
B. hookeri
nitida,
and
follows
materialsfrom
Fiji
a
generic
by
a
new
question-
and therefore
descriptio generico-speciflca.
a
5—7(—10);
5;
stamens
Inflorescences
3.
around
Chang,
black
conspicuous,
Brackenridgea
is followed
Ko
tropical
Bot. Chall.
(Hemsley,
Remarks: The
2.
Ovaries
little
a
hemispherical
and red in fruit.
enlarged, fleshy
5,
reddish,
New Guinea,
everwet
1906, 569;
2,
floating because
Guinea seadrift
1.
curved
—2(—5),
recorded from the Kapuas R. (Beccari,
1.
slits.
longitudinal
Celebes,
birds because of
mainly by
it is
Sepals
(or cv>); filaments subterete;
io
the Andamans,
in
species
5
Confined
Ecology:
mark,
cymes,
caducous,
ripe.
Borneo, the Philippines,
of
±
small.
stigma
in fruit.
Stamens
by
epitropous,
camptotropous,
the
joining
base of inflorescence with small, axillary buds; pedicels filiform,
white or
the
to
shortened
growing and turning red in fruit, ultimately caducous. Flowers with ±
torus,
ones
less
or
of umbelloid
thyreoid but
distichously arranged,
after
more
curved
strongly
the margin, the
to
Inflorescences
transverse.
compound,
vegetatively
on
Stipules
glossy above,
cm
5.
axillary,
made
long
10—45;
ovaries
never
Inflorescences
7^—20
stamens
10;
up
of
ovaries
(Sect.
but
TO
B.
THE
5—10.
SPECIES
(Sect.
A.
Notochnella).
.
.
1.
B.
fascicularis
Brackenridgea).
sometimes
many-flowered
terminal
cymes,
on
the
short
pedicels
side-branches.
in
2
or
more
tiers;
leaves
2.
B.
mostly
hookeri
42
BLUMEA
VOL.
XVI,
No.
i,
1968
cited
from
gen ralised
slightly
only
are
areas
Outlined
areas.
neighbouring
and
Malesia
in
Brackemidgea
of
species
the
of
map
Distrbuional localites.
6.
Fig.
A.
made
Inflorescences
3.
Kanis
A
:
of
revision
the
Ochnaceae
of 3(—j)-flowered cymes, the
up
the
of
Area
Indo-Pacific
pedicels
in
43
leaves
tier;
I
mostly
long
2.
inflorescences
Axillary
4.
Anthers
4.
Anthers
always
ij by J
c.
hy J—f
2—3
when in fertile
present
about
mm,
than
more
mm,
long
as
Cymes
±
length,
the
in
Flowers
in
Stamens
<x>,
minute,
papillose.
species
1
For
1
1
the central
in the
5—10;
Corolla
yellow.
irregular,
style correspondingly ribbed; stigma
excluding Palawan
Philippines,
the establishment
on
unequal
sometimes
shortened,
±
flower; pedicels articulate, the short basal
flowering successively.
cyme
whorl. Ovaries
discussion
a
distinct; branches
±
nitida
B.
5.
Tiegh.
v.
—
of
than
more
Distribution:
Remark:
Notochnella
nov.
peduncle
longer overtopping
persistent.
part
stat.
remote;
filaments
as
forbesii
B.
4.
long
as
A. Notochnella
Section
(v. Tiegh.) Kanis,
3 times
cm
palustris
state.
filaments
as
4—12J
B.
3.
of this
Prov.
section
new
under
see
B.
fascicularis.
fascicularis
Brackenridgea
I.
Fam.
Gen. Pi.
&
Malpighia
Gen. Pi.
Tiegh., Bull.
403.
105;
v.
—
1023
fr.
Tree
4—-8
of
mm
mostly
attenuate
cymes
Torus
acute to
base,
at
c.
1
mm
Philip. J.
0.5—0.7
Ecology:
In
2
entire
1—2
to
3 —4
mm
mm.
mm
style
c.
4
a
203.
—
Blanc.
Sp.
Rev.
Kuntze,
Notochnella fascicularis
Nat. Bot. VIII,
(1902)
16
fl.
287; En. Philip. Fl.
(1920)
17
on
Dipterocarp
account
(1845)
±
at
long,
up
0, in fruit
long,
in the
at
to
3
up
shortened
±
in
cm
to
to
Pi. 3
(1923)
once
up
4
mm
cymes,
high,
7
2f—3 by
to 7
along
described this species
mm
a
f
c.
in the
to
a
little
terminal, made
sometimes separate
to
1—5
0.
6—8
mm.
Fruits
5
by
mm
2
mm,
persistent.
Sepals elliptic
by 2f—4f
Ovaries
up
to
7
0.7
by
6
to
mm.
—0.8
mm.
Palawan Prov.
(Merrill, 1923).
Once
reported
stream.
genus
consequently
placed this species closer
sometimes
cm,
sometimes
up
mm
long.
Ochna L., since it has
Fernandez-Villar (1880)
of its annular embryo;
(1902)
2—5
acute,
fruit, the basal
Philippines excluding
s.s.
to
laciniate. Leaves
mm,
by
obovate-lanceolate,
anthers
in fruit
and
by if
mostly triangular,
low and medium altitudes
forest
5
5—15
finely denticulate. Inflorescences
Petals obovate
mm
to
obtuse
apex,
bracts
leaves;
up
lanceolate,
to
transferred it
he emended the
Most later authors have followed Blanco rather than
Tieghem
separate
Stipules
cm.
long filaments;
subspecies
Remarks: Blanco
genus.
30
or
latter floral parts thanBrackenridgea
Van
(1902)
549; Ann. Sc.
many-flowered,
cm
primary forests
from logged
up
of normal
if—2f
mm;
Distribution:
this
Sc.
little acuminate
number of
high, if
by
6—8
a
margin
the axils
in
Stamens with
genus
16
O.
—
Bartell.,
79;
29;
(A, L), Luzon, Bulacan Prov., Angat,
p.p.
high; dbh.
m
varying
a
obovate,
by
(1902)
8
(1886)
(1905)
27
Diporidium fasciculare
—
Bot.
J.
long petiole; lamina oblong
caducous; pedicels
±
in Morot,
Filip.
Vase.
Publ. Philip.
40.
Vidal, Syn.
92;
25-IV-1919.
up to 25
obtuse,
up
68.
Pi.
App. (1880)
(1878)
3, 2
Type: Ramos & Pascasio B.S. 34479 (A, K, P, PNH holo f?), Mindanao, Surigao
Prov.,
with
(1923)
3
Tiegh.
Ouratea mindanaensis Merr.,
68.
Govt. Lab.
Nov.
Filip.
ibid. ed.
245;
Rev.
27A;
t.
19,
Fl.
F.-Vill.,
(1845)
2
Mus. Hist. Nat. Paris
Neotype: Vidal
—
(1883)
162; Merr.,
Philip. Fl. Pi.
En.
(1891)
1
Filip.
(1901)
15
(1918) 263;
v.
(Blanco)
fascicularis Blanco, Fl. Filip. ed.
Ochna
Brackenridgea
to
more
the latter
description of
Fernandez-Villar, but
giving it the
status
of
genus.
The indefinite number of the floral parts
certainly indicates
a
relationship
with Ochna.
VOL.
13LUMEA
44
the other hand, the annular embryo,
On
blade,
of the venation,
especially
intermediate stages between the
No.
XVI,
1968
I,
and the characters of the leaf
the free stipules,
refer
The
Brackenridgea.
to
branched
distinctly
inflorescences represent
that
thyrses
found in
are
and the essentially similar, but strongly reduced inflorescences of Brackenridgea
somewhat intermediate position of the species is best reflected by placing
section in
it in
Ochna
s.s.
The
separate
a
Brackenridgea.
In have indicated
Vidal
1023
original material from Angat
from the
originates
same
area
neotype, since
a
as
and
it is
not
there is
as
is
doubt about the
no
that Blanco's
likely
very
existing. Vidal's collection
still
is
suitable
very
it
as
of this
interpretation
species.
KEY
I.
Stamens
I.
Stamens
a.
fascicularis.
ssp.
Inflorescences
up
Distribution:
LUZON.
Bulacan
SING),
Agr..
Luzon:
P,
Ramos B.S.
14923
(L, P).
Norte
Prov.:
Andrade
Albay
Prov.,
Cagraray
(CAL,
—
Romblon
Vernacular
:
Merrill
Merrill
Stamens
Zambales
—
1023
(A,
K,
Blanc.
Sp.
Bur.
B.S.
Quezon (Tayabas)
L).
fascicularis
ssp.
ssp.
mindanaensis
long
mm
Ovaries
—45.
Subic:
Prov.,
—
Rizal
Oro F.B.
Vidal
28669 (A, P).
—
Ahern’s
BO, K,
48803 (A, BM, E, K).
Prov.:
Bur.
Merrill
Prov.:
916 (A, BM,
837,
Paracale:
28144 (SING);
7
peduncles,
—10.
L,
Agr.
coll.
2099
Laguna Prov.,
30684,30979,31067(SING).
692
(K, L);
Catanduanes
P,
(BO);
Pilea:
San Antonio:
Camarines
—
Makahadok: Riviera
Prov.:
-
K,
Merrill Bur.
P); Tanay:
Ramos B.S. 2119
—
(K).
(BO,
3077
8
(A).-
B.S.
Ramos & Edaño
73269
Petals
(3 —)j.
Distribution:
MINDANAO.
B.S.
to
(Merr.) Kanis,
2—6
cymes
(3—)5.
according
bitas, mala-kiting, masalisi (Tag.); Visayas:
Merrill
stat.
with j—io(—20)
Stamens
(1923).
nov.
—
Ouratea mindanaensis
often indistinct by
long rachis,
cm
mm
(8 —)io—15.
long peduncles,
Ovaries
Distr.:
34479
Mallonga
F.B.
San
Mateo
27003
was
Bo.:
Mendoza
referred
genus
to
case
due
in
bracts
to
P.N.H.
iron
41896 (L, SING).
deposits
at
NE.
—
coast:
Sungao
Ramos &
duplicates
as
rather similar
to
Ochna fascicularis
Ouratea Aubl. because of the small numbers of flower parts,
and similar
The type collection is
androecium. In
probably
of
7-flowered. Sepals
(A, K, P).
but in other characters of flower, inflorescence,
from that
up to
—7.
(A, BM, BO, K, P);
Remarks: Ouratea mindanaensis Merr. was described
Blanco. It
6
Merr.
development
Mindanao.
Agusan Prov., Butuan,
Prov., Surigao
18831 (K)
F.B.
Luzon: aniatan, dirigkalin,
names
Inflorescences with
normal leaves;
I.: Franco
Prov., Sibuyan
mindanaensis
b. ssp.
the
2—J
20
Agr. 2691 (BM, K), 2693 (K),
Edaño
I.: Bautista F.B.
names:
bansilai (Bis.). All
Pascasio
with
cymes
5 ——10).
153d (A).
issbis (K),
SING),
F.B.
a.
b.
long peduncles
mm
SING).
VISAYAS.
to
Petals
San Andales:
3296 (BO);
long peduncles
Visayas.
Boso 2
(A, K, SING);
B.S.
SUBSPECIES
long rachis;
cm
5.
Vidal
Vidal
Angat:
(BO, K,
3174
2297
Ramos
—2\
I
Luzon, N.
Central
Prov.,
THE
Ochna fascicularis Blanco.
—
with
15-flowered. Sepals
to
TO
mm
2 —5
with 5 —io(—20)
Cymes
10—15.
with
Cymes
20 —45.
to
Brackenridgea,
and leaf venation it is
especially
B.
fascicularis
remarkable for the strong reductions
from
A and
connation between
1
K, I found
or
2
3
or
4
sepals
pairs of the original
in
calyx,
per
5.
clearly different
(Blanco) F.-Vill.
corolla, and
flower, which
The
same
the corolla, judging from Merrill's description, but all petals
is
might be
were
shed
on
A.
the sheets
A
The
Gray,
s.s.
distinct
caducous, leaving
regularly
peduncle
10,
45
occasional deviation of little
an
for
young,
or
no
of its leaves
many
palustris).
Tiegh.
v.
and branches much
white. Stamens
5-merous,
Area
Indo-Pacific
Brackenridgea
B.
Flowers of
scars.
the
of
apparently
Campylopora
—
Cymes much conferted;
only
is
under B.
also
Section
A.
Othnaceae
specimen
denticulate (see
Brackenridgea
the
of
revision
I expect that this is
me.
importance.
distinctly
are
by
seen
taxonomical
Kanis:
in
I
shortened;
whorl. Ovaries
1
pedicels ultimately
Corolla
flowering simultaneously.
cyme
style 5-ribbed; stigma
5;
small, 5-lobcd.
Distribution:
species in the Andamans,
4
Borneo, Palawan,
Celebes,
hookeri
Brackenridgea
2.
Proc. Am. Ac.
(1857)
&
Rev. Gen. Pi.
of W.
Isl.
excl.
233,
Fl. Br. Ind.
,
106.
Tiegh.,
(BO, CAL, E, K,
7310
1
(1849)
(1875) 526,
I
Ouratea hookeri
—
Type: Phillips
—
42.
offprint of
6,
in
4
(1859) 675,
1, 2
King, J. As.
525;
Mai. Pen.
Ann.
Flook.,
(1922) 366;
1
Gomphia umbellata Hook.
—
met.
nom.
Bat.
(1875)
1
Fl.
Ridl.,
Ochna hookeri O. Kuntze,
—
Kew Bull.
Burk.,
(1853)
Fl. Ind.
Miq.,
(1961)
3
Pi.
(1854) 361; Walp.,
1
Br. Ind.
/, Fl.
corymbosa;
var.
Gen.
Bot.
Gomphia hookeri Planch,
—
182;
Fiji.
(1935) 318;
Diet. Econ.
Herb. Hook. (K holo), Malaya,
s.n., ex
Penang."
P.
v.
Ann.
182.
Benn. in Hook.
(1935) 1615.
2
=
perakensis
B.
f.
(1891)
1
Prod. Mai. Pen.
P.
Walp.,
(1962)
19
New
Expl. Exp.
the Malay Peninsula,
Sumatra,
Queensland,
Gray,
U.S.
51;
List Trees & Shrubs ed.
Chit,
U
in Hook.
Benn.
ex
3;
(1893)
II
Beng. 62,
Hundley
f.
(1847)
6
A.
(Planch.)
(1857)
3
‘G. glaberrima Planch.';
errore
Soc.
Bot.
J.
Sc.
Arts&
Chang,
Ko
Guinea, NE.
Furt., Gard. Bull. Sing.
421;
Lond.
New
Ann. Sc. Nat.
(1902) 396.
16
Bot. VIII,
holo, SING), Malaya, Perak, Larut,
P
—
Type: King’s
1500 —2000
ft.,
fl.
coll.
fr.
&
II-1885.
Gomphia corymbosa
Fl. Mai. Pen.
Tree,
with
to
to
up
acicular
±
rachis
entire
often
or
sessile
growing
not
mm
3
high,
5
mm
c.
±
by
c.
\
mm.
up
Ovaries
c. 0.7
by 5L
to
ANDAMANS.
Middle
KO
Penang: Phillips
2134
(SING),
=
s.n.
2134
mm;
often
if
a
up
Torus
20
c.
\
mm
mm
or
—
laciniate. Leaves
at
i
i| —2
obtuse
of
up
the
small, triangular,
those of
mm.
in fruit
up
one cyme
in fruit
0,
mm
Petals
long filaments;
long,
cm,
base, margin
simultaneously,
fruit,
high,
by
i—mm
style i\ —3
in
mm
4—5
typus;
183.
terminal, made
bracts
flowering;
to
obovate,
to
mm,
little tapering
Inflorescences
after
Stamens with
0.5
Ko
Andaman:
Chang:
MALAY PENINSULA.
P.
by
ovate
by
flowering successively
long,
mm
(1962)
lanceolate, ~]\—20 by 2\—6
treelets.
young
cymes,
8
excl.
p.p.
19
Timah, fl. II-1900.
Bt.
to
up
33,
Sing.
ovate
anthers
to
5
mm
up
to
—2
long.
mm.
Distribution: Andamans,
THAILAND.
mm.
to
acute,
distinct tiers.
Sepals
0.
obovate, 3|—5 by i-J- —if
Fruits
apex,
10 —15
3
Stipules
m.
oblong
vegetatively
on
pectinate; pedicels
fixed close together in
to
at
denticulate in
many-flowered, shortened,
1
to
up
(1910)
Soc. 54
Gard. Bull.
holo), Singapore,
lamina
long petiole;
sometimes acuminate
acute,
entire,
SING
dbh.
high;
m
33
mm
5 —12
B. denticulata Furt.,
—
10738 (BM, K,
Type: Ridley
Str. Br. R. As.
(King) Ridl., J.
(1922) 367.
1
Schmidt
Thailand,
Chang (NE. Gulf of Siam),
Parkinson
Surat
P,
(DD,
K),
1203
Malay Peninsula,
Borneo.
(CAL, DD).
614 (C, K), 665 (C).
Circle,
(K); Haniff306 (BO),
(BM, E,
53
the
SING).
—
Ko
2409
Phangan:
Put
(BM, SING);
Perak,
Larut:
King's
769
(K).
—
Malaya: Hervey
Government Hill: Curtis
coll.
7310
(BO,
1147
s.n.
(K).
-
(K, SING),
CAL, E, K, P,
SING).
—
BLUMEA
46
G.
Negri Sembilan,
S.
(SING);
935
Corner
S.F.
Ridley
28993 (A,
Sinclair
Res.:
Seletar
(BM),
s.n.
BORNEO.
Semengoh
K, L,
(A,
SING).
Vernacular
From
Remarks:
his
Kanis
in
Malacca:
G. Pulai:
P
Gading
G.
5649 (BO,
Kanis
King
the
Peninsula:
(Sarawak).
level
to
up
730
The
real
B.
H.B.
(1962)
came
except
King's coll.
are
183.
Banka
s.n.,
A.
s.n.
Tiegh.,
v.
ft.,
B. serrulata
forests
on
luru
kayu
sandy soils,
and hill forests.
forests,
Gomphia
on
material and Ridley based
former
as
the type colfor all sheets
the type of both,
as
to
different
Gomphia corymbosa
Ridl. is
me
hookeri,
B.
as
belonging
as
a
Beccari
Type:
but he again
It
hookeri,
as
was
new
little from
based
on
five
Timah. In
could observe
I
Timah) that leaves
Bt.
very
and four from Bt.
locality
his
typified
species differs
of its leaves.
margin
B.
Singapore:
Gray
v.
(BM, K, L,
P
B. rubescens
P
fl.
&
var.
of juvenile
my
the field
in
specimens
rule.
(1901)
15
(A,
3472
165,
t.
10;
holo, K, P),
FI
Furt.,
Gard. Bull.
Borneo, Daiiao
W.
fr.
corymbosa King,
var.
Ann.
Soc.
BO,
Sc. Nat. Bot.
54
(1910)
CAL
Tijd.
N.I. 32
K,
(FI
Ann. Sc.
Tiegh.,
Soc. Beng. 62,
(1902)
16
p.p.-,
33,
holo,
J. As.
VIII,
15
(1901)
holo, K),
395.
Fl.
Mai. Pen.
P,
SING),
L,
163,
W.
Nat. Bot. VIII,
holo), Malaya, Perak, Larut,
v.
Nat.
(1873)
—
411.
—
Paratype:
—
I
II
(1893)
233.
—
Gomphia corymbosa
(1922) 367.
Malaya,
Perak,
—
Type:
Gopeng,
VII-1883.
Beccari 3469
Tiegh.,
leucocarpa Scheff.,
(BO holo, L), Banka, Djebus, 'mensulung kaju', fr.
Bartell., Malpighia
Type:
(CAL, K,
(BM,
4673
500—1000
kingii
maskam,
(not seen).
Str. Br. R. As.
King's coll.
B.
Samarinda
while the Calcutta sheet gives
regarded by
to
of unknown
one
denticulate
—
(Planch.)
corymbosa
—
any
He stated that this
conspecific with
Gomphia hookeri Planch,
184.
bunga
be regarded
to
conclusion,
same
denticulate
finely
For. Res.;
are
Lectotype: Teysmann
Ridl., J.
CL);
13387
fr. V-1867.
B. hookeri
Buddingh
Borneo,
described by King,
one
4673 has
Brackenridgea palustris Bartell., Malpighia
(1962)
E.
evidently based
was
mention
handwriting,
same
Ridley 10738.
denticulata is
Lamadgian,
B.
in the
about the
to
in the
(Sarawak: Semengoh
19
—
16a
16, B
Res.: Muas
For.
of this species.
of the latter species
3.
(BO).
p.p.
merah,
(1910)
not
10738 is referred
only Ridley
collections from Singapore,
Sing.
SING); Kuching
K, L,
species.
B. denticulata with
opinion, B.
8465
kelat
(King) Ridley
Evidently
the
synonym
hookeri,
Sabal
lowland Dipterocarp
collections, mentioned by Ridley,
Furtado
species
corymbosa
var.
and
Since
species.
no
13589 (A,
(K, SING);
910
(E, SING).
39573
Kuala Serait: Kanis B
Ridley 10738 and King's coll. 4673, indicating the
on
the author.
2
S.F.
reported from kerangas
m,
forests, primary
swampy
name
as
436 (SING),
=
(L); Sadong Distr.,
bunga
empodat
Gomphia corymbosa
variety
4334
SING);
lection. However, the latter collection is the only
bear the
Hullett
For. Res.: Paie
Bio
Hudang: Derry
NW. of Mawai:
(SING);
s.n.
(SING); Bajau -.Ridley 3987 (SING);
s.n.
Sinclair
K, L,
Madjang: Teysmann
P.
coll.
S.
Maingay 1369 (K);
Henderson
Bt. Timah:
(L);
21
1968
I,
L).
Malay
sea
Distr.,
B
corymbosa King (1893). King did
var.
species
(K,
—
Singapore: Cantley's
(SING);
10
W. Borneo,
—
s.n.
names:
hillocks
—
Gondol: Shah
Bt.
(SING).
39136 (E, L, SING);
Lundu
Div.,
1st
Borneo:
Ecology:
hookeri
SING).
S.F.
Park,
Kostermans
dry
BO, K,
6760
=
3167
No.
XVI,
(SING). —Johore,
s.n.
Res.: Mead Sar
For.
(Malacca);
Burkill
Alvins
10738 (BM, K, SING);
Sarawak,
Distr., Bako Nat.
reg.:
Tampin:
Ujong:
VOL.
Ann.
holo, SING),
t.
Changi,
Ochna foxworthyi Elm., Leafl. Philip. Bot.
5
Furt., Gard. Bull.
(1902)
16
100
Sing.
395.
—
19
(1962)
fr. V-1867.
Type: King's coll. 6396
ft., fr. VI-1884.
Sc. Nat. Bot. VIII,
Singapore,
9;
Borneo, Danao Lamadgian,
16
fl.
(1902) 396.
—
Type: Ridley 2072
1890.
(1913)
1823; Merr., En. Philip.
Fl. Pi.
3
Kanis
A.
(1923)
68.
Tree,
to
up
with
4
by
—
3 —io
i|—5
acicular
±
cm,
5 —io(—25)
Pulgar,
Puerto Princesa, Mt.
bracts
long,
mm
Torus
c.
\
by
0.7
by
6
I
style
mm;
in
fruit, those of
Petals
mm.
mm
(1 —)2-|
by
3|
to
up
0, in fruit up
mm
made
anthers
long,
little
a
\\—3 by
pedicels
in
Sepals
i
c.
one
ovate
mm.
Ovaries
mm.
in fruit.
growing
the
in
together
0.
mm
the
succession,
branching
fixed close
6
3(—5)-
flowering
laciniate;
±
mm,
high,
mm
4
of
up
peduncle,
obovate-lanceolate, 3! —7 by i|- —3
ovate to
mm
2\
one cyme
to
long filaments;
—4
base, margin entire,
sometimes
vegetatively,
acuminate,
acute to
at
long
cm
(ovate-)lanceolate,
to
terminal,
\
to
up
entire or laciniate.
mm,
of different stages in close
on
linguiform,
(|—)i—2(—2-3r)
0.5
with
or
growing
mm
20
high,
oblong, 31 —6 by i|—3
Stamens with
by
treelets, mostly
young
inflorescences
2
rachis
to
up
8
to
up
(ovate-)oblong
treelets. Inflorescences
sessile,
broadly
mm
in
long
cm
young
cymes,
long
Stipules
lamina
rounded to acute, often a little tapering
sometimes
mm
parts;
8 —15
8
47
184. —Type: Elmer 13074
(1962)
19
Area
Indo-Pacific
f?, U), Palawan,
1.20 m.
petiole;
to 20
up
apex,
denticulate in
simultaneously,
tier.
at
to
up
long
mm
shortened
flowered,
lower
high, dbh.
m
30
sometimes obtuse
c.
the
of
IV-1911.
Leaves
to
Ochnaceae
PNH holo
(A, BM, BO, CAL, E, K, L, P,
fl.
the
of
revision
A
foxworthyi Furt., Gard. Bull. Sing.
B.
—
:
Fruits
up
to
mm.
Distribution:
Remarks :
in Sumatra, the
subspecies
3
Up till now generally
not
Malay Peninsula, Borneo, Palawan, Celebes.
recognized
different from B. hookeri in Sumatra
as
and Borneo.
serrulata Bartell.
B.
together with
published
was
palustris Bartell. Studies
B.
herbarium and also observations in
type material and other collections in the
Sarawak have convinced
in
form of
juvenile
a
denticulate leaves,
palustris,
that material under the former
me
the latter.
Young
often found
are
KEY
i|—2
I.
Anthers
1.
Anthers 2—3
a.
2.
Filaments
2.
Filaments
c.
c.
—
B.
Style 2J —4
2—2J
I
mm
mm
THE
anthesis
long during
long during
long during
anthesis.
Ovaries
c.
0.7
by
0.5
long during
anthesis.
Ovaries
c.
0.5
by
0.3
var.
SchefF.
leucocarpa
B. serrulata Bartell.
—
long,
name
B.
a.
palustris
ssp.
anthesis.
mm
B. hookeri
the
SUBSPECIES
mm
—
have chosen
I
represents
and relatively
species.
J
palustris Bartell.
only
name
larger
—
—
B.
b.
mm
mm
ssp.
c.
Gomphia
hookeri
kingii
Tiegh.
v.
foxworthyi
ssp.
kjellbergii
corymbosa
var.
B. rubescens
—
Tiegh.
v.
Cymes
1 —2
3-flowered, sessile,
treelets. Sepals
young
long
mm
2-J —4
SUMATRA.
N.
Langga Pajung:
Toroes
W.
3752
Sumatra,
(BO);
222:
near
81
to
up
J,—
Sumatra,
Bila
(A,
K,
bb
Beguin 314 (BO,
de
L,
i|—3
3799
(A, L);
287
741
U);
(BO);
(BO);
P.
S.
Kuantan
Voogd 348
=
Peninsula,
Si Mandi
Bengkalis:
of Selat
by
0.7
3491
in
Stamens with
0.5
mm;
Rahmat
3498 (L).
Bruinier
64
—
=
(L);
Kota
si Toroes
4162 (A), 4187 (A,
B. Sosa
Riau,
NIFS hh
NIFS bb 26482
.
13603 (BO).
—
S.
Pinang Distr.,
(A), 3616 (A, L); Sibungan:
724
near
(A, BO, L).
Mentawai,
P.
Dalu 2
(BO, L);
Pandjang: Beguin 326 (BO, L);
Distr., Djake:
mm.
style
mm.
14290
(A, L),
Angin:
Vermeulen
c.
long peduncle
cm
Borneo.
Lor zing
3432
6
\
by i|—3
Ovaries
by
8
to
up
3 Is —6
long.
to
Labuanbilik:
NIFS bb
Petals
mm
up
(A, K, L),
Pariaman: Meijer&
Begum
NIFS
Palembang:
L),
near
near
3293
mm.
Fruits
the Malay
Rahmat si Toroes
Tiku
=
S.
5-flowered and with
by t|—3
anthers
filaments;
Sumatra,
Tamansari:
Bruinier
3
long during anthesis.
mm
Distribution:
si
long.
long. Style
in other
occur
TO
already.
2
palustris.
ssp.
King.
mm
mm
bearing
be fertile
to
denticulate leaves also
since
still
trees
of the
the field
P.
—
S.
P.
:
v.
Kanan,
Rahmat
K,
L).
—
d. Voort 27
Tebing Tinggi:
Rangsang, Panglang
Sumatra, S. Sekanak
Siberut: Iboet 214
(BO, K, L,
BLUMEA
48
SING,
U);
Kloss
MALAY PENINSULA.
Kloss
coll.
NIFS bb
C
663 (K,
Malacca:
—
L,
SING).
K,
R. :
Tampinis
K,
(CAL,
BORNEO.
Haoiland
L, P, SING).
246 (A,
Sarawak,
1st
(BM,
K,
BM, K,
K,
SING),
(K, L);
jojó (K,
Telok Asam:
Sandakan
Be ccari P.B..3469
S.
bb
2
reg.
B
Vernacular
names:
7780
S.
k.
k.
lude,
(Banka); mensolongang
with
humic
0.5
mm,
P.
with
if—2\
style
1^—1\
12423
Ecology:
c.
latis;
J
sepalis
4
Borneo,
0.3
16
SING);
2072
H.B..
For.
Djanan
8465
reg.:
(K, L, SING),
(K,L);2ndDiv.,
Res.:
West
p.p.,
Plateau:
Pasir Puteh:
Bt.
—
Haviland
Matang:
Merurong
Ashton
Sati
Nicholson
Borneo,
Danau
(BO, K, L, P);
Sambas
Lamadjang:
8466, 8495 (BO, L);
Kostermans
9943 (BO,
K, L);
madu
timur bĕsih
to
up
(P. Madjang).
1000
reported
m,
sometimes rather
sandy,
on
luai, mĕnsulung
(Pahang) ; pĕndorah,
mura
occasionally
forests
barat
dahan, sĕniang (Malay); kaju
Peninsula: lidah
being used
as
stat.
Sepals
nou.
—
5 —by
long
during
SING);
2888
rocky
in
housebuilding (Malacca, Sarawak).
Ochna foxworthyi
2
anthers
anthesis.
2—3
Fruits
Elm.
Petals
mm.
—3
mm
long.
to
up
5
by
5 —7
by
i\—2
Ovaries
4
c.
mm.
by
0.7
mm.
with
style
Mt.
(A, BO, L,
level,
sea
ssp.
(BO,
along
Victoria
Elmer
Pulgar:
Mts., Karaniogan R.:
13074
Sulit
(A, BM, BO, CAL, E, K,
SING).
river
in
forest,
and from
150
m
altitude
mm
1—2
mm
holo, S), Celebes,
i|-
—2
mm
Matana
latis; petalis
longis; antheris 2-J,-— 3
mm
\ —f
nov.
L
longis
sessile.
Distribution: Celebes.
177
(A, BO, K); Panacan,
Princesa,
riverbank.
mm
1 —2
B.S. 903
Puerto
P.N.H.
■
mm
\—f
latis; stylis
mm;
Lao
but
lowlands,
mm
from
Cymes 3-flowered,
Stamens
Gopaladen
(BM,
7711
Distr.,
(L).
undang (Sambas), sĕmukau,
mata
filaments;
near
—
fdamentis
mm
22187
S
Paie S .17904
Marintaman
Patrick San
&
Bt.
Brunig
Brunei,
—
' 2209
long
kjellbergii Kanis,
Type: Kjellberg
Flos
Baloh:
Res.,
Labis:
SING); Kuching Distr.:
near
Bintulu
(SING).
Hallier B
mm
Edano
Reported
rocky hillside
ssp.
Park:
L,
saholat, k. topa (Kota Pinang Distr.);
twice
sessile.
Foxworthy
BO, L,
(A,
L, P, U); Bacungan:
on
Kuantan
s
98563
SING); Changi: Ridley
K,
656
Palawan.
PHILIPPINES. Palawan:
P.N.H.
K.F.N.
Kang: Ridley 5896
(K, L);
Madjang: Teysmann
East
—
(Elm.) Kanis,
3-flowered,
Distribution:
Nat.
Res., Sipitang Dist.,
(Belitung); Malay
reported
foxworthyi
Stamens
C
Gopeng: King
podsols.
Use: The wood is
ssp.
Bako
S 3054
from peat-swamp forests and from kerangas
Cymes
Robinson &
Chang
&
Kochummen
Pahang,
—
SING); Lintang path:
L,
Anderson
K);
found in the
Ecology : Normally
b.
Jerai:
(BO, K, L).
thĕharahan (Malacca); Borneo:
soils,
G.
Kinta,
P);
R.:
9148 (BO,
13137
8516 (BO, L); 3rd Div.,
Kenepai:
FI,
(A,
(K,
Anderson
Sumatra: majang, mampat, rampat
laut, k. galugus badak,
kaju
L,
(BO).
14739
Kedah,
SING). —Johore,
L,
Choa Chu
Bujang
SING);
Res.: Fabia
For.
(A, BO, K, L).
'295
Kostermans
:
L,
Interior
Leila
P.B. 3472
Senibung:
49'9
Baram:
(BO, L);
15328
P'
Anderson
Sabah,
Res.,
(FI, K),
Semitau: Hallier
Sanga
—
K,
Ridley 4807 (BM,
Lemidin:
(BM,
Purseglove
Res.:
—
L, SING).
NIFS
Paloh:
NIFS bb
Telok F.
876 (SING).
(SING);
Road:
S.
(K);
9715
(K); 4th Div., Bg.
8704
27988 (K, L);
reg.,
3284 (BO, L); Djebus: Teysmantt
99967 (K), Cheang
6396 (BM,
(A, BM, CAL,
1957
Distr., S.
2704
Brooke
L, SING);
Distr., Saribas For.
Bruti
coll.
Alvins
L);
1192
Goodenough
Div., Lundu
Betong
S
Gul:
King's
K.F.N..
Selangor, Klang,
—
(SING); Tampinis
s.n.
S 12052
Brunig
H. B.
Pandan:
SING).
2224
(BM,
1021
Ridley
P,
Teysmann
Kochummen
Larut:
Perak,
—
K.D.
Maingay
Singapore, Tg.
—
1968
i,
Belitung, Tg.
—
(SING),
20734
Yeop 835 (K, SING); Praman, Pekan: Ridley
(SING).
Banka:
—
No.
XVI,
Thailand, Surat Circle, Tako-Langsuan: Put 1738 (K).
4673 (BM, BO, CAL,
(K).
SING).
11823 (BO)-
(K, SING), Mustafa
5989
(SING),
BO, K,
(BM,
14530
Rindik:
(BO, L);
s.n.
VOL.
longis.
Sepals
long
long
Fructus ad
4
—
filaments;
by
5
mm
—2
anthers
during anthesis.
mm
Lake,
4—4JI
longis;
longus
mm.
2J —3
Fruits
up
m,
400
fl.
longis
mm
ovariis
c.
\
to
5
long.
by
if —2j
mm
mm
longis
latus.
4 mm
Petals 4
—4J by
mm
18-XI-1929.
4
if
Ovaries
mm.
—mm.
c.
o.j
by
A.
CELEBES.
Malili
Nat.
Bot.
].
up
3 —5
16
Suppl.
7.
to
m
30
—
at
to 2
of
one
by
axillary,
z\
to
up
more
up
to
NIFS bb
little
a
or
(L),
E.
3
(BO, L, S).
2808
altitude
m
228
TGH
10768 (L) ;
:
the
at
N.
237
of
Bot.
edge
4515
Schodde
Sogeri :
3
(1857)
A.
51;
A.
U.S.
made
Sogeri
laciniate.
Leaves
by f
mm,
5 —15
by
Torus
bracts
at
by
i
linguiform,
in
cm
mm;
those
fruit,
mm.
mm
c.
style 1^—2
0,
mm
—4f by if—if
Stamens with
apex.
0.5
3
mostly growing
high, \—§
mm
elliptic,
to
Inflorescences
broadly
up to
J
c.
acute
cm,
5-flowered, shortened,
long rachis
long,
cm
0.6
c.
to
ij— 5
entire.
margin
mm
3037
or
ofSarmi: Koster BW
'
S.
4816 (K, L, SING);
and Lake
Bupul
R.,
Fly
Lake
(BO, K, L, SING);
371
long,
mm
(BM,
E,
Royen
van
Brass
Daviumbu:
Distr., Sogeri
Kokoda
trail,
Gjellerup
519
(BO, K,
4750
7910
12262,
(Hollandia):
KotaBaru
ofBivak Hollandia:
Warn:
Central
K, L);
of Sorong: Schram B W'
8085 (L);
L,
(A, BO, L);
reg.:
Forbes
Ower's Corner:
208
Hartley
(K, L).
jobias (Je); obaisang (Mooi);
rainforest
peat,
on
on
serukdeho
flat country
sites which
New Gen. Pi.
Gray,
Bot.
Expl. Exp.
Arn. Arb.
Smith, J.
C.
v.
Queensl. Pi.
Campylopora
Sc. Nat. Bot. VIII,
apex,
New Guinea,
to
(Manikiong).
slopes,
steep
be inundated
may
to
up
during
the
season.
B. australiana F.
Rockingham
lamina
base,
acute
mm,
Middle
P),
primary
clay, sand,
on
SE.
B W'
Distr.,
FI, K, L,
1
up
to 20
m
M., Fragm.
(1913)
(1902)
Bay,
up
of
a
a
to
3- to
4
36 (1955)
(1865)
—
—
29;
(1870)
284.
—
offprint of
6,
t.
t.
42;
Bay,
v.
1
von
Mueller
s.n.
(A, K,
4
Ac.
(1857)
10—14,
errore
Expl. Exp.
s.n.
1838—42.
(1899)
222;
M., Fragm.
Bull. Mus. Hist. Nat. Paris 8
Type:
Ann.
2—7,
U.S.
fl. & fr.
australiana F.
Proc. Am.
Walp.,
248, f.
Type:
Bailey, Queensl. Fl.
Gomphia
Tiegh.,
404.
(1902)
MEL
Compr.
5
(1865)
547;
Ann.
holo?), Queens-
1865.
up
to
8
by
(ovate-)lanceolate,
little tapering
few
v.
high. Stipules
(ovate-)oblong
acute or
fl.
5
f. 70.
85,
australiana
16
(1853)
(1854) 362,
Vanua Levu, Sandalwood
(A holo, K, P), Fiji Islands,
Tree,
It
c.
Seem., Fl. Vit. (1865) 34; Schnizl., Iconogr.
land,
Ann.
667; Rendle,
mm.
5
between
R.,
R.: NGF
(BM,
Reported from
altitude,
linearis;
—
nud.;
nom.
(1912)
8
holo),
Vogelkop Peninsula, Warsamson Plain, E.
(L); BodenR.,
Western
Brackenridgea nitida
Cat.
6
axillary;
ovate
Ovaries
by
6
to
up
Distr.,
Laloki
(BM, K, L),
Arts& Sc.
421;
P
to
up
at
tier.
Sepals
mm.
Vernacularnames: W. New Guinea:
Ecology
wettest
(1902) 46,
Guinea
lanceolate,
to
one
—4f by 1—if
Fruits
Guinea,
Distr.,
Moresby
Stipules
tapering
in
0.
mm
r| by £
c.
BW 12793
New
—
(BM),
49
49
New Guinea.
Merauke
L, U);
m
Nova
sometimes when
25646 (A, BO, K, SING); Versteegh
K,
SING).
29.
Area
Kjellberg
400
16
J. Bot.
laciniate; pedicels
4
NEW GUINEA. W. New Guinea,
Moll BW'12761
cm.
55
each other
to
high,
mm
mm.
3
to
up
less
or
fixed close
Distribution:
B.
Lake:
(BM, FI, K, L,
lamina oblong
terminal,
long filaments; anthers
5.
Matana
simultaneously, the 2—5 (—10)
Petals obovate-lanceolate,
750
Indo-Pacific
made up of a varying number of mostly 3-
when
2 mm,
cyme
in fruit
in fruit
in Morot,
Forbes 237
Type:
acute
apex,
vegetatively
Port
the
and from
swamp
descr.; Pulle,
395,
dbh.
high;
sessile cymes, flowering
up
Tiegh.
v.
(1902)
long petiole;
mm
terminal and
(BO,
level in.
sea
forbesii
VIII,
acuminate
on
of
1885—6.
Tree,
with
Bot.
(1923)
fl.
reg.,
to
Ochnaceae
lake.
a
Sc.
the
of
revision
Lampea: Kjellberg 2046 (BO, L, S);
reg.,
Brackenridgea
4.
A
Reported from
Ecology:
of
Kanis:
at
1 mm.
5
base, margin
many-flowered
—18
Leaves with 3 —8
by
entire.
cymes,
2—cm,
Inflorescences
sessile
or
mm
acute to
long petiole;
acuminate
at
terminal and axillary,
shortly peduncled, flowering
BLUMEA
VOL.
No.
XVI,
the
simultaneous,
i
c.
6
long,
cm
Stamens with
style i\—3
c.
mm
Remark: For
obovate,
fruit
in
2
subspecies
a
discussion
to
up
Anthers
c.
1.
Anthers
c.
a.
by J
3
on
australiana
ssp.
sessile
Cymes
sepals
mm;
(F.
5 —6
by f
3
Style 2|
mm.
Distribution: NE.
Bellcnden
White
8992
Ker:
by
long
—2f
2
Distribution
FIJI.
Home
Peak:
Brass 2313
Cibbs
mm.
Petals 35 —4 J, by
:
596 (A, K, P).
A.
C.
—
Viti
Greenwood
K,
P,
Dept. Agr.
446A (K);
Smith
Expl. Exp.
S);
12860
Vernacular
S. Nausori
483,
W. of Mbutha
nitida
ssp.
M.
v.
to
3-
4^—5 by 2\—3
mm.
many-
Anthers
Tableland, Boonjie: Kajewski
Mueller
von
1273
(A,
Mt.
(A, K);
s.n.
BM,
Spec:
(light) rainforest.
in
altitude,
Anthers
by J,
c. 2
Style 1J—mm
mm.
Vanua
—
R.:
Parham
Seemann
483A (K);
Distr.,
Bay:
A.
mbelembele (Vanua
sea
level
country, in
Hook., Lond. J.
Home
Levu: Parham
(K);
13429
(K);
Bola
E. Yanawai R.
Smith
1
(13M);
Mbua
lower Wainunu R.:
Natua:
(A, BM, K, P);
93
Naketei:
C.
Navosa: Greenwood
Rakiraki:
(A, K);
893
92
A. C.
S.
Tothill
slope
drainage: Degener
Mt.
Smith
6643
1834
813 (A, BO, K, S);? Seagaga:
(A,
(A, K, L, S),
446
(A, BO, K),
Numbuiloa:
& Ordonez
483C
& Ordotiez
Sandalwood
Distr.,
A. C.
Smith
Nanduri:
(K);
Degener
14113
Watkins
A.
C.
(A,K);
383
(K);
(K).
name:
Brackenridgea
Jack;
Petals
F.
branches,
Highlands, Nandronga and
Na Sau: Parham 35
Thakaundrove
to
up
dry
500
open
elegantissima
Wall, in Roxb., Fl. Ind.
in
b.
long
mm
australiana
ssp.
many-flowered. Sepals mostly obovate,
mm.
Wai-wai:
L, S);
Macuata Coast:
S);
313
From
rolling
m
5- to
if—2f
Levu).
m,
in dense and
Bot.
Benn. in Hook,
2
5
(1824)
Br.
forest,
in
patches
of forest in
SPECIES
(Wall.) Kanis,
305;
in
Hook.,
(1846) 647; ibid.
f., Fl.
open
hillside thickets, and in scrub vegetation.
DOUBTFUL
6.
3
a.
....
long
mm
Rockingham Bay:
and 700
483B (K).
(A, K, P);
Greenwood
K, L,
Home
Ecology :
open
(K);
Peninsula,
Rambi I.:
s.n.
Levu,
K,
Macuata Distr., Korovuli
Lambasa:
6331 (A,
Natewan
to
mm.
long
mm
Islands.
Fiji
13693 (A, K); Penang:
BO,
description.
B. nitida A. Gray.
—
4608 (A,
U.S.
mm.
J
under the generic
B. australiana
—
up
Atherton
(A);
(BM, K);
6322
Smith
Bay:
mm;
Is.
Fiji
see
mm.
0.6
anthesis.
during
(A, K, L),
4 J—5
nov.
3 —4
Cymes sessile, with shortened branches,
4—5
by
0.8
high,
2—3
long during anthesis.
from 300, 600,
Reported
nitida.
ssp.
c.
pedicels
mm
SUBSPECIES
with
by
by
6
3
3^ —5 by
Ovaries
to
up
to
up
terminal,
mm;
(A, P).
Ecology:
b.
Fruits
i|
Queensland.
Thornton
QUEENSLAND.
K, L, P);
stat.
in fruit
0,
mm.
and the
long; petals 3J—4J
—6
5
mm
—
THE
long; petals
mm
mm
4—5
flowered. Sepals mostly ovate,
c.
TO
M.) Kanis,
v.
—f
the date of publication
shortly peduncled,
or
long.
2J, by
Petals obovate,
by
2—3
to
up
mm
i
mm.
Queensland
in NE.
sepals
mm;
by J
2
2—4
mm
5
high,
mm
by
—6
4
KEY
I.
I
c.
when
vegetatively
on
linguiform,
to
long filaments; anthers
long,
mm
Distribution:
in fruit. Torus
cm
ovate to
\
mostly growing
axillary; bracts triangular
to 2
Sepals
0.
mm
up
rachis
long
mm
3-—10
sometimes when
1968
i,
50
Ind.
1
6
comb.
(1847)
(1875)
nov.
Bot. Misc.
526,
2, err.
err.
itt
in
—
2
syu.
syn.
Euthemis ? elegantissima
(1831)
of
77,
note;
Gomphia
idem; King,
Planch,
sumatrana
J. As.
Soc.
Kanis :
A.
Beng. 52,11 (1893)
Mai. Pen.
syn.
of
idem.
Gomphia
Euthemis?
Wallich
The
Jack; King, I.e.
2518
what
this
genus
in
233, err.
He also
named Euthemis pulcherrima.
Pierre 5877, 'Insula
the views of
In
The former
Under
be distinct. In
to
concluded from both Wallich's
is rather
however,
uncertain,
obviously
taken from
by
Singapore
me
in
other hand I
occurring in Singapore
The combination B. elegantissima
all other specific
(Wall.)
in the
names
Schreb., Gen. Pi. ed. 8,
Paris
Ind. Bat.
1
(1875)
(1859) 675;
1, 2
Kurz,
525;
Fl.
Ridl.,
Engl.,
Planch,
Jahrb.
Bot.
in E.
&
Bot.
16
(1902)
40,
33
18
(1903)
(1893)
17
194,
Ochna Linne,
Enc.
Meth.
excl.
syn.
Ouratea
Pi.
1
=
Mus.
(1834)
15
&
153;
Endl.,
197,
A.
made
21
Gray.
On
the
B.
palustris
it has
priority
to
2.
Ouratea
ex
5
2,
21
(1754)
Adans.,
v.
Fl. Br. Ind.
N.I.
—
232;
sect.
‘Gruppe
Reticulatae’
Reticulatae Gilg
Tiegh.
73.
(1890)
Gomphia
in Morot,
Reticulatae
sect.
(1925)
1
(1893)
Sc. Nat. Bot. VIII,
(Gaertn.)
serrata
/,
Beng. 62, II
422.
Ann.
1142; Miq., Fl.
in Hook.
420,
—
Ann.
s.s. ;
ed.
(1799) 569; DC.,
(1840)
Palaeouratea subsect.
Fam. ed.
Gomphia
2
Campylospermum
—
Pi.
513; Gen.
(1901)
6
Sem. Pi.
16
(1811)
Gen. Pi.
(1775) 397] Baill.,
1
155;
(1931)
cf. Kanis, Taxon
17
identity,
16
J.
(1902)
Gilg,
Bot.
Type species:
—
Kanis.
excl. typus; Lamk.,
229, p.p.,
Fam. Pi.
2
(1763)
364,
p.p.,
illeg.
Ceyl.
Paris
sect.
Soc.
(1967)
16
4,
Benn.
As.
(1847)
6
Jabotapita Plum,
cf. Kanis, Taxon
illeg.,
as
sp.
large, serrulate leaves
(Planch.)
Gen. Pi.
subg. Palaeouratea
Nat. Pfl.
Hist. Pi. Gui. Fr.
nom.
142.
Ouratea
—
DC.
—
Bot.
illeg.; ibid.
[non Aubl.,
nom. cons.,
specific
Boerl., Handl. Fl.
206;
King, J.
Ouratea
—
notn.
Handb. Fl.
Its
Jack,
sumatrana
Brackenridgea
a
made, because
Pi. ed.
(1862) 318;
1
(1877)
234;
(1895)
6
Ochna L.,
DC., Ann.
Or.
80.
(1753)
509.
Meesia Gaertn., Fruct.
150,
28.
1
Bartell., Malpighia
in Trim.,
B. hookeri
P:
I.
365; Kanis, Taxon
nom.
(Lamk.)
Sp.
is found in
XI-1877
much the collection B
very
Kanis has been
Pi.
Burma
(1893)
1
From this I
in Wallich's edition
name
specimen.
type
Willd., Sp.
p.p.;
Gen.
H.,
in E. & P.,
(1797)
4
s.l.,
(1903)
250;
Gomphia zeylanica
same
certainly
it is
(1824) 736; Endl.,
1
Br.
as
GOMPHIA
Hook., Lond. J.
in
P., Nat. Pfl. Fam. 3,
296; ibid.
Jahrb.
173,
&
Fl.
(1922)
1
291,
Prod.
B.
For.
Mai. Pen.
Gomphiastrum
p.p.;
(1789)
I
414;
Trim., Handb. Fl. Ceyl.
174;
193,
(1811)
17
to
name
doubt
genus.
3.
Mus.
I refer
in
syn.
for the collection
conspecific with Gomphia
and
the
some
exclude the possibility that Wallich's species belongs
Bartell., both species
over
name
validly published
It resembles
that
seedling
a
manuscript
the material is sterile and with
as
under
Catalogue
2318 hue referenda'?'
No.
opinion
my
description
juvenile plant.
a
of
can not
is
authors this species
most
err.
in
IX-1822.
ad chasserianum monticolum',
Singapore,
but King thought it
was
err.
2803 he listed ‘Gomphia sumatrensis
no.
alternative
name
/, I.e.,
idem; Bartell., I.e.,
A similar collection under the
Flora Indica.
Roxburgh's
an
idem-, Ridl., Fl.
syn.
‘Gomphia sp.?’, expressing
added
species really belongs.
in
err.
51
Benn. in Hook,
syn.
ster.
Jack', adding the remark 'ad Euthemis elegantissima
of
160,
ex
Wallich's
in
Area
Indo-Pacific
(1901)
15
listed
was
conclude that Euthemis elegantissima is
the
of
2318 (K holo), Singapore,
collection
‘Euthemis? pulcherrima Wall.'
to
Ochnaccae
pulcherrima Wall,
—
sumatrana
Type:
—
Remarks:
the
Bartell., Malpighia
235;
(1922) 368.
1
of
A revision
Lecomte, Fl.
42;
1
(1788)
(1967)
16
421;
Back.
&
—
419;
Prod.
1
1142;
rej.;
Walkera
(1967)
(1840)
Bakli. f, Fl.
344, nom.
419.
Hist. Pi.
non
Java
Schreb.,
737;
W.
1
1
(1911)
(1963)
&
703;
Alst.
327.
Muse.
(1801)
Gen. Pi. ed. 8,1
(1789)
Hedw., Sp.
Willd., Sp. Pi.
(1824)
(1873) 367, emend.,
4
Gen. Indo-Ch.
ed.
4,
1
(1797)
1145;
A., Prod. Fl. Pen. Ind.
Planch, in Hook., Lond. J. Bot.
5
(1846)
BLUMEA
52
Walkena Mill,
non
593;
Gaertn.
serrata
Cercinia
162.
Campylocercum
Tiegh.
v.
Leaves
wavy
base,
with
not
leaving
in fruit.
globular
2
reduced
±
apical
1
Ovaries
pores.
One
Distribution:
in Africa
S.
to
up
conspicuous
The
Only
who has
(Planchon,
two
In
a
American
and
to
the
which
name
including those of
a
type
species
Gomphia
differs
and seed
free, subulate
In Ouratea
caducous,
was
the
or
ones.
s.s.
and
articulate
everwet
an
a
distinction
the
genera.
in
at
mostly
to
±
not
(Bull.
for
a
it
long
was
in several
subgeneric
or
were
level
respectively
often divided
1895, 1925).
criteria and
Soc. Bot.
name
to
propose
Suisse 73, 1963,
Tiegh., Rhabdo-
v.
Ouratea Aubl. for the
characters, derived from embryology, leaf anatomy,
to
me.
Recently
(1967)
genus
as
as
a
I
the Old
segregate
lectotype
have proposed
World
species,
from Ochna L.,
Gomphia
zeylanica
of Ochna jabotapita L., the type species of Ochna L.
Gilg
(1925)
from
intra-axillary,
the ovule and seed
as
the ideas of Van
Worlds
Africa, Campylospermum
the
use
published
genera,
sectional
Tieghem's
Farron
in
dry
not
are
coloured.
not
accept
1893; Gilg,
the American Ouratea by
connate,
Moreover, the sepals in Gomphia
scale-like, rarely
moderately
a
are
the African species
indicated. I have chosen
segregate
30—35
(1775), with which
Gomphia Schreb. for the majority of
a
stigma punctiform.
or
and New
Old
(Engler,
quite convincing
never
creamy,
opening with
and Celebes;
torus
Aublet
reconsider Van
genera
new
base. Flowers
turning sub-
yellow,
5,
has been commonly
poorly defined
34
subsections
partly
are
terminal
few bracts
a
Thailand, Indo-China,
E.
climate
and
calyx
Ouratea
from
species
different
irregular
blue-black when ripe.
Most authors did
into
to
at
anthers
short;
in
possibly by birds, but the fruits
since
by
However,
tendency
according
and
sometimes
in fruit. Petals
India, Ceylon,
Asia. Schreber has described his
which is
(Lamk.) DC.,
ovule
3
He used
species.
blastogeny,
resurrect
but
or
united,
and/or
and sometimes
very
Tiegh., and Idertia Farr., correctly reserving
v.
lateral
camptotropous, epitropous;
with
areas
moderate division into distinct
196 —217) recognized
phyllum
a
Nat.
conspicuous, somewhat
a
Inflorescences
enlarging
1895, 1925). Furthermore,
there is
Sc.
Madagascar.
replaced
1907.
three sections
years
and
nerve
filaments terete,
synonymous.
between
(1907)
straight, curving upward
pedicels filiform,
SW. Peninsular
Ouratea s.l.
and
1847; Gilg,
or
recent
more
scars;
gynophore,
Brackenridgea,
was
be
to
split
1901
often made
it
(1902)
small, intrapetiolarly
almost
persistent with
±
Gomphia Schreb. (1789)
name
gradually
between
papers
into
Stipules
parallel,
ones.
altitude. Dispersal
Ochna and
generally considered
Tieghem,
in
tropical
m.
8
5
Kanis.
Ann.
546;
distance, veinlets reticulate, joining
the primary
of the Sahara and
to
(Gaertn.)
serrata
(1902)
8
Paris
ibid. IX,
31;
Malay Peninsula, Borneo, the Philippines,
1500
in
as
Remarks:
was
at some
obovoid; ovule
species
the
Confined
Ecology:
monsoon,
time.
Nat.
inconspicuous marginal
whorl;
I
(1903)
18
Gomphia
tinged pinkish, enlarging
5,
5,
ibid.
=
—2(—5) yellowish, turning dark purple
Hainan, Sumatra,
species
close,
branches; peduncle
in
io
Meesia
Type species:
—
Kanis.
nerves
5-ribbed
Sepals
white. Stamens
Tiegh.
branches.
distinct annulus of
a
131.
198; Bull. Mus. Hist. Nat. Paris
309;
Hist.
(Gaertn.)
to
short, columnar,
a
Fruits
an
194,
Mus.
(1763)
53
Kanis.
(1902)
v.
spreading
the margin
to
parallel
nerves
with
thyrses,
or
with
chartaceous,
parallel
nerve
secondary
at
304;
the margin, forming
near
16
1968
i,
ibid. 18 (1903) 28. —Type species: Campylocercum striatum
serrata
treelets
or
caducuous.
194,
Gomphia
=
Shrubs
Bull.
Tiegh.,
v.
(1902)
16
Bot. VIII,
Trans.
(1902)
16
Cercinia thorelii
Type species:
—
Bot.
J.
Sc. Nat. Bot. VIII,
No.
XVI,
(Gaertn.)
serrata
in Morot,
Tiegh.
v.
376; Ann.
Ehret, Phil.
ex
Gomphia
=
VOL.
are
are
incised
and subulate.
curved
lateral,
persistent.
straight, the stipules
persistent
a
stipules, rarely
never
Sepals
connate,
are
mostly small,
caducous.
A.
Campylospermum
Kanis
species
of the latter
Tieghem (1902) has
v.
of Gomphia
Schreb.
generic
wrongly defined
was
certainly
it has been
Walkera
which
whereas
not
(1789)
considered
the
as
Gomphia
(1797)
Ceyl. Pi.
50—51.
Pi.
(1824)
W.
(1849)
Benn. in
Diet.
194.
(1906)
ex
Thw.,
Sylv.
&
(1921) 387;
6
3
(1895)
illeg.
—
ed.
Ochna
angustifolium
v.
14.
Mus. Paris
‘ceylanica
Tiegh.,
19.
Fl.
—
’;
17
W.
Ceyl.
France
Filip.
(1811)
&
6
42;
f.
legitimate
a
This
was
name
status
Walkera
was
Schreb.
under Art.
Meesia
75
(1824)
1
Pi. ed.
(1834)
1
Taxon
Robs.,
4,
I
Moon, Cat.
737;
Ind. Or.
Pen.
serrata
Gaertn.,
serrata
Willd., Spec.
153;
(1962)
11
1
non
Fl.
(1916)
2
Ridl.,
68;
Gilg,
Bot.
Vahl
J.
16
Bot.
(1903)
9
74;
107;
&
Sol.,
Ind.
Brand.,
197;
106.
Sc.
&
P.,
1
Str. Br. R. As. Soc.
(1930) 76;
Ann.
Ann. Sc.
J.
in E.
Gamb., Fl. Madr.
Kirt.
(1903) 269,
33
(1891)
i
Vid.
39;
Watt,
79;
(1886) 607; Gilg
34; Merr.,
197;
411;
(1880)
Back., Schoolfl. Java (1911)
Kew Bull.
(1902)
235;
(1864)
1
(1873)
32
App.
(1893)
(1902)
Walp.,
3;
Filip. (1886)
1
1
(1912)
2
N.I.
2
(1839)
&
nom.
Basu,
homon.
Campylospermum
—
Nat.
Nat. Bot.
Bot.
VIII,
VIII,
18
16
(1903)
(C holo, P) Ind. Or., Ceylon, fl.
Meth.
(1797)
4
(1824) 736;
1
9
1
512.
(1834)
(1903)
153.
123,
t.
Syst. Veg.
—
77; Ann.
Kew Bull.
(1737)
Gomphia zeylanica
—
Spreng.,
Ann. R. Bot. Gard. Perad.
Airy Shaw,
Thes. Zeyl.
214;
Jahrb.
O. Kuntze, Rev. Gen. Pi.
Enc.
Ceyl.
(1909)
Java
(1847)
6
Nov.
Filip.
Pres. Bomb.
34,
(1923)
3
516;
Morot,
Fl.
Handb. Fl.
Exk.
Bot.
Rev. Pi. Vase.
19;
Cat. Pi. Bomb.
Tijdschr.
Nat.
(1799)
2
Drury, Handb. Ind. Fl.
71;
Laness., Pi. Util. Col. Fr.
ex
Koord.,
Fl. Pi.
Grah.,
152;
8,
Syst. Veg.
(1824) 736; Spreng.,
1
F.-Vill.,
525;
(1883)
Willd., Spec. Pi. ed.
49;
Scheff.,
4;
Bomb. & Sind
Prod.
415;
Gomphia malabarica DC.,
—
Hook., Lond. J.
(1858)
A., Prod. Fl. Pen. Ind. Or.
(1931)
Prod.
(1834)
1
in
1
Bot. Centralbl.
zeylanica Alst.,
holo?) cf.
417;
Cooke, Fl.
101;
142;
in Hb.
Lamk.,
422.
serrata
Ouratea
Prod.
533; Trim.,
(1933)
in
8,
Bull. Mus. Hist. Nat. Paris
Ouratea
1801
years,
Ehret (1763)
ex
Prod. Fl.
—
(1791)
2
Pi.
Mus. Hist. Nat. Paris
Type: Koenig
—
1
Tiegh.
Ochna zeylanica
v.
2,
angustifolia
(1902) 298; Bull.
593.
(1875)
1
For. Fl.
(1895)
t.
51,
(1890)
Philipp.
En.
Pi.
Gaertn.
(1788)
Gaertn.
Meesia
In later
Furthermore,
(1967)
421;
A.,
&
Bot.
Ceyl.
En.
Gen. Pi.
(1915) 167; Hall./, Beih.
86
17
angustifolia Baill.
3,
W.
(1846)
5
16
Walkera
(1811)
17
(1811)
Br. Ind.
Talb.,
129;
Fam.
Ind. Med.
be used since
can not
Coll. Sem. Hort. L.-B. (L holo) Ceylon, 'Walkaera', fl. & fr.
(1869)
3
Philipp.
Ouratea
—
for
Meesia
Gaertn.
—
(1842) 526; Planch,
1
182;
Fam.
Cum.
Nat. Pfl.
Bot.
6.
A., Prod. Fl. Pen. Ind. Or.
Prod. Ind.
Econ.
Trees
Paris
‘Walkeria’;
err.
Hook./, Fl.
Vidal, Syn.
Phan.
f.
70,
t.
344,
Ann. Mus.
17,
&
Fl.
Bedd.,
221;
it
and hence illegitimate.
Meesia
Taxon
(Gaertn.) Kanis,
(1788)
1
38; Walp., Repert.
1
flower. Meesia
10 stamens per
new name
later homonym of Walkeria Mill,
Ann. Mus. Paris
(1825) 318;
a
as
superfluous
angustifolia Vahl, Symb.
569; DC.,
Ann.
a
Type: Koenig
—
Gomphia
flower. The holo-
stamens per
5
contain
for Musci, making
rejection of
Hook., Lond. J.
in
synonym
Code.
DC.,
1145;
Planch,
to
of Meesia Hedw.
homonym
became
later
serrata
Sem.
introduced
was
later
a
Schreb.
the
by
should be considered
&
superfluous
a
Gomphia Schreb., but
of
synonym
starting point
Walkera
of the present
1.
older
an
Schreb.
changed
Fruct.
53
rejected.
was
appointed
becomes
having
as
of its only species has been found
type specimen
Gaertn. is
Area
Indo-Pacific
been typified. Since it includes the type
never
automatically
it
(1789),
the
of
name.
(1788)
Meesia Gaertn.
the Ochnaceae
of
A revision
:
(1940)
11
2
Ann.
318,
err.
Campylospermum zeylanicum
Sc. Nat. Bot.
(1929)
249.
DC.,
(1825)
—
209;
Type:
VIII,
18
(1903)
in Trim., Handb.
Burman
(Inst,
de
56.
Arm. Mus. Paris 17
(1811) 416; Prod,
x
(1824) 736; Spreng.,
BLUMEA
54
Syst. Veg.
cf.
(1825)
2
(1903)
Nat. Paris 9
(1846)
N.I.
J.
Ridl.,
Benn.
4";
Beng.
44,
Mai.
1
Pen.
(1925)
5
(1961)
(1921) 387; Ridl.,
—
J.
in E.
&
Morot,
J.
Beccari
As.
525,
(1930) 76.
Beng.
30,
2 3 3 ">
Ridl.,
Chit,
&
(1891)
1
Fl.
79;
List
106.
Bartell., Malpighia
142;
—
15
Merr., J. Str. Br. R. As. Soc.
35;
Campylospermum
—
Kiirz,
(1893)
129;
Hundl.
226;
(1895)
(1916)
’;
App. A,
62, II
O. Kuntze, Rev. Gen. Pi.
3, 6
181;
Tijdschr.
16; Ridl., Kew Bull. (1925)
(1952)
33
77;
(1849)
1
Nat.
‘sumatrensis
err.
Ind. Trees (1906)
Suppl.
2
34,
Soc.
Ann. Sc. Nat. Bot. VIII,
197;
Ann.
Scheff.,
534;
(1875)
1
King, J.
(1924)
sumatrana
Bartell., Malpighia
Kew Bull.
Ouratea
(1902)
16
Bot.
P.B.
P.B. 3414
(Fl
3138
neriifolium
197;
Ann.
holo, K, P)
sumatranum
Tiegh.
v.
(1902) 298; ibid.
16
Tiegh.,
Ann.
Ouratea beccariana
Nat. Bot.
S. Kanta, fr.
18
(1903)
III-1867.
158,
t.
301.
—
(1902)
sphaln.
7,
Kew Bull.
16
(1902)
16
VIII,
Str. Br. R. As. Soc.
borneense
v.
—
Tiegh.
in
Lectotype:
Paratype:
Beccari
(1940)
304.
249.
‘neerifolia’;
Merr.,
—
Beccari P.B. 3347
Type:
—
Campylocercum
fl. IV-1867.
Bartell., Malpighia
Ann. Sc. Nat.
J.
6; Merr.,
V-1867.
(1901)
15
t.
Marop, fl.
Sarawak,
Sc. Nat. Bot. VIII,
(1921) 387; Airy Shaw,
Tiegh.,
156,
Campylospermum
—
(1921) 387; Airy Shaw,
86
(Fl holo) Sarawak, Tg. Datu,
86
Sc.
neriifolia Bartell., Malpighia
v.
(1901)
15
(1930) 76.
Pontianak Prov.,
(FI holo)
J. Str. Br. R. As. Soc.
v.
209,
(1830)
2
Type: Jack, Sumatra, Sibooru f.
(1921) 387; Ridl.,
Soc.
Br. Ind.
206;
Bot.
preserved,
Walp., Repert.
t. 712;
Walp.,
2;
(i860)
Misc.
Bot.
(1845)
4
(1847)
6
1
P., Nat. Pfl. Fam.
Kew Bull.
(1902)
16
Bot.
Ouratea borneensis
86
Ochna
—
II,
(1893) 285; Brand.,
3
Hook.,
29;
Ic. Pi.
Merr., J. Arn. Arb.
296;
42.
Gilg.
sumatrana
in Morot,
21.
Bot.
(1877)
1
Hist.
Mus.
not
Prei. Rep. For. & Veg. Pegu (1875)
160; Hall./, Beih. Bot. Centralbl.
(1901)
86
141;
(1922) 365; Rendle, ƒ.
Trees &c. ed. 3
Ouratea
Hook./, Fl.
in
(1875)
II
Trans. Linn. Soc.
Mai. Pen.
Fl.
(1859) 675; ibid., Suppl.
1,2
Fl. Br. Burma
35; For.
App. B,
(1821)
5
Hook.,
in
Ticgh., Bull.
v.
—Type: prob.
19.
‘poeatsjetti’.
52,
I,
(1903)
18
Planch, in Hook., Lond. J. Bot.
399;
Soc.
t.
Misc.
(1842) 526; Planch,
1
Ind. Bat.
(1873)
32
As.
(1685)
5
1968
I,
malabaricum
Campylospermum
Jack, Mai.
sumatrana
Walp., Repert.
5
—
No.
XVI,
Ann. Sc. Nat. Bot. VIII,
Hort. Malab.
Rheede,
Gomphia
Miq., Fl.
318.
77;
VOL.
(1901)
15
Kew Bull.
Bot. VIII,
(1902)
16
301.
t.
159,
(1940)
249.
Campylospermum
Type:
—
Merr., J. Str. Br. R.
9;
—
As.
beccarianum
(FI holo,
Beccari P.B. 4025
_
K) Sarawak, Bintulu,
Cercinia thorelii
8
v.
Ann.
(1902) 376;
fr. IX-1867.
Tiegh.
Ouratea thorelii Lecomte, Fl.
and
fr.
178.
—
Bien-hoa,
Type:
Gen. Indo-Ch.
Thorel 643
1862—66.
—
i
310;
(191 1)
Godefroy
P
ibid.
704;
(1946) 671, f.
1
(C, CAL, E,
Paratype:
(1902) 198; Bull.
(1902)
16
VIII,
246; Gagn., Fl. Gén. Indo-Ch. Suppl.
(i960)
16
in Morot, J. Bot.
Nat. Bot.
Sc.
82,
IX,
Mus. Hist. Nat. Paris
5
(1907)
Fl. Siam. En.
13; Ho
&
184.
1
Du'o'ng, Fl.
—
(1931)
Vietn.
between
holo) Cochinchina,
355
162,
Craib,
Saigon
Route de Pursat,
(K, P) Cambodia,
VI-1875.
Campylocercum
striatum
striata Lecomte, Fl.
Masam., Fl. Kait. (1943)
7 —12.
—
Campylocercum
3073
Henyala,
fr.
zollinger i
(BM, BO,
v.
Tiegh.,
Type: Harmand
XII-1875.
205;
v.
Ann. Sc.
1
&
16
Ann.
holo)
Gen. Indo-Ch.
Sc. Nat. Bot.
(1902)
304.
Chun, Sunyats.
Suppl.
(P holo) Annam, Tourane, fr.
Tiegh.,
P
Nat. Bot. VIII,
703; Merr.
Fl.
Gagnep.,
291
L,
(1911)
VIII,
Sumatra, Lampongs,
1
2
Ouratea
—
(1935) 282;
(1946) 673, f. 82,
I-i 837.
16
(1902)
305.
—
Type:
between Tarabangi
and
IX-1845.
Cercinia brevis
—
Tiegh.,
Type: Gaudichaud
Zollinger
184.
v.
Gen. Indo-Ch.
131
Ann. Sc.
Nat. Bot.
(BM, E, K,
P
VIII,
holo) Laos,
5
(1907)
Treng,
16
(1902)
310; ibid. IX,
Bassin de Sc-Moun, Mt. S.
A.
Campylospermum
Nat!
Sc.
Kanis
A
:
retinerve
16.
of
Bull. Mus.
Tiegh.,
v.
(1903)
18
Bot. VIII,
the Ochnaceae
of
revision
Type:
—
the
Hist.
Hb.
ex
Area
Indo-Pacific
Nat.
Vaillant
Paris
(P
55
(1903)
9
Ann.
75;
'Gurunda
holo) India,
Karpata'.
Campylospermum vahlianum
Nat."Bot. VIII, 18 (1903)
Nat. Bot. VIII,
Sc.
281; Fl. Mai.
p.p.
fr.
excl.
Pen.
typus.
(1925) 296.
5
—
Hist.
Mus.
Ouratea
Paris
Nat.
P
E, K,
(1903)
9
Kew Bull.
Kew Bull.
(Griff.) Burk.,
crocea
Ann.
(1903) 76;
9
holo) India.
P
Gomphia oblongifolia Ridl.,
—
Wallich 2803 (A, BM,
Type:
—
17.
(C?,
s.n.
Bull.
Ticgh.,
v.
(1903)
18
Paris
Mus. Hist. Nat.
Type: Breyn
—
holo) Singapore
76;
(1925)
(1935) 318,
&
Dindings,
1822.
leschenaultii
Campylospermum
Sc.
Nat!
Bot. VIII,
Nat. Bot.
walkeri
Nat. Bot.
K,
P
holo)
VIII,
rheedei
Campylospermum
48,
t.
Mus.
(1903)
9
76;
Ann.
C.P.
Paris
2412
(1903)
9
77;
Ann.
(BM, BO, CAL,
p.p.
Hist.
Nat.
Paris
(1903)
9
77;
Ann.
Bull.
Mus.
Type: prob.
Hist.
not
Nat.
Paris
preserved,
cf.
(1903)
9
78;
Ann.
Hort. Malab.
Rhccde,
19.
Tiegh.,
v.
(1903)
18
20.
Bull.
Mus.
Hist.
Type: Hombron
—
—
Bull.
Mus.
Nat.
(A,
s.n.
Hist.
P
Paris
(1903)
9
78;
Ann.
holo) Singapore, 1838—40.
Nat.
Type: Derry 987 (CAL,
Paris
(1903)
9
79;
Ann.
holo, SING) Malacca,
P
1892.
VIII,
ft,
fl.
Tiegh., Bull.
v.
(1903)
18
21.
Mus.
Hist.
Nat.
perakense
Sc. Nat. Bot. VIII,
Tiegh., Buil.
v.
(1903)
18
21.
(1903)
9
P
Ann.
79;
Sc.
holo) Perak, Larut,
Mus. Hist.
ft,
0—500
fl.
fr.
&
abbreviatum
Campylospermum
Sc. Nat. Bot. VIII,
(1903)
18
Paris
9
(1903)
80;
Ann.
„
VII-1883.
v.
21.
Nat.
Type: King's coll. 4343 (A, BM, K, L, P holo, SING)
—
„
Perak, Larut,
Paris
Type: Kings coll. 3370 (K, L,
—
IX-1882.
Campylospermum
Campylospermum
fl.
Nat.
Hist.
Thwaites
Bull. Mus.
Ticgh.,
v.
(1903)
Campylospermum kingii
300—500
—
19.
strictum
Bot. VIII,
Nat. Bot.
Nat. Paris
Type: Thwaites C.P. 2412p.p. (P holo?) Ceylon 1854.
—
Tiegh.,
v.
plicatum
18
Campylospermum
Batu Tiga,
Type:
—
Hist.
Ann.
‘tsjocatti’.
Sc. Nat. Bot. VIII,
Nat.
Mus.
(1903) 76;
9
(P holo) Ceylon, fl.
Walker 34 (P holo) Ceylon, fl. 1837.
Type:
Tiegh.,
18.
(1903)
18
Campylospermum
Sc.
18.
v.
(1903)
18
Sc. Nat. Bot. VIII,
Bull.
s.n.
1854.
nodosum
Campylospermum
—
Tiegh., Bull.
v.
(1903)
18
Ceylon,
Sc. Nat. Bot. VIII,
(1685)
18.
Mus. Hist. Nat. Paris
Leschenault
Type:
—
Ticgh.,
v.
thwaitesii
Campylospermum
Sc.
17.
(1903)
18
VIII,
Tiegh., Bull.
v.
(1903)
18
Campylospermum
Sc.
5
17.
wallichianum
Campylospermum
Ann.
Tiegh., Bull.
v.
Sc.
—
Tiegh.,
Bull. Mus. Hist. Nat. Paris
(1903)
9
Type : King's coll. 6398 (P holo) Pcrak, Larut,
80; Ann.
o—100
ft,
IX-1884.
Campylospermum
cumingii
Sc. Nat. Bot. VIII,
Campylocercum
Sc. Nat. Bot.
Canara,
VIII,
Tiegh.,
29.
Bull. Mus.
Hist.
Type: Cuming
—
Nat.
1320 p.p.
Bull. Mus.
Hist.
Nat.
Type: Hohenacker 89
Paris
9
(1903)
80;
Ann.
(P holo) Luzon, Manila,
Paris
9
(1903)
Ann.
P
holo)
(BM, E, K, L,
p.p.
fl.
86;
hohenackeri
18
v.
(1903)
Tiegh., Bull.
29.
—
Mus.
Hist.
Nat.
Type: Hohenacker 89
Paris
p.p.
9
(1903)
86; Ann.
(BM, E, K, L,
P
holo)
Mangalore.
Campylocercum
Nat. Bot.
VIII,
Cercinia
Bot. VIII,
Ceylon,
v.
—
Mangalore.
Nat. Bot. VIII,
Canara,
22.
(1903)
18
Tiegh.,
v.
(1903)
paucifolium
Campylocercum
Sc.
18
fr.
metzii
18
wightii
18
31;
III-1836.
Bull. Mus.
Tiegh.,
29.
Tiegh.,
v.
(1903)
v.
(1903)
Bull.
ibid. IX,
Hist.
Nat.
Metz
2223
(P
Hist.
Nat.
Type:
—
5
Mus.
(1907)
162.
—
Paris
Type:
Paris
9
(1903)
fr.
holo) India,
9
(1903)
Wight
87;
K.D.
86;
Ann.
Sc.
1858.
Ann.
396
p.p.
Sc.
Nat.
(Pholo)
BLUMEA
56
Cercinia doumerii
Tiegh., Bull.
v.
(1903)
18
Bot. VIII,
VOL.
32.
Cercinia annamensis
Ann.
Tiegh.,
v.
Mus. Hist.
Doumer
Type:
—
s.n.
Sc.
No.
XVI,
Nat.
Paris
Cercinia
elongata
Hb. Pierre
=
Nat.
Bot.
Ann. Sc. Nat. Bot. IX,
Tiegh.,
v.
(P holo) Cochinchina,
7026
Gomphia microphylla Ridl., Fl.
Fl. Siam.
Craib,
En.
Dayang Bunting, fl.
Ouratea
Fl. Gén. Indo-Ch.
Bakh. ƒ, Fl.
IX,
Ouratea
163, 184.
Ouratea
(1907) 163, 184.
(1922)
1
365, f. 38.
Bull.
(1926)
Ridl.,
Kew Bull.
Siam.
Fl.
341;
En.
Dansk Bot.
(1931)
1
Ark.
(BM, BO,
var.
(1930) 76.
Type: Ridley
—
Fl. Siam. En.
Craib,
nervosa
holo, SING) Puket,
K
lobopetala Gagnep., Fl.
Gen. Indo-Ch.
s.n.
Shrub
or
tree, up to
scrambling.
to
Stipules
to
finely
—35)
io(—15)
or
pedicels
\—f
sometimes
Fruits
long,
to
in fruit.
to
by
up
5
—0.6
SW.
INDIA.
(BM),
Breyn
s.n.
Metz 2225
Hohenacker
Kerala:
mm
mm;
Cu-bi R.,
Wight 163
(E).
—
CEYLON. Fraser
Macrae
12
(BM),
CAL, K, P),
(E),
K.D.
Kandy :
?
Hb. Pallas
Walker 9
396 (P).
Macrae
28
33
(E, U),
Central
—
India,
Doumer
(BM);
Bourne
(BM, Fl, K),
little tapering
up
(2^ —)
usually
ovate
by
—8
in
rounded,
—6
white.
or
by
high,
elliptic,
to
sometimes
creamy
mm
mm
0.7—1
mostly
—j(—6)
to 7
to
up
usually
conferted flowers;
±
Torus
obtuse,
to
base,
at
rachis
inflorescences
sometimes
long,
plants,
young
of lateral inflorescences
Sepals
0.
acute
Ceylon,
Hb.
s.n.
E.
in Hb.
Thailand,
Hohenacker
4815
Bourdillon
(K);
129
Hb. Hermann
(P), 89 (U),
Prov., Sigiriya:
Brogniart (P);
Vaillants.n.
89
0.5 —0.8
fruit; stigma
mm,
Stamens
mm.
minute.
Hainan,
Indo-China,
(BM,
K.D.
—
Hb. Madras
E, K,
P,
L,
U),
Ramakan
396p.p. (K).
—
=
South Canara:
Mysore,
Wallich
2802B
Beddome
1086
Wight K.D. 396p.p. (K).-
934
(DD),
Hb.
Wight 469
=
Madras, Tirunelveli, Kuttalam:
(K).
s.n.
427
(P).
Travancore:
(L),
(BM), Pamplin s.n. (E),
54
a
in
cm
many-flowered;
filaments; anthers
Quilon: Wight 164 (E),
Colatinpolay:
113
m,
lamina
long petiole;
broad-spatulate,
to
mm
10
acute or
ones
mm
yellow,
3 —5
by
deciduous.
mostly
—800
fr. 18-V-1922.
m,
Borneo, the Philippines, and Celebes.
s.n.
Calicut:
(E, GL, K, P);
35
(1 —)3 ——cv>),
5
7
—
l'Enfant,
mm.
(C, P);
Royen
long
style
(BM); Mangalore:
Belanger s.n. (P);
Wallich1 2802A
Vahl
van
2225
mostly
Peninsular
in Hb.
(P);
high,
mm
—
apex,
fruit,
mm,
%
to
up
with
obovate
truncate,
or
i\
to
terminal
of
in
cm
mm
up
primary
or
by z\ —4
Sumatra, the Malay Peninsula,
(E, K);
branches
1
at
1 —6.
de
et
Branchlets of young plants sometimes
cm.
cm,
Type:
—
(1946) 671, f. 82,
1
massif de la Mère
Inflorescences
obliquely
4—7
0.4
to
8( —10) by 6(—8)
Distribution:
40
2—6
cymose,
to
up
Petals
with
0.7 —1
long,
cm
obovate,
or
by
6—20
primary
long,
in fruit
rounded,
up
1
cm
0,
to
Leaves with
mm.
I —2
chartaceous.
denticulate,
up
mm.
Ovaries
by
long, secondary branches
reduced
subsessile
high, dbh. up
m
acuminate, sometimes obtuse
cm
cm
enlarging
25
3 —4
245.
8-XII-1917.
Paratypes: Poilane 12256 (P) Annam,
—
obovate-lanceolate,
acute
obtuse,
Klong
(K holo) Sarawak,
(1931)
1
fl.
Kopah,
Suppl.
(P holo) Arrnam, Nhatrang,
fl. 20-V-1923.
15—1700 m,
0.7—i
Back. &
70;
Chang,
Ko
fr. 28-VII-1925; Poilane 3354 (BO, P) Annam, Nhatrang, Song Co, 700
5-—20 (
Gagnep.,
245;
(1963)
23
6849 (K holo, SING)
Kerr
Type:
—
(Jack) Gilg
sumatrana
Ouratea
margin
microphylla
Ouratea
—
I-1915.
Lectotype: Poilane 6604
mostly
Harmand
Type:
—
1876—77.
(1946) 673; Lars.,
1
327.
& Nur 2047
ovate-
Type:
—
1877.
Type: Robinson 6197 (K holo) Lankawi Is., P.
—
(1963)
megacarpa
Matang, fr.
Haniff
(1907)
5
Nat.
1821.
XII-1916.
Kew
Suppl.
1
Mai. Pen.
255.
5
87; Ann. Sc.
(P holo) India, fr.
fl.
Mayum,
Mt.
Java
fr.
&
Craib,
arcta
(1931)
1
(1903)
9
Hb. Brogniart
ex
Harmand in Hb. Pierre 7027 (P holo) Annam, Hue, fl.
144
1968
i,
(K),
König
Rottboell
1059
Worthington
(BM, C, L, P),
s.n.
s.n.
(C),
(E, GL),
33
(BM);
s.n.
Thwaites
(E, GL,
Nalanda:
Leschenault
C.P.
K,
2412
s.n.
P), Wight 165,
Worthington
(BM, K); Nawalapitiya: Worthington372 (BM); Hanguranketta: Simpson
(P),
(BM, BO,
1034
9208
166
(BM);
(BM).
—
Kanis:
A.
Southern
Rd.:
Prov.,
Udon
Circle,
Lakshnakara
600
Smitinand
(C, K), 603 (C),
Laos.
K,
Poilane
Savannakhet:
Schmid
CAMBODIA.
(p).
31994
Bejeaud
Mb
Godefroy
Thu Due:
(P);
Phung
Tri
CAL, P);
(P),
(P),
129
waterfall:
Huyen,
Nha
Robinson
Trang:
Nui Hon Heo Peninsula:
(K, P,
[
SING), 4801
8000
Eberhardt
Poilane
Eberhardt
2328 (P);
(P);
31273
Dent
(P);
du
Poilane
Hainan. HOW & Chun
Sumatra. W.
b.b.
S.
6069 (BO, L).
(BO, L).
Hooker
—
S.
Radja:
W.
and
3631 (BO);
Niru:
Ba
Tukang
Bumi:
Gusdorf
de
Voogd
wang:
L,
Muntok,
Kostermans
19131
Ton:
P.
Kerr
Curtis
Paull
s.n.
s.n.
S.
Corner S.F.
(SING).
(SING),
—
S.F.
221
Evrard
Van Dieu 40
Poilane
Ko R.:
(BO, P);
3354
C6 Inh Massif: Poilane
Da
Quang Nam,
Nuages:
Ka:
Bai
(P);
4611
Nang
Poilane
7842
Hue: Clemens
Eberhardt
(P);
2473
Sa
Lung:
Che Massif: Poilane
11280
& Finet
(P);
Khanh
(P);
2333, 2398
Col des
61
Ismael
?
(K),
7599
(P)-
1313
Yai Distr.:
Liang 62377 (E),
(K, L), Meijer 6989a
(L); Solok,
Elbert
s.n.
A.
1340
Arta:
Sugi:
Is., P.
Telok
& Anta
P.
(L),
and
H.B.
Teijsmann
s.n.
Klabat,
708
Hb.
s.n.
(L);
(K, L),
Steenis 3397
van
P. Bau:
(BO);
between
(BO);
Natar:
19
S. Liat: b.b.
34229
(BO,
Ulu
Kota
Menggala
Teijsmann
Teijsmann
(L);
(BM),
3033a
s.n.
Dorst
s.n.
Tubuan:
3961 (BO, L, U);
Sepatuhu:
Air
in
s.n.
(BO, L); Bajung Lentjir:
(Mangala): Teijsmann
Elbert
S.
Posthumus
Marsden
12043
60
Singkep,
Djcmadja,
S. Manau:
Kostermans
(BO, K, L, P, SING);
Kostermans
H.B. 4234
(BO); Bangka,
s.n.
(L);
1346 (A, BM, BO, K, L, SING);
(BO, K, L,
G.
Betong:
Lobok Besar:
SING).
Circle, Krabi,
Pattani
K,
Tambon Kao Panom: Kerr
Nur
Circle,
2074
221,
F.
R.:
Terai: Corner
3195,
(BM, CAL, SING);
s.n.
18776 (K); Trang,
(BM, BO, K, SING);
Kao Ri Chow:
SING); Jerai
6197 (K); Tg.
=
s.n.
Rahmat
Durian:
(BM, CAL), 1776a,
472
Mas
to
Hb. Hasskarl
F.R.:
Simpang
Forbes
G.
P.
(BO, L); Tulung Bujut: Engles-Julius
s.n.
.35144 (A,
road
Ogan: Teijsmann
between
(BO, K, L);
1319,
Curtis
in
s.n.
(BO); Tjaban
Ulu
Anambas
(BO);
(BO); Palembang: Teijsmann
s.n.
Arch.,
Riau
(L);
Selajar: Biinnemeijer 7427 (HO);
Jack
Pengubuan, Gedong
—
66
P.
37813 (A, BM, BO, K, SING);
Penang:
Curtis
Hoa:
(BM,
Tenajan: Soepadmo 167 (C, E, K, L); Upper
al.
(BO, L); Menggala
s.n.
Puket
Robinson
Bien
Zi Tinh: Pierre 323
p.p.
(FI);
Djambi, Bangko,
Lampongs:
14464 (K).
Dayang Bunting:
Selat Panchor:
—
(K); Kopah-Janjan: HanijJ&
Kedah.KohMaiF.R.:Kiah
Is.,
40
Mangol:
Thailand,
Pierre 323 p.p.
(C, K, P);
p.p.
7216 (P); ThuaThien,
33331
Poilane
(P),
P); Tay Ninh,
E,
(Fleury) (P),
(= Baria): Phung
Lecomte
Wang
Biinnemeijer 7380
SING).
448 (A, BO, K, L, P,
MALAY PENINSULA.
Soi Dao: Kerr
G.
bis
283
s.n.
(P);
7027
Soai:
(Fleury), 36944 (Chea
6604
s.n.
Chevalier
P); Kg.
K,
(E).
i
&
Lake,
W.
Ulu
(BO,
31731
12236 (P); Dong
Tjampo:
(BO, L);
Ranggal:
(A, BO, L);
& Anta
(K); Khlong
Baru:
Teijsmann
Menumbing: Teijsmann
s.n.
S.
4375
Amiroeddin
Brooks
Maras: Kostermans & Anta
SING);G.
Teijsmann
231
Le
Ca Na:
7109,
Thua Lun:
Zollinger: 3073 (BM,BO, L, P);
Telok Bt.
U);
G.
& Tso
Pursat:
(P);
7023
649
(P); Song
Pierre
Poilane
?
—
of Ranau
Teijsmann
(BO);
185
Terbanggi Besar:
(BO,
Benkulen:
N.
(BM, E,
131
(K, P), 29843 (BO, P); Quang Tri,
27794
1963 (P),
Teijsmann H.B. 3696 (BO, L, U);
H.B.
SING);
Poilane
in
Siak, Pakanbaru,
K,
(K, P); Talang Ampat: Ajoeb
Teijsmann
L,
S.
(BO,
Muara Enim:
Harmand
(BO, L); Padang Lawas,Purbasinumba: b.b. 6198 (BO); Sidjundjung:
(BO); Kg.
20433
Sumatra,
T1IP703 (BO, L);
Batu
Hb.
Taram,
Pajakumbuh,
7147
Schmidt
Hahn
Thorel
Poilane
Harmand
Chun
Taratuk Air Hitam:
S.F.
2888
(C, L); Nipple:
643 (BM, C, CAL,
Phuoc
(P),
291
Teijsmann s.n. (BO, L); Lingga Arch.,
P. Karas:
Henderson
l'Enfant:
Gaudichaud
(P, SING).
Riau, Upper
—
8221
Poilane
Poilane
Ma:
Shi:
Put K.D.
Schmidt
14112
Chevalier
(P),
Thuan,
(E, K, P), Liang 64840 (K),
Vreeden 233
van
Manggu: Biinnemeijer
Maras:
70110
Wen Fa
Sumatra,
Buurman
Indragiri, Taluk,
(BO, L);
to
(P),
Pierre
Chevalier 30211
Tuy,
4266 (K, P),
Eberhardt
10410
Phuoc
Mere et de
Clemens
2078
Sangka:
(L); Ubon,
32311
6849 (K, SING),
Treng:
(P), 6224 (BO, P), 6238, 6834 (P);
6221
Nui Bach
Valley:
Cu Bi
Tigre:
62704 (P); Dung Ka,
Silunkang:
Ba Na:
630 (K);
Tha Khum:
7119
(BO, SING);
s.n.
Annam, Ninh
—
(BM, K, P, U),
3377
(BM, K, P, U),
3417
Toang:
4896,
S.
Kamchay:
313, 533
Poilane
Ninh Hoa:
(P);
1490
at.
Put
R.F.D.
=
Kerr
Poilane
Thorel
(P),
s.n.
(E);
323 p.p.
Massif de la
(P, SING);
Srok
Thu Dau Mot:
(P),
212
=
(E, K, P).
Poilane
(P);
Clemens
Tourane) :
=
(BO, P),
Murutallawa
(BM, K);
Popokvil (Elephant): Geoffray
Mt.
Lefivre
Point A:
in Hb.
Ksan:
Kralanh F.R.:
31938 (P);
(P), Talmy
144
Pierre
&
Mt.
Basin,
Harmand
of Cheom
(P); Bokor,
Pierre 323 p.p.
908
17690 (K);
Sorensen
(L),
36188 (P); Prey Veng,
(P);
323 p.p.
Dinh: Pierre 323
p.p.
Mt.
Hoa,
Pierre
Van Dieu
168
Surin:
Circle,
Phenkhlai
Kerr
Knap:
Se-Moun
(P);
(K,P); Long An,
Pierre 323p.p.
Ubon
Chang, Klong Mayum:
22832
W.
Thorn,
Vietnam. Cochinchina: Harmand
Mt. DeonBa:
Gardner
57
(E).
p.p.
Chevalier
Chevalier
(P); Kep:
22972
Kao
Ko
12086
74, 283
io) (P); Kampot: Geoffray
214,
Hautane:
(K);
54
Reap, Angkor:
Siem
(P); Kg. Chhnang:
148
?
—
Area
Indo-Pacific
(K).
649
(P); Kg.
333
Circle,
R.F.D.
=
19
11713,
Thorel
(P);
s.n.
3633
Murton
P); Kang (= Khong?):
Anuwat
(C, E, K);
17927
Nun:
Chang
Thani:
Chantaburi
877 (K);
Kerr
Ko
(P).
212
Srisaket, Kanthalak, Nongtha:
(C, E, K, L); Klung:
667a (C);
Pierre
(E),
s.n.
(K), 8270A (C, K);
8270
Warin :
Dubuc
the
of
Madugoda: Simpson 8984 (BM).
;
Udon
THAILAND.
Kerr
Galle:
Simpson 8099
the Ochnaceae
of
A revision
Lakshnakara
722
Mat Sant C.F.■ 17934
S.F.
'■
37854 (DO,
P. Terkam: Corner
(SING),
Bt. Penara:
Satul: Lakshnakara
Kunstler
Yahya
S.F.
.
K, L,
s.n.
1320
(K).
331
Malaya,
(SING);Langkawi
SING);
(SING);
(CAL);
21430
—
Kuo
P.
P.
Batu
(A, BO,
Chupa,
Songsong:
Feringgi:
K, SING);
VOL.
v
L,
P.
(SING);
Scortechini
near
Ulu
Wai:
(K,
Bt.
C.F.
Sadanan:
F.R.:
Musang,
S.F.
33841
S.
Ketil:
Hashim
Henderson
Hume
Smith
S.F.
C.F.
. 906 (K);
F.M.S.
G.
22662
33897
Derry
Kemaman, Bt.
of S.
Raub, Gorge
S. Nilam:
C.F.
Nong
SING).
Ridley
—
Castlewood
Johore:
&
98237 (K), Santiago
G.
Lambak:
Corner
BM,
(A,
E, K, P);
JAVA.
P.
Sarawak,
Bartlett
(K, SING);
s.n.
Ilias Paie
L,
8316 (K,
(K, L), Purseglove
2nd
Haviland
& Hose
Pelagos :
Daud &
S.
.19941
(K, L, SING);
Tachun
SING), 2488 (A, BO,
Muput
F.R.
Nyalau
Bt.
S.
Lambir,
Haviland
& Hose
(K, SING);
Lawas:
Hotta
Road
12794
S.■4877
near
Tada
S..18026
(K, L);
(A);
P.B.
'.
S.1594 2
21298,
S.
Div.,
374L
Kapit
Mt.
Ulu
(K,
Distr., Bt.
L,
Bt.
(FI,
(L);
Riam
Hose
450
Ra., Long Kapa:
Ulu
(BM,
Dapoi:
L,
Rd.:
Au & Ashton
(K);
G.
4601 (K, L, SING); Bario,
—
Bt.
Luang
S. 1170
(near
S. Marario:
(K),
Kuala
Belalong:
Ashton A
1
(K).
—
K,
Bt.
S.
K,
Ulu
Miri
Distr.,
L) ;
BO, K,
Distr.:
Baram
Sibat
Tisam:
SING);
Ulu
Kalulong,
Matu?):
(A,
Anap,
22167 (K, L),
(BM,
Moulton
ak
Tinjar,
N.
slope:
S.F.
666}
S.20043 (K, L); 5thDiv.,
(L) ; Kg.
37058 (K, L);
—
F.R.:
S. Paku:
1920
S.
(K, L);
(K, L);
Ashton BRUN1185
K.F.
17818
& Kadim
Raya
Richards
SING);
507
Bok
Anderson
Suhaile
S.
Sinclair
S.
SING),
K,
Ashton
S.
.21977,
=
(K, L);
Lio
Sepadang:
G.
(BM,
(K,L, SING); Nanga
S..8681
(A, BO,
Bt. Bangar:
.7841 (BO, K, L); Temburong,
13548 (KYO);
L,
S..
.16788
Ashton S.
Hotta
(L),
(SING);
s.n.
(K); Kuching:
s.n.
P); Btg. Saribas,
Luang
Hose
S..
.22988
Sewabok
s.n.
N. Amau: Othman b. Haron
(K,
Mentagai, S.
1525, 1957
Brunei, S. Belait, Kg. Ingei:
(KYO, L); Tutung: Brunig
ak
K,
Koyan:
Brunig
Haviland &
Sibat ak
SING);
F.R.:
Niah:
Richards
Ridley
507
B.■ 1
15
.18938
Ulu
L,
BO. K,
36492 (A,
Asam:
S.
S..11909
Sibat
Mersing:
Similajau
SING);
Chai
E,
Burkill
(A, BM),
Smith
s.n.
2803A
39639 (BO,
12482 (K, L);
(FI,
Hombron
Wallich
(SING);
=
Telok
Mujong,
Dulit,
Brunig
K.F.N.
SING);
L).
0088
Anderson
Raya:
Bt.
Distr.:
Mering:
K,
Baru:
Lundu:
South: Kanis
Ulu
G.
Tinggi: Feilding
s.n.
S.F.
? P.
Haviland
Beccari P.B.■3138
(BM, K);
Anap,
SAR
16244 (K, L);
Gondol:
S.F.
SING),
(FI);
P.
Matang): Ridley
SING),
Hose Mts.,
Bintulu
4025
21305
(K, L);
37^1
(BM, L, SING).
Labu:
Bt.
(K);
C.F.
18496 (130,
(BO,
246 (K),
(K), Ridley
(BO,
40374
L,
S.
Anderson
2130
(En-)toyut:
Dulit
S. 23435
(SING),
(K,
Btg. Lupar, Marop:
Haviland
Beccari
Anderson
Api:
Nat. Park:
L); 3rd.
L,
879
Muas
Abang
13475
Camp:
Yeob
Chelliah
(SING),
s.n.
Walker
P.B. 3547
Serapi (=
Rd.:
Penrissen
SING); 4th Div.,
SING);
Suib
Brookee9939
Anderson
L);
L,
3724, 3734,
G.
12621,
507
=
Long Dapoi: Haji
SAR
ak
G.
35673 (K, L, SING);
Belaga:
Bakam: Fuchs
Luang S..23206 (K,
Pickles
K,
S.F.
Bintulu:
Brain
:
Datu: Beccari
F.R.:
SING);
coll.
(BM, SING);
s.n.
Koorders
SING),
Distr., Tg.
Chai
Ulu
Kanan: Ashton S..19356
(K> L, P);
22179
Bako
Div.,
K,
(BO, L,
(K);
K,
456 (SING),
Ridley
—
SING); Mersing F.R.:
36482 (A, BO, SING), Ngadiman
Ubin:
P.
(E,
9375
?
(P),
5874
S.F. .
24931
(E,
CAL,
(K), Cantley's
4984,5045 (K, L, SING); Lintang Path
(BM,
3179
S.F.
Santubong: Bujang
Paku:
Telok
(BM, L);
376
507
G.
Pierre
& Kiah
(K, SING); Mawai-Jemaluang Rd.:
s.n.
24964 (BO,
178
Kelantan,
(SING),
Ledang &
G.
S.F.
39551
(CAL,
1054
—
Rhinoceros
13101
Saik: Henderson
Holttum
s.n.
(K, SING).
3°470
Tahan,
(K, SING);
S.F.
25974,
Sinclair
(A); Kuching Distr.,
SING);
P. 4910,
(L, SING);
Brooke 9371
5231
Sinclair
Holttum S.F.
(BM),
s.n.
Liew
Lundu
Div.,
Haviland
(BM),
s.n.
Haviland
1st
Native coll.
Sampadi:
10307
Karta
Karimundjawa:
BORNEO.
Pulai:
Deny
Brang: Moysey
(SING); Banang
Sembrong:
Bt. Timah: Hullett
(SING);
s.n.
G.
47
4058
Corner S.F.
(BM, BO),
20
Ridley?
SING),
SING); Tampenis: Ridley
S.
coll.
S.
Soh C.F.
Tioman, Tg.
Kelsall
Singapore: Anderson
(SING);
s.n.
K,
(BM, BO,
2029
Gul:
Tg.
Pelandok:
—
Kunstler
(SING),
132
SING);Tengah: Ridley
S.F.
P.
(SING).
s.n.
&
(K);
71522
Sedili, Danau:
(K, SING);
Burkill
Hullett
(A, P),
S.
Lake
(A, SING);
9373
(K);
77930
S.F. 32991
(SING),
s.n.
S.F.
Holttum
Kochummen K.F..
Pahat:
(K);
372, 374
(A, SING),
P.
15663 (E, SING);
K.F.N.■
Lindong
C.F.■ 3715
(A); Pinerong: Cantley's
s.n.
Batu
Kuala
(SING);
3227
C.F.
Poore
Perting:
Kuantan: Mahmud
Bidin
3630 (SING); Rompin:
S.
(K, SING);
55'S
16897
2480 (K),
16949 (SING); Bentong,
Haniff S.F.
&
■
Bt.
Ridley 2458 (CAL, SING);
S.
Corner S.F.
Kajang:
Burkill
Tras:
Ulu
Trengganu,
—
C.F.
(BO, L);Bt.
s.n.
(SING).
249
Kuala
SING);
SING), Ridley ?
BO,
Brisu:
Ridley
L,
(K,
Hamid
(SING), Griffith
Bt. Panchor:
L, SING);
Ahmad C.F.
Res.:
11646 (BM,
1166
K,
Selangor,
—
HMB. 1034
(CAL, P, SING);
987
(BO, SING).
(SING);
Wild Hills
567
SING),
31652 (K, SING),
(BO,
250
(SING); Kepong:
7205
Alvins
(K);
295 p.p.
HMB.
3370
SING); Batang Padang
(BO, SING).
s.n.
Nur S.F.
Angsi:
Malacca:
—
& Shah
L,
coll.
Scortechini
Para:
K,
99b (BO,
Kings
BM, K, L, P,
(A,
Corner S.F.
Burkill & Shah
65520;
Larut:
Arang
K,
463 (A,
Pangkor:
Seimund
Buloh:
K.F.N.
Tiga:
K.D.
S.F.
P.
Bur kill
Lalang:
(SING); Sg.
Batu
=
MK.
;
Pahang:
1581B
(A, BO, SING),
P.
906 (SING).
Deny 384 (CAL, SING);
Kadim
&
G.
Sinclair
«p
(CAL, SING),
coll. 4545
Hamid
Aling:
/'
99
(SING);
11592
Telok
(SING);
(SING);
3125
Gopeng: King's
Pinang:
Negri Sembilan,
—
C.F.
Maingay
S.
Gates:
Wyatt
S.F.
Kinta,
Allen
(K);
s.n.
Scortechini
Wray
Reservoir:
(SING);
s.n.
4
.x
SING); Dindings,
(K); Kg.
(K); Klang
474
Tampin Res.:
Gua
L,
Ahmad C.F.
■ 5415
Lagong
Bt. Baru:
Henderson
8694 (BM,
2803B
12649 (SING).
G.
SING);
coll.
6598 (P),
(K, SING),
s.n.
Malaka:
Bujong
G.
Rd.:
C.F.
Hashim
(SING);
Hill:
For. Guard
t/
coll.
King's
SING),
.
Pahang:
T.
/.
4168 (E, SING);
Is., Pulau Rumbia: Kloss
SING);
Ja'amat
coll.
Wallich
(E),
s.n.
Maxwell
Lumpur,
Perak:
—
(K); Taiping
3552
567a (SING);
Sembilan
S.
P),
Sarajah: King's
Selangor: King's
Scortechini
x
1968
i,
K,
_.
37657 (A, BO,
(SING);
s.n.
(BO, CAL, SING), 562 (U),
537
(BM, K,
Curtis
C7
39306 (E, K, L, SING).
SING),
Nauen S.F.
(SING),
221
Bahang:
,.
.
,
=
Suloh
(K);
2 jog
S.F.
Curtis
Hill:
Penang
No.
XVI,
■
BLUMEA
58
Hotta 13100,
Sabah:
Mendaram:
Berakas
Agama
13105
474,
F.R.:
(KYO,
479
(A),
Wood
(BO, L),
595
Kuala
(S);
Penyu
SAN 50054
Gaya:
Distr., P.
Sittanggul
SAN
(BO, L), 27706 (K, L);
Labuk Rd.
Distr.,
BO, K), Creagh
Tahir
(A, BO),
Chinese
SAN
22606
SAN
7187
Arsat
F.R.:
L,
Distr. :
Tawau Res.,
Chin
31632 (K, L);
Silabukan
Kennedy Bay,
33409
SAN130441
Maidin
Rd. mile
Abu
L,
(K,
P.
(K, SING);
Ahmad
K).
S.
(BO, K);
2714
De Vriese
?
S.
1J
mating:
2092
b.b.
(BO, L);
ülu-u:
18893 (A,
P.
Bclajan, Tabang,
K, L,
(BM, BO,
Kostermans
Bt.
S.
Meijer
2504,
6606
SING),
Cettabre
(A, BO, L);
Melville:
Merrill
1520p.p.
San
Agr.
611
gi38o (L).
Zambales,
(K); Busuanga
Luzon,
—
(K); Cagayan:
=
P.N.H.
61—10
4613
(BO,
E,
E,
FI);
(A, L);
SING);
Puerto
?
(A);
Paracale: Ramos & Edano
Bical
Galera:
Sulit
Romblon I.: Escritor
Masbate I.: Merrill
F.B. 313
(BM, K);
B. Gov.
33541
Canicosa
5710
B.S..
Lab.
Elmer
2968
=
12740
(BM),
=
Manilla?:
Escritor
Creek:
(A,
B.S.
Alcasid
—
3611
=
P.N.H.
11751
8981,
Kasian:
Bt.
Kelindjau,
East
—
Lakes,
=
Borneo,
(BO, L); Berau,
Amdjah
(BO), Meijer
(BO);
197
(BO,
3577
Bentuk:
11
2910
Lianggagang:
9070
Endert
L);
Kostermans
G. Beratus: Kostermans
S.
S.
12629
Sulit
3785
P,
(A), 2278,
Claveria:
Ramos
Cuming
1520 p.p.
P.N.H.
U);
2279
Mt.
L, P,
Mindoro
Yagaw:
SING),
Iloilo:
Vidal
(A, L, SING).
P.N.H.
& Cordero
SING);
Madre,
Gutierrez
7658 (A);
Camarines
40, 45
=
264
=
Norte,
F.B.
1520 p.p.
EdaHo
(A);
Sulu
Cape
Mulanai: Alcasid F*
Cuming
Conklin
c
I.,
(A,
Palanan:
(A);
Canicosa
155
(A),
12349
Cuilon I.:
(A, K);
(A); Isabela, Sierra
Ramos &
—
Balabac
(P); Batangas: Cuming
4683
I.:
P.N.H.
j2g66
(BM, L), Quisumbing
Visayas,
=
Mendoza
B.S.
78184 (K, L);
P.N.H.
7456 (L).
Cettabre & al. F. B. 27890
CAL, E, K, L,
Edano
CAL,
Garson
L); Kapuas
(K).
923
(K), 20842 (BO, SING);
32829 (A, L);
BM, BO,
Merutai:
Abu Bakar
Rd.:
b.b. 11266
Kemul:
155 bis
2iig7
&
SING);
Pinayas: Orolfo
Mandai,
L),
Agelar F.B. .28883
=
3085 (BM, K, P); Panay I.,
Buenavista: Sulit
Bt.
Sur, Sipaco, Sagonoy:
9767 (A).
P.N.H.
&
20774
Camarines
S.
(A, BO, SING); Lipuun I.:
towards
61—222
(BM),
P.N.H..
1211
K,
14226 (A),
BO,
(A,
SAN
13306 (BO, K, L); Samarinda, Loa Djanan:
I.: Vidal
Cenabre
77991,
Baler:
(A);
S.
L);
P.N.H.
B.S. 41219
L); Rizal,
20721
Dimilngen
B.S.
N.P.:
B.S.
K,
G.
Chai
48754 (K);
K,
(BO,
Salimbatu:
Mahakam,
36066
(BO, L, SING); St. Barbe I.
28125
8677 (BO,
SAN
SAN
Penangsoo: Belajadia
Ulu
SING); Balikpapan,
SING); Paragua
18
W3
(L); Aurora,
Escritor
K, L,
Edaho
K,
b.b.
15589
(FI);
(A, BM, BO, CAL, L, P, SING),
P.N.H.
=
34'4
?
1459,
Chai
East:
Apas
Mt. Lucia,
SING);
Bala: b.b.
Kostermans
Norte, Bangui
(A,
155
L,
(BO); Kutai,
12796 (BO,
I.: Ramos
Ilocos
Alvarez F. B.
782gs
Quezon, Guinayangan:
Edano
Vidal
Gutierrez
P.N.H.
K, P,
Ulu
(BO);
(A, BO,
Mts.:
35893 (L);
Bandjarmasin: Motley
Kostermans
Mandu:
Pulgar:
Mt.
1565s (BM,
Santa Cruz:
Mariano:
61—
347
(L);
Puerto Princesa,
Fettix B.S.
Bur.
P.N.H.
30100
Victoria
SAN
19651 (K,
Maidin
F.R.: Aban Gibot
L,
SAN
SING);
SAN ;23m
16169 (L, SING);
F.R.
Kuala
L,
(K,
Segama:
(K,
5240
& Norditi
Nicholson
SING),
Ambullah
Agam
Pereira
SAN
Kalabakan
Otik
:
(BM, BO);
SAN
Lokan:
Dam
(A,
P), 1687
Kapur: Meijer
A2800
S.
Sugut
F.R.: Meijer
Wood SAN
40605 (L);
(BO, L); Bulungan,
2571
Kostermans
2
BO,
Buaya: Meijer
(K);
SING);
Singh
SAN
Beccari P.B.
Magne: Jaheri 614, 654
Palimasan:
K, L,
Uinas
F.R.:
South Borneo;
1047
Pmrippmns. Palawan! Merrill B.S.
*234
F.B.
SAN
P. Nunukan:
L);
SING); Sangkulirang, S.
6498 (BO,
Kuala
Sinanggul
Kantu:
—
(BO),
1023
BO,
Tarakan:
Jaheri 418 (BO);
L, P,
Melawi, Kalawaideras:
(P, SING).
239
Semenggaris: Amdjah 996,
Du
S.
2742, 2863 (BO, L);
. Pedjantan: Latiglassd
(K, L);
Lubuk
Valera
95'9
Sim Sim:
K, L,
Kaluinpang
C, K,
(L); Mempawa,
s.n.
S.
Ulu
S.
Kadir
Sakar:
P.
BM,
Buli
(K);
(A, BO,
SING),
L,
(K,
47633 (L);
Beradaya
mile 24:
(K, L);
<
Sinkadjang: Teijsmann HB 8462
(BO);
Hallier B..
.2631,
SAN'
(A, BM, BO,
Simpang Tiga: Meijer
West Borneo:
—
Talip
7289
Segaliud:
Bt.
9438 (A, K),
(A,
(K, L);
38818
2754^
Ranau
Labuk &
(Melegrito)
1149
1691 (A, BO, K);
(K, L);
SAN 21661
Chai
Clemens
SING);
Matunggong:
Kebun China
SANJ 37851
Mujin
Kinabutan Kechil: Aban Gibot
SING);
Sebatik,
Silain:
Bakar SAN;24982
Mail
(A, BO, K),
Maidin
(BO, K),
Bt.
Chai SAN 29735
SING),
Elmer 21070
: Brand SAN 21453
tt
SAN 25040
BO,
L,
F.R.:
(K, L, SING);
2710
(K,
Tawau Distr.:
(K, L);
1897
Distr.,
Yu Concession:
24574
F.R.,
(L); Elopura
(A),
L,
Res.,
1557
20050
(L);
SAN
Elleh
(A, SING), 968 (BO); Bilit,
879
Datu
SAN'38883
•
(K, SING), Enggoh
10410
SAN 42924
Burot Ho
Lahad
Lok
Brand SAN
(K, L);
Puteh:
Batu
SING);
Elleh
31780 (BO), 30858
(K, SING), Belajadia
S.
Clemens
Ramos
SAN
P, SING), Meijer
Sapi: Sayuh
Batu
Enchai
Evangelista
(SING),
818
SING),
4326 (L, SING);
(K,
5919
(K, L);
Distr.:
(A), 836 (CAL, SING), 867
809
al. 1013
Sandakan
BO, K);
Sandakan
(K, L);
684 (A), Sayuh
al.
&
3184 (A,
(A), Punching
661
(K, L);
34^77
j
Sam
Mail
BO, K, L,
(A,
(K); Kabili-Sepilok F.R.:
1460 (K);
(A,
Leila
21198 (L, SING),
Kinabatangan
(K,
Keith
Som SAN
34605 (K, L),
Wood
675 (SING),
(K);
41 059
Monarca
Clemens
Apostol
Brand SAN 30904
(K, L),
J 36223
Bakar SAN
&
Chew
F.R.:
Distr., Kedayan
Batu:
Bay, Tg.
(BM, K),
Cemetery:
Kudat
A.
(BO); Dallas-Tenompok ridge:
E. Shoulder:
Jesselton
33561a (K, L,
Smith
Wyatt
s.rt.
Madius
SAN 43013
West Coast Res.,
SAN
Penibukan:
32670 (BO);
30323
Kinabalu,
Apostol 7680
mile 82: Abu
s.n.
Clemens
Clemens
Mt.
(K, L);
173
Marudu
SING);
L,
4687 (K,
Dallas:
Treacher
Hill:
Masirom Rundi
South:
'41444 (L) ;
Wood &
I.:
Beaufort
Distr.,
(K), Mujin
59
Labuan
Res.,
SAN
1443
Distr., Tabilong:
Lobang:
Distr.,
F.R.:
Labuan
(K, SING);
.
4045
Ranau
Darnton
Resthouse:
near
above
L), 40849 (A, K, L);
K,
(A, BO,
Belud
Interior
Beaufort
Ra. F.R.
Ampuria
Haviland
Area
Indo-Pacific
(K, L);
Crocker
F.R.:
(K, L),
4i392
the
of
(A, BO, SING);
SAN'40809
15096 (SING);
Kota
(K, L);
lit. Nunkok
Kinabalu,
Mt.
SAN
Ampuria
Ochnaceae
2120
(K, SING); Rayoh
10507
40134
1865 (A),
Batayan: Ampuria
Wood SAN
(L); Pangi:
the
of
revision
1210,
Distr., Bt.
(K, L); Melap: Angiatt
A
,
Castillo
:
Kanis
.
A.
30306
(A,BM;
P.N.H.
18679
B.S. 31545 (A);
Guimaras
I.:
Gammill
Archipelago, Tawitawi:
BLUMEA
60
Ramos &
F.B..
Edaüo
B.S.
Hutchinson
F. B.
b.b.
Vernacular
Madras?:
Laos:
ton
prote.
so
aluan
ang
quan.
—
( a)may,
Sabah:
biobi,
burgang
Palawan: anduyong,
(Tagalog);
kĕlutak
are
branches
tribes
ondogong
mampa
Ecology:
used against
from
Repoited
peat
swamp
level land
but
to
sandy,
it is
its
remote
also
to.
Brunei:
antimagas
djambak, sebalusi
bunga kĕlimbing besi,
mĕramong,
murmagong,
names
were
a
—
with ubah
(Iban), pinis
(Murut), kolambang
tampalanuk
(Tidong),
melindingan
bakan
(Dayak);
(Malay).
—
East
Philippines,
(Zambales), simahima (Bicol), sasahit
morosisio,
as
names
chĕnaga lampong
gimbaan
alas,
—
chmaul,
(Annamite),
quân
compound
wulisi
poles
or
—
Celebes: lebani,
(Tobela).
mapute
for the construction of
planks
Sarawak. Roots and
and
the Malay Peninsula,
stomachic and anti-emetic tonic. Young
are
chewed by jungle
or
less
to
but
to
1200
m,
up
strong, apt
to
1500
m
lowland and submontane,
part of the
a
on
cliffs
usually acid; limestone
is understandable.
separated
make
river banks and
is
to
split
Mt.
on
in
drying.
Kinabalu;
mixed Dipterocarp
open
scrub vegetation;
near
the
Soils
sea.
are
reported occasionally,
soils in the localities concerned have been leached
two
characters
of the specific
western
up
out.
under rather different ecological conditions, the variation
grows
characters
more
impossible
red-brown, hard and
near
clayish,
or
the distinctive
parts
In the
konkea
several compound
West Borneo:
used
being
or
probable that the
very
morphological
it is
(Singhalese),
Bangka: mentungging,
(Panay), minsaray (Mindoro).
vegetation:
standable that in certain floras
area
madjang;
(Kenyah);
Luzon: salactoc
level
sea
slopes,
steep
loamy,
geographically
although
trung
forests, high kerangas forests, ridge forests, and
Remarks: As this species
in
roi tia,
pling; Malaya:
(Kinabatangan),
(Malay);
(Rangkas),
as
(BO),
Chantaburi: kapi
tooth-aches in Cambodia. The leaves
primary and secondary
rocky,
aam
(Kadazan),
caranan
Properties of wood: dull red
011
mal) kaera
pu;
kompuat chwuk,
cay
bĕsi;
tampang
decocted in Kerala for
are
tong
161
Matalena:
vilavu, chokatti, puachetti;
anai
wal (or:
lakodjong,
mat,
(BO);
Malaya.
in
forests,
Kerala:
gioi,
2413
(BO).
1172
gass,
IIII34 (BO, L),
2386,
(= Cratoxylon), jambu and kelat (= Eugenia).
Vietnam, Sumatra,
bitter and
are
b.b.
batu, mulak, tengkedjing kering
bibingo, huisac;
The wood is reported
Uses:
cay
(Iban),
tulangkara
(Dayak),
(Bugi),
houses in Ceylon,
in
bep,
gĕronggang
Visayas: bulocauan,
parasinga tjila
Rona:
Ubon: hang kwa
nao;
kaju
majang
pcsoon,
malatangor,
amir
Borneo:
Kjellberg
recorded occasionally;
were
quintalai,
(Rungus Dusun),
bitanag,
La
kĕtam batu, mĕnarah, mĕmbatu, mĕndapor,
mura, mata
(= Eugenia)
ubar
(K); Cotabato, Bango: Whitford &
Usu: Cel.
IVI194;
bo kaera
dua
cay
sepah,
Borneo, Sarawak: kĕladang,
leaves
Lasao:
lĕbah, pokok luas, pĕnarah, pĕngling,
(Malay);
Miranda
22684 (A),
.
Cambodia: domchung chhkê, kompes,
recorded occasionally with
or
Cel.
SING);
Malay Peninsula, Thailand:
—
& Acutta F.B.
Sumatra, North: kaju ndolak (Batak); West: kalek
—
chinta mula, lidah
pokok
K, L,
Ceylon:
—
South: kaju
(Menankabau);
mesulung putih.
Beliti
Villamil F.B. 22060
13584 (BO, L),
Thailand, Udon: chang
(Mo'i).
am
h.h.
Celebes, Kendari,
Vietnam:
—
Zamboanga:
India, Mysore: punde gida;
names:
—
1968
i,
«
II/438 (A, BO,
gurunda kirpata.
(Tamil?).
go
Cel.
SE.
(BO, L);
24099
Mindanao,
12461 (BO, CAL, L),
Malili:
Celebes,
Kawata:
(BO, L);
—
No.
XVI,
(BO).
1021
CELEBES. Central
190
(A, C).
44192
Tarroso F.B.
(BO),
22781
VOL.
key
'races'
or
three
Apart from these variations,
be recognized.
different
hardly be used
can
to
can
species
at
a
It is
have been
specific level.
the 'races', because the
same
some
therefore under-
recognized,
Over
differences
a
large
occur
in
area.
area,
a
distinction could be made between
a
race
‘zeylanica’
'
and
a
race
malabarica',
the former with slightly longer sepals, petals, and anthers, and
hardly branched inflorescences,
the latter with relatively small panicles.
In the basin of the Mc-Kong there is
a
race
‘thorelii’ with relatively
large leaves and
A.
profuse
flowering
Hainan, the
the
has smaller
flower parts
‘arcta’ is
race
of
A revision
Ochnaceae
the
unbranched, lateral
±
at
‘striata’
race
and smaller
Kanis:
like
and
the
of
Indo-Pacific
inflorescences.
those in Ceylon.
61
In Annam and
In the Chantaburi Circle
found with long flower parts and terminal,
in
possibly
terminal inflorescences
leaves,
narrow
more
Area
of Thailand
unbranched inflorescences.
The diffeientiation in Malesia is less clear. In Sumatra and the Malay Peninsula plants
are
In
greatest variation is found in Borneo,
relatively tmiform. The
exposed
more
habitats,
on
cliffs, and
small leaves. On limestone in the
small
leaves and
found elsewhere
Langkawi Islands,
reduced inflorescences.
very
the
along
The holotype of Meesia
coasts
serrata
having
as
considered the
Gomphia
all
sumatrana
back
go
to
stamens.
5
correct
Jack
(J.
Kurz
Gaertn. has been found in the collections of the
The species is doubtless
Jack's
but without
species,
but from the
original specimen,
with Ochna integerrima
known
to
(Lour.)
Nat.
Bot.
Syst.
Engl.,
ed.
Nova
2
Acta
in E.
&
P., Nat. Pfl.
in E.
&
P., Nat. Pfl. Fam. ed.
morphology
16
Trees, shrubs,
Inflorescences
not
(1966)
5.
short
Staminodes
with
0—eo.
2—00
Distribution: The
neotropics;
Remarks:
tribe
2
tribe
The African
(1895,
I consider its
natural, because
—
Stamens
ovules
As. Soc.
1854, 465)
Beng. 62,
synonymous
succeeded in tracing Griffith's
of specimens
per
15
other
conspecific
is
from Burma
&
Flowers
free
—
are
not
genus
(1903)
316;
Phyto-
distichous,
connate
curved,
functionally
or
at
base.
Fruit
a
decurrent
the lamina ±
slightly
or
Carpels
berry
or
parallel.
polygamous.
2—5,
fused
capsule;
torus
as
Banks
was even
representing
to
to
Gaertn.
ex
considered by
was
on
and is
confined
are
treated
various
in the
Africa
a
as
a
as
related
Central
a
the
to
separate
Ochneae
distinct family by Van Tieghem
and anatomical
grounds.
tribe within the Sauvagesoideae
separate
leaves and carpels with
a
5 —10
good resemblance to those
genera
The tribe Lophireae is distinct from other Sauvagesoideae
unequal enlargement of
genera
and
included in
was
him
morphological
that of certain Malesian
seeds. Like in several neotropical
mainly represented
West
(Euthemideae) respectively.
the shape and venation of tire leaves show
but
‘ser.
Albuminosae’
H., Gen. Syph.
straight
pantropical
is
Lophira
which
of its non-distichous
considerable,
—
Albuminosoideae Decker,
bisexual
or
21.
Sauvagesia L.
genus:
numerous,
area
(1901) 169—194
and the pollen is similar
138; D.-T.
Ochnaceae
—
'
(1874)
2
Type
5,
Sauvagesiae’.
37,
alternate,
tribes
Malesian
1925),
position
‘subordo
Akad.
carpel; style apical.
Sauvagesieae
monogeneric
in Morot, J. Bot.
a
4,
(J.
likely that this species
(1925) 63.
21
2,
nerves
because ofits exalbuminous seeds. It
by
is
Seeds albuminous.
and the West
by Gilg
sub
64,
paniculate.
or
distinctly enlarged.
(Lophireae)
epithet
SAUVAGESOIDEAE
(1895)
6
illeg.
nom.
petiole;
racemose
ovary,
Fam. 3,
46,
2.
undershrubs. Leaves
or
the usually
i
not
more
its
these records
Probably
and King
other records
Leop.-Carol.
Gilg
into
it is
No
(1836)
Gilg
Petals
times.
many
have
arguments. I
Rijks-
wrongly described
Gomphia and
true
Griff". (Not. Pi. As.
description
Merr.
a
were
me.
albuminosae’
into
are
genus.
II, 1871, 49)
40,
crocea
SUBFAM.
Lindl.,
species of this
has been listed for Burma
As. Soc. Beng.
II, 1893, 233), who considered Ochna
with
forms
Similar, but less pronounced
of the Malay Peninsula.
for the type
one
have
found with
is
‘microphylla’
a race
herbarium in Leiden. It could be demonstrated that the flowers
originally
in Sarawak.
especially
soils, specimens often
kerangas
on poor
the
outer 2
ovules.
in the
more
Besides,
Sauvagesieae
of that tribe.
by the
or
3
1 -seeded
sepals,
of the Sauvagesieae, there
are
fruit, winged
and exalbuminous
many
stamens.
BLUMEA
62
VOL.
Tribe
Planch, in Hook., Lond. J. Bot.
Gen. Pi.
37,
(1874)
2
(!9°3)
16
(1862)
1
21;
Gilg
3 j6;
(1966) 46.
—
319;
Gilg
in E.
in E.
&
Type
Shrublets. Lateral
2
cell. Fruit
per
Distribution:
I
7
Euthemis
genus:
of leaves
opening by
5,
Jack, Malay
Misc.
Miq., Fl. Ind.
221;
Handl. Fl.
in E.
in E.
&
Adansonia
Shrubs
P., Nat. Pfl. Fam.
I
or
glabrous,
(1961)
59.
sidewards, straightly ascending
terminal,
many-flowered,
polygamous,
Staminodes
pinkish.
ovules
5-celled;
Ovary
with
5
Seeds
pyrenes.
Distribution:
Two
1
152;
nerves
2
species
400;
(1868)
7
(1893)
(1901)
(1952)
234;
166;
Gilg
Vidal,
224;
caducous.
from
at an
bracts
c.
8o°.
caducous.
anthers
pendulous,
axile;
Cambodia,
Sumatra, the
stigma
curving
Inflorescences
<3
Flowers
red in fruit. Petals
free;
5,
angle of
small,
turning purplish
coriaceous,
Leaves
the midrib
5,
white
subsessile,
minute.
or
or
rostrate.
Fruit
berry
a
cell.
per
in
15
Arn. Arb. 33
free,
nerves
racemes,
60,
(1873) 360, 369, f. 383;
4
Soc. Beng. 62, II
parallel,
numerous,
5,
cell,
per
2
J.
Hook.,
in
Ann.
Walp.,
319;
Hist. Pi.
Jack
(1846)
5
leucocarpa Jack.
E.
filamentous. Stamens
or
5-celled; ovules
303;
Bartell., Malpighia
Stipules
the marginal
to
(1862)
1
86; Merr.,
species:
(1824)
2
Walp., Repert.
1143;
King, J. As.
branched.
Sepals
o(—-5),
Staminodes o(—5).
nerve.
pyrenes.
Roxb., Fl. Ind.
17$;
compound
actinomorphic.
H., Gen. Syph.
&
Ovary 5-carpelled,
H., Gen. Pi.
(1925)
2, 21
denticulate,
margins
the marginal
pore.
(1840)
(1895)
6
3,
sparsely
H.,
(1925) 64; Decker, Phytomorphology
(1875) 526; Baill.,
1
Lectotype
—
shrublets,
&
(1890)
1
P., Nat. Pfl. Fam. ed.
&
B.
Br. Ind.
Ned. Ind.
&
B.
EUTHEMIS
Gen. Pi.
(1859) 674;
1, 2
Hook./, Fl.
Benn. in
BoerL,
Gilg
Bat.
D.-T.
152;
179;
Leop.-Carol. Akad.
Malesia.
15; Wall, in
(1821)
5
1,
(1831) 69; Endl.,
2
(1849)
1
Acta
Jack.
usually i-seeded
in West
genus
Nova
(1895)
6
3,
2, 21
apical
an
Engl.,
543;
reaching
4.
Bot. Misc.
647; Walp., Ann.
593,
(1868)
P., Nat. Pfl. Fam.
&
berry with
a
(1846)
5
Ann.
1968
I,
EUTHEMIDEAE
2.
P., Nat. Pfl. Fam. ed.
nerves
anthers
free;
Stamens
Walp.,
No.
XVI,
SW.
Malay Peninsula,
and
Borneo.
Ecology: Confined to
on
low
ridges
everwet
tropical
areas
forests, and
in peat-swamp
in
below
open
1250
soils. Dispersal probably by birds because of conspicuous
(Ridl., Disp. Pi.,
1930,
Inflorescence
distinctly
1.
I.
2
branches
4—15
303;
Hook.,
a
very
cm
Jack
Ic. Pi.
(1859) 675;
Benn. in
Ridl.,
kerangas
well
TO
THE
developed
slender,
often
white, rose-pink,
Hook.,
II,
ibid.
4
8—40
cm
Bot. Misc.
1
t.
711;
(i860)
Br. Ind.
Trans. Linn. Soc. Bot.
3,
1
9
nearly
all branches
reduced with
Misc.
2
Walp.,
leucocarpa
conferted flowers.
2.
1, 5
(1821)
16; Wall, in
(1831) 69; Walp., Repert.
208,
long, margin
E.
denticulate
Jack, Malay
(1845)
Suppl.
Hook./, Fl.
red berries
or
SPECIES
with scattered flowers. Leaves
cernuousraceme,
long, margin faintly
in
forests,
mostly sandy
1.
Euthemis leucocarpa
(1824)
in
panicle,
in
on poor,
denticulate
Inflorescence
Leaves
a
altitude,
410).
KEY
1.
m
ridge forests,
Ann.
533;
1
(1849)
Scheflfer,
(1875) 526; King, J.
l79>
Nat.
As.
1
Soc.
Roxb., Fl. Ind.
N.I.
Beng. 62,
15
minor
2
(1842) 528; Planch,
Miq., Fl. Ind.
Tijd.
(1893) 285; Bartell., Malpighia
E.
32
II
Bat.
1,
(1873)
411;
(1893)
234;
(1901) 167; Hall./,
Kanis: A
A.
7. Distributional
Fig.
cited
Ridl., Fl. Mai.
Pen.
Prod. Mai. Pen.
(1952)
of Euthemis.
species
sphalm.,
92,
the
of
Outlined
Area
Indo-Pacific
63
from
only slightly generalised
areas are
(1922) 368.
1
H.B.
Teijsmann
Shrub
up
6
to
8
Flowers
2
Anthers
3 —5
1^—3
sectors
of
by
a
CAMBODIA.
c.
1
c.
SW.
Kg.
1 —2
by
in
Som:
Smitinand
P.
388;
Diet. Econ.
Index Kew.
ex
Wallich
Neotype:
—
33
Suppl.
Bat.
Suppl.
apart.
up
1
ovoid
cm
at
0
to
to
8 —10
ovate
(1868)
544;
Ridl.,
32;
533.
Type:
—
by
to
via red
c.
2
mm, acute
oblong
margin
to
mm,
by
2 —4
and
8—20
cm
lanceolate,
elliptic, unequal,
4—10
linear
distinctly
panicles,
2 —4
bottle-shaped,
,
by
lamina
erect
spathulate,
fleshy,
208,
4 —6
base,
Inflorescences
obovate
Ovary
to
(i860)
1
ovate,
base; bracts
Sepals obliquely
pairs.
7
(1917)
fl.
Stipules
tapering
apex,
at
2
34,
(K holo, P), Borneo, fr. 1853.
s.ti.
green.
mm
Ann.
Walp.,
z\ —5
by
turning white.
c.
1
4 —7
mm.
mm,
Seeds like
mm.
Cambodia,
Sumatra,
at
obliquely
yellow.
2
163;
long, winged petiole;
cm
long, articulate
globular,
4
stout,
2 —5
acute
cm,
mm,
Fruit
sphere,
N.
Type: Lobh
with
ciliate. Petals
mm.
(1862)
23
holo, U), Banka, Plangas,
L
(1921)
86
Burk.,
Merr., J. Arn. Arb.
249;
Hook. /
Type: Jack f, Singapore.
Linn. Soc.
nerves
mm
(1940)
‘E. jackiana
Soc.
311;
168; Hall./, Beih. Bot. Centralbl.
—
<3, erect, often
Distribution:
Sumatra.
2 —10
4—10
mm,
—
Leaves
denticulate,
long; pedicels
Kew Bull.
—
As.
(1926)
60
1822.
high. Branchlets
m
by
—40
irregularly
acute.
(BO,
3375
60.
—
Jahrb.
Bot.
latifolia Miq., Fl. Ind.
var.
acuminate, ciliate.
oblong,
nudum.
fr.
(1901)
15
leucocarpa Jack
E.
&
Hook, f., Trans.
Fl. Mai. Pen.
style
nomen
(1961)
1
J. Str. Br. R.
Merr.,
30;
(1922) 368; Diels,
fl.
(K), Singapore,
E. robusta
(1917)
2
34,
1
Adansonia
Bartell., Malpighia
by
Ochnaceae
(1935) 987; Airy Shaw,
1
Vidal,
224;
(1947)
2516
to
the
localities.
Beih. Bot. Centralbl.
10
of the
map
of
revision
Sumatra,
6456
Musala:
R.
F.
the
D.
—
Batten
Pooll
s.n.
Malay Peninsula,
24420
(K),
(SING).
Schmid
—
W.
s.ti.
and Borneo.
(P).
Sumatra, Indragiri Uplands,
S.
BLUMEA
VOL.
No.
XVI,
Bankar:
P.
Buwalda
6373 (K, L).
de Haan
Rangsang:
Henderson
Djemadja:
H.B.
Teijsmann
S.F.
U);Belinju: Teijsmann
S.
Selan:
Berkhout
Fl,
MALAY PENINSULA.
HUI:
Maingay
2207
K.D.
=
coll.
Tampin: Ridley
SING);
Sinclair
Nur
s.n.
s.n.
Murai:
S.
Clemens
coll. E
Browne
(K),
22333
(E),
212
P. Bruit:
S.
Anderson
Marudi:
(K, L);
(K);
Baram:
(K);
G.
K, L);
B
tRamos
(L);
(BO);
Teijsmann
H.B.
79lS
Amdjah
—
S. Tiram:
(BO),
G.
Ecology :
293
coll.
G.
K,
Kelam:
G.
s.n.
L),
Borneo, Sampit:
Hub.
S.
Winkler
Sebakis:
12749,
P. Nunukan:
SAN
(BM, K).
297
Hallier
Kenepai:
NE.
B.
\2363
Hallier
Buwalda
3266
W.
(BO, L);
St.
Bario:
Jaheri
237a
Anderson
9267
(BO, L);
E.
Rahman
—
I.
(BO, L);
Bt.
K,
Binai:
79
F.R.:
(A),
Wood
Paloh:
(P); Liang
Pedjantan?):
L),
G.
Rutten
(BO,
Sabah:
Singkadjang:
P.
(=
(BO,
1870
20108
1026
G. Silam:
s.n.
Borneo,
—
S.
S.
Wood 3881
&
(BO);
Barbe
17260
Synge
Res., Beluran, Sapi
7641, 7662 (BO), 7671
L).
S..1157
towards
Au S.
Dulit:
Bt.
Ashton
Borneo:
1239
(BM, L),
Othman b. Har on
Hills:
Tawau Res.,
Manan: Polak
Native
(FI),
(L); 3rd Div., Binatarig,
(K);
Sandakan
(BM, BO, K,
Kostermans
107
(BO, L), B2467 (BO), Langlassd
B1461
(L),
20
10789
Mujong:
Baram,
43416 (K);
—
(BO, L);
B
N.P.: Brunig
Lambir
23432
1693 (SING),
Chai S..17131
10639 (BM, L) ;
8369
Ulu
(BM, CAL,
Bridge: Jacobs 3687 (K, L).
(A, K);
3401
Kanis
Bako
Po: Brooke
Mts.,
6440
(F).
Beccari P.B.
SING), 9827 (L),
S.
Burkill
s.n.
8714 (L),
1316 (BO), Orolfo 693 (A, SING),
F.R.: Pereira
Leila
(BO,
Kostermans
(BO);
1038
Melegrito
Native
Brooke
L);
33376
Singapore:
(K); Bajau:
p.p.
Langlassd 63
F.R.: Smythies,
Telamba
2316
s.n.
Burkill
Kuswata
S.F.
—
(A, BO, L), 1634 (A),
Ulu
K);
E,
Badas
Brunei,
(K, L); Tutong,
(Tawau?): Creagh
Central
Palimasan:
—
(BM,
173
Mandi:
Anderson
CAL,
(K, P),
Goodenough s.n. (SING);
(BM, SING),
Hose
Suib
SING),
(SING).
s.n.
Wallich
K,
793
Panti:
G.
BO, K,
Timah:
486 (BM); Miri,
=
K.D.
33369 (SING),
(BM, SING),
F.R.:
(BM,
Ridley 1367
(A,
S.F.
s
Kostermans
Djempanga:
21
(BO, U);
12771A (BO, L), 12789A (L), 12906 (BO, L);
Samarinda,
P. Tarakan:
Meijer 2487
Kostermans
$7/1,
9001
(BO, K, L);
(BO, K, L).
From
Vernacular
iur iur
—
Saribas
8837 (BM, L);
Hose
Weston:
Sandakan:
Borneo, Djihi:
Meijer
2313
pĕlanduk
(K, SING).
21192, 21199
Mondi
s.n.
1318
King
Negri Sembilan,
(FI); Changi: Ridley
Bt.
West.
S.F.
Malaka:
SING); Jurong Rd.:
486 (BO,
=
—
Bell &
(A, FI, K, L), Ridley 601,
p.p.
Bt.
(BM, L,
9701
& Hose
Tinjar, Long Dapoi:
209
coast
Mandor:
(L),
Betong,
Brooke
3180 (K),
Res.,
K,
s.n.
s.n.
Burkill
Ophir: Griffith
(FI), 1378 (FI, K),
Hullett
(K).
Jerai:
1768 (SING),
=
Bujong
10907
Goodcnough
Native coll. 661
Bartlett
Rd.:
G.
Pekan:
G.
Corner
(BO);
(BM);
1289
486 (K),
Haviland
Div.,
(A, K);
(BO, L); Bulungan,
717
Belajan,
East
Teijsmann
Penang,
—
Curtis
Ra.
S.F.■ 32215
(BM,
s.n.
s.n.
Beccari P.B.
Haviland & Hose
B2698 (BO, L);
S.
coll.
(BO, SING),
Beccari?
(K, L).
308
(SING),
Rifle
297
Kiri:
14192
3118 (FI, K),
Hose
Interior
(P).
232
(L).
&
8461 (BO, L);
Hallier
Langlassd
2660
Moulton
Bianchi 33
L,
(BO, L);
(BM, SING),
3220
10636 (BM, SING);
Purseglove 4898 (K, SING); Tg.
(L), 1316 (A, BM, L, P);
1144
Gagang:
2nd
Niah:
23463 (K, L);
SAN 13470
Asam:
Corner
1941
Woodlands:
(K); Kuching:
K.D.
=
Runto:
(SING),
273
Haviland
s.n.
(L); Kelapaan:
Hills: Fuchs
(K, P);
SAN
(BO,
s.n.
G.
Kedah,
—
(SING);
s.n.
Holttum
Tanah
(SING), Ridley
486 (K),
=
(L);
17
9014
Sa
s.n.
Pahang,
—
Pelepah
Kang: Ridley
Beccari P.B.
Telok
Richards
Murud:
s.n.
Meijer
S.
Haviland
Haviland
Lumut
(P).
(SING),
S.
(BM, L), 8340 (BM), 8331
14386 (K, L); 4th Div.,
Lobb
Chu
1941
(A), Ridley
s.n.
(SING), Ridley
s.n.
(A, BO, E, K),
18013 (K);
Kuala Serai t: Kanis
(BM); Djebus:
Pandan:
Tg.
(K), Ridley
s.n.
(L),
Bubong: King's
7144
(SING);
772
Serapi (Mt. Matang):
8017
9664 (K, L),
Chai S..
Div.:
ist
3081
Brooke
(FI, K),
G.
(A, P);
Billiton,
11893 (K).
267
Curtis
Belumut:
(L);
4819
(BM), Ridley
273
Goodenough
Sarawak,
Borneo.
1929,2170
(K),
Corporal
G.
(K);
Panjang: Ridley 14136 (BM, SING);
SING);
Ulu
Ridley
Holttum
(SING),
Chua
(SING);
(BM); Kranji: Goodenough
Pasir
Bruas:
(K), Maingay 2207B
s.n.
11032
coll.
(P), Cantley's
s.n.
& Burkill
Selangor,
Stone et al.
Buloh:
S.
106
(BO),
Arch., P.
H.B.
• 3375
(BO), Teijsmann
1793
Lobh
Bireh
Hill):
Govt.
(K);
s.n.
Malacca:
—
ILobb
(E),
4739
(BO, K, SING);
Anderson
Corner
(K),
3193
(L),
424
Samsuri Sa
(K),
(=
Kerr
(SING),
Ngadiman S.F. 36635 (A, BO, K, SING). —Johore,
Penggaram: Ridley
G.
H.M.B.
(K). —
?
(SING).
Gordon 61
s.n.
Hill
Scortechini
1870
(BM, SING),
830
(SING);
Biinnemeijer
(BM, FI, K, P), Ridley 1768 (SING);
297 p.p.
Perak:
—
s.n.
Pontian:
IHullett
3967 (K, SING),
Scortechini
(K),
7159
G. Bui:
Biinnemeijer 2468 (BO);
coast:
Evans &
(SING),
s.n.
1768 (K, SING); Penang
(BM, BO, SING).
tHorsfield
Teijsmann
Sumatra,
(BO);
Ridley
6602
Anambas
(BO, L);
(BO, L); Plangas:
Thailand, Surat Circle, Tako-Langsuan:
Boden Kloss
&
Curtis
Southwest
Biinnemeijer 6600,
s.n.
13
P. Batam:
(SING);
Banka:
S.
60
Teijsmann
de Haan
Bengkalis:
s.n.
Daik:
G.
Lingga,
(BO),
—
Berkhout
(BM), Ridley
L).
Haniff s.n. (K), Flippance
Robinson
Petar:
P.
(BO, L);
397
s.n.
P.
(BO, L);
s.n.
Biinnemeijer 6874
(BO, K, L, SING);
(BO);
1039
(BO,
iiog3
Hullett
20446 (BO, K, SING).
s.n.
Polak
Rengat:
Karimun:
Teijsmann
(BO, L, U);
3211
P,
H.B.
P.
Tanda:
G.
(K, SING);
s.n.
Riau, B.
—
(BO);
P. Bintan:
(K, SING);
12577
Hullett
60
1968
i,
64
(Banka);
level up
sea
names:
to
iooo m, on poor
Malaya: pĕlawan
balong
(W. Borneo).
(Biliton).
bĕrok.
soils, preferable
Sumatra:
Borneo:
tambu
in moist,
shady places.
bĕlusung putih, kaju padang,
(Sarawak);
ranggas
hutan
mata
(Sabah);
A.
Use: Medical
Remarks:
probably
Jack's plants
original
lost
Edinburgh
in
leucocarpa
The
‘E.
the
Among
Fructus
has
genus
'Java'.
2.
Euthemis minor
in
Hook.,
N.I. 32
(1873)
412;
Béng. 62, II (1893)
235;
As. Soc.
Wallich
Am.
J.
2517
Str.
Lobb
Br.
s.n.
(1901)
15
(1940)
E.
R. As.
Soc.
in
E.
5 —12
65
up to
Leaves
by
3
33
Fl. Mal. Pen.
(1952)
&
224.
island,
18; Wall. in
fr.
1
(1901)
1
34,
I
the
the
as
collection
1
(i860)
209,
Fl. Ind.
2
(1824)
Ann.
1
(1849)
Scheffer,
534;
168; Ridl.,
J.
Str.
Br.
Merr., J.
32;
Soc.
R.
As.
Str. Br. R.
Kcw Bull.
Jack f, Singapore.
Nat.
As.
(1875) 526; King, J.
1
(1922) 368; Airy Shaw,
Type:
1812 —1813.
in
(1842) 528;
(1917)
2
Roxb.,
number,
this material
presume
and Palembang
Fl. Br. Ind.
—
however,
Probably
date and collector's
no
in that
15
have been
1920.
(1940)
Neotype:
—
1822.
23
(1862)
Ann.
163; Walp.,
Beih. Bot. Centralbl.
(1921) 389; Airy Shaw,
86
P., Nat. Pfl. Fam.
&
Kew Bull.
10
i—2
Bot.
cm,
up
m.
to
Jahrb.
34,
Kew Bull.
3,
S.
60
3;
2
(1868)
7
(1917)
(1940)
f ?,
in B
Hall.
2
152,
(1917)
f. 78;
544;
33; Merr.,
250.
Type:
—
rj
(1926)
33;
Bot. Centralbl.
Airy Shaw,
fl.
at
apart.
apex,
mucronate,
caducous,
on
branches
34,
2
(1917)
fl. XI-1897.
Kew Bull.
tapering
branches ±
c.
c.
lamina
racemes,
trigonous,
Bull.
(1940)
250.
—
n-VI-1923.
long, winged petiole,
Injlorescences lax
Kew
Labuan I.
/, Beih.
310;
Borneo, Sampit,
cm
Bartell., Malpighia
Airy Shaw,
SING), Sumatra, Siak, Penassa,
drooping; rachis slender;
mm,
(1895)
6
34,
Branchlets slender, blackish. Stipules
obtuse
1 mm
(1906)
(K holo, P,
(B f?),
( —5 ?)
nerves c.
sometimes
by
in
with his
one.
doubtful,
very
bearing
Centralbl.
Bot.
regarding
comparison
undoubtedly
Philippines.
(1821)
5
and
the Herbier d’Alleizette and
from
of Banka
Bartell., Malpighia
gooo
with
1^—4
denticulate,
lateral,
the
occur
Hook./,
Benn. in
in
sheets that
two
in
unknown collection
Type:
—
Hackenberg
Shrublet
4 —15
neotype
the expression
major'. Obviously,
1144 is
169; Hall. /, Beih. Bot. Centralbl.
hackenbergii Diels,
serrulate.
i,
(1901) 169; Hall./,
15
Type: Ridley
Type:
from the
as
(K holo, P), Borneo, fl. 1853.
250.
—
The Kew
Hooker /, Trans.
to
published
were
leucocarpa)
E.
(1859) 675; ibid. Suppl.
2
1,
Arb.
E. ciliata Pearson,
33.
most
among
Jack, differt omnibus partibus duplo
annotation is
Hook. /, Trans. Linn. Soc.
engleri Gilg
E.
Misc.
(K), Singapore, fl.
Bartell., Malpighia
J.
species
duplicate
a
reference
(1831) 69; Walp., Repert.
2
Hall./, Beih.
34;
obtusifolia
E.
This
native Ochnaceae
(1921) 388; Ridl.,
86
Merr.,
250;
are
of Ramos
Horsfield and
Jack, Malay
Tijd.
(1910)
and
1819
1954, 219—227).
certainly
be mistaken for another
cannot
been collected
Bot. Misc.
Miq., Fl. Ind. Bat.
54
and it
(=
Sandakan and vicinity
no
179;
Soc.
21,
is
species
sed duplo
Euthemis’
during Horsfield's exploration
collected
Jack
at
two new
leucocarpa,
Jackianae,
specimen
never
Since
is
304;
E.
collected by
specimen,
is labelled
'Ab E.
the Philippines.
otherwise
A Kew
in
mention this
not
in 1822.1 accept this collection
Singapore
examined, there
specimens
made by Ramos
was
65
In Brunei the fruits
Singapore
at
in Index Kew. with
for ‘Jack's
( E. robusta)
from
reported
at
is cited
mislabelled. The Leyden
is
collected
was
1824. Cowan does
In the latter paper
used
was
species
own new
Area
reported from Malaya.
the type of this species,
stated:
was
(....).
Jackianae’
Indo-Pacific
R. Bot. Card. Edinb.
(Not.
as
jackiana’
(i860) 163.
Euthemis robusta it
majoribus,
of the
Jack.
‘E.
name
Linn. Soc. 23
Ochnaceae
is
roots
in
collection Wallich 2516 p.p. made
of E.
the
specimen
by shipwreck
that is marked
specimen
of
revision
A
eye-diseases.
Jack's
was
:
application of the
used against
are
Kanis
by
3
at
base,
5—30 cm
shortened;
1
mm
I mm,
oblong
long,
to
acuminate,
oblanceolate,
margin
long,
bracts
often
on
faintly
pseudo-
the rachis
acuminate. Flowers
66
BLUMEA
complete, often
functionally polygamous,
by 2\
4
—
2\—3
in $
<? flowers
or
ovary;
5-lobed,
up
c.
6
to
by
0.2
SUMATRA.
P.
6703
Ridley
Hullett
Batu
Teijsmann H.B.
Hullett
conical
widening
base. Fruit globular,
at
c.
by 2\
4
mm.
G.
G.
Wallich
(BM, SING),
12525
G. Daik:
Tanda:
s.n.
Biinnemeijer
Biinnemeijer 6859
(BO,
Sumatra, Banka, Djebus: Teijsmann
S.
—
Ridley
(BM, SING);
5725
CAL, SING);
(BO, L).
(P),
long,
mm
by
yellow. Ovary
mm,
c.
P. Batam:
SING);
Rajah:
Bui:
Biinnemeijer
De Leeuw 5
1256 (BO, K, L),
5876
1
6—8
depressed globular, shallowly
semiannular,
SING, U);
& Anta
Pierre
Singapore:
L,
(BO,
3373
P,
c.
unequal,
lanceolate,
and Borneo.
5699 (BM,
s.n.
style
long, broadly
(K,
gooo
(SING);
s.n.
by
—
reduced,
red. Seeds
Petals
Chua
(K);
2517
1787
s.n.
(BO, L), Teijsmann
(BO, L, SING).
Chu
Kang:
Ridley
14191
SING).
Sarawak,
BORNEO.
1712
Brunig
S
(FI, K),
Lintang
(K, L);
S
Path:
Bt.
Bako
Jacobs
Tambi:
1662
1647 (K),
Fuchs
Sanaboh:
F.R.:
W. Borneo:
Jaheri
Parret
Main
Mas :
B2193 (BO, L);
Amai Ambit:
(BO).
Hub.
—
Central
(K, SING).
(K, L);
Remarks:
(BO);
1890
(BO, L);
level
sea
drier and
pĕtikawo
Polygamy
was
specimens which
K,
to
ex
Hist. Pi.
4
Winkler
K,
L); Tabang,
Main 1704
H.B.
often found
8463,
Kostermans
near
Pasir:
G.
(SING);
3924
Ramos
1315
(A),
26305 (K,L).-
B1445 (BO, K, L),
8464 (BO, L);
Teijsmann H.B.
—
S.
G.
1599
Borneo, Djihi:
(BO); Bulungan,
159
Wyatt
Richards
Lobok
(BO, K, L); Sintang,
Hallier
L, SING), 7738 (BO).
G. Palimasan:
S.
SAN
P. Karimata:
Amdjah
Bt. Dulit:
Res.:
Sam
Kenepai:
(L);
1421
Miiller Mts.:
S.
Sebakis:
12938 (K, L).
the previous
species,
but
places.
Kedayan);
of this species
recorded in Euthemis
are
doubtlessly functionally
mata
pĕlandok
meets
with
(Banka).
Borneo:
buah
(W. Borneo).
the
same
difficulties, already
have bisexual
or
before, but
male. I
in this
am
female
functionally
species
not
I examined
certain whether
many
fruiting
flowers, but these possibilities
both realised.
DC., Prod.
Walp.,
(K, L),
156 (BO); Tajan:
Sibu:
near
5544
Chai &
Path:
Res., Sipitang, Menggalong
Sandakan
K);
G.
(L),
2987
Madjang: Teijsmann
P.
not
1
(1824)
Pi. Vase. Gen.
148; Meisn.,
120,
BO,
2364 (BO, L);
m,
SAR
SAN 47343
7646 (A, BO,
Tribe
Ging.
Pandan
(SING); Marudi,
2882
bĕlusung mèrah, kĕtjing pĕlanduk
(Sabah,
Telok
(K, L); 4th Div.,
Interior
(K, P);
s.n.
26oj (A,
Hans
Borneo,
1250
S. 21206
S.
Brooke
28331 (L);
leucocarpa.
generally
probably
Bika:
exposed
The typification
E.
S.
E.
Sumatra:
names:
B
Smitinand
Purseglove 4888 (K, L, SING),
(K, L);
Pickles
(SING),
s.n.
(K); Bungoh Ra.:
(K, SING); Kuching:
s.n.
JCj
■It 3 (K, L),
Asam:
18012
Plat.:
(K, L),
Dunselman
Hallier
Buwalda
up
more
Lobb
(BO, L);
—
(BO,
9273
Apau
SAN 20031
G. Kelam:
Borneo, Sampit:
Unyotig
Pantai: Lumau
Meijer
Main
B13408
(BO, L),
for
specimens
Bt.
Usun
Sabah:
—
S..
Baram: Anderson
Bg.
(K, L);
Telok
Enoch
Bartlett
s.n.
(A, BO, SING); 3rd Div., Kabang
coll. 5123
(K);
2131
11220
Pladjang:
(Sarawak);
explained
Native
(E), Ridley
Haviland
4« (E),
Chai
Tajor:
(Mt. Matang):
E.
'.211
coll.
Carrick &
(L);
14
Serapi
Native
C
Ulu Tiau: Asahak
Mts.,
F.R.:
13, B
Telok
G.
(A);
coll.
Native
(BO, L); Pangkalan:
s.n.
1794
From
in
Vernacular
itĕk
Leila
2226
(SING),
7669 (SING),
Kanis B
4679 (L);
3268 (K, L, SING).
9251
Ecology:
generally
6772
Hallier
Winkler
Kostermans
S.
S
Brunig
Stupong:
Hose
SAN .21807
(BM);
s.n.
s.n.
(A, K, SING),
Moulton
Meijer
Rycroft
G.
coll.
(K, L),
Sleumer
(K);
Native
Haviland
Santubong:
G.
N.P.:
(K), Yacup
21244
(FI),
5504
17285 (K, L);
.
Smith C.F • 7933 2
Div.:
1st
2525
7627 (SING);
(L, SING);
9779
are
Chie
(BO, L, SING),
PENINSULA.
Beccari
Paie
Penassa:
Siak,
3!
mm;
ij
much
Malay Peninsula,
S.
acuminate.
Anthers
| by
2
mm
c. 0.2
G. Maras: Kostermans
(BO, K, L);
MALAY
(K,
c.
1968
i,
conferted. Sepals obovate,
more
apex,
very
SING, U), 6875, 6887 (BO), Teijsmann
(BO, L); Plangas:
s.n.
S.
Lingga: Nottg
(BO), 6713
L,
K,
Riau,
flowers
style
0.4 mm;
Distribution: Sumatra,
(SING);
5-ribbed,
in $
acuminate, 5-ribbed,
0,
mm
obovoid,
ovary
2 or
the
near
No.
XVI,
in $ flowers.
distinctly reflexed
mm,
into the
ciliate
mm,
—
VOL.
Ann.
(1873)
7
(1868)
339;
1
315,
Engl.,
err.
(1836)
220;
3.
SAUVAGESIEAE
‘Sauvageae
21;
ibid.
2
’;
Lindl.,
(1836) 64;
Intr.
B.
&
Nat.
Le Maout & Decne., Traité Gén. Bot.
Nova Acta
Leop.-Carol.
Akad.
Syst.
Bot.
H., Gen. Pi.
37,
2
1
(1830)
(1862)
(1868)
432;
Baill.,
(1874)
22;
Eichl.,
A.
Bliitendiagr.
burgieae
Pfl.
the
Soler., Syst.
Hook., Lond. J.
D.-T.
145;
Ochnaceae
Anat.
H., Gen. Syph.
&
shrubs,
undershrubs. Lateral
or
10
—
in
00
or
1
67
95—97.
&
P., Nat.
in E.
&
P.,
(1966)
16
Luxem-
—
in E.
Gilg
(1903) 316; Gilg
of leaves
nerves
opening by
ovules
the style,
which represent
dehiscing
along
sutures
archingly
00.
adnate
lateral slits.
2
Fruit
1-celled with intruding placentas;
Nat.
39—55.
—
a
joined
base
at
2
acuminate
capsule,
of the
the margins
short
a
—3-carpelled,
Ovary
coriaceous
the
near
to
by
Seeds
carpels.
small.
CSD,
Distribution:
genus
595]
593,
whorls. Stamens ±
more
anthers
tube of inner staminodes;
1
(1899)
Dicot.
(1846)
Bot. 5
Area
Indo-Pacific
Sauvagesia L.
margin. Staminodes
usually
the
of
(1925) 64, emend, illeg.; Decker, Phytomorphology
2, 21
genus
Trees,
(1895)
6
3,
of
revision
257 —259;
Planch, in
Pfl. Fam. ed.
Type
(1878)
2
\non
Fam.
Kanis: A
6
genera
confined
to
SE. Asia, the Malesian
area,
and the
occurring in Africa and America, and several others confined
deviations from the
Remarks: The following
found in American
Pacific;
the neotropics.
given above
characteristics
general
western
to
are
genera:
Staminodes absent.
(a)
(b)
Stamens
(c)
Ovary
3-celled.
(d)
Ovary
5-carpelled,
10
or
cv>.
5-celled.
SINIA
5.
Notizbl.
Diels,
347,
350.
—
Shrublets.
Type
entire. Nerves
with
the
at
an
compound
at
base.
Ovary
of
angle
Bot.
Fr.
(1965)
111
10
in
2
slightly curved.
70 —8o°,
branches
racemes;
of 2 types, the inner
co
connate
stamens
Bull. Soc.
Vidal,
888;
ascending
terminal, many-flowered,
Flowers bisexual. Staminodes
and
(1930)
10
Sinia rhodoleuca Diels.
species:
Stipules
Inflorescences
Berlin
Gart.
Bot.
much shortened.
alternating whorls, mutually
stigma
3-carpelled;
Fruit and
1.
seeds
unknown.
Distribution: One
I.
(1965)
hi
shan, Lohsiang,
Shrublet
600 —1000
up
8—13 by
serrulate,
c.
\
cm
—3"|
2
(?)
Stipules
2—5
narrowly
long
mm
acuminate
at
apex,
lanceolate,
±
tapering
erect
nerves
papyraceous,
by if—3
by i—1]
10,
mm,
nerve.
mm
Distribution:
c.
apart.
gland-like
I
Stamens
long; style
SE.
mm
or
Petals
at
Bull. Soc.
elliptic,
at
base,
those
with
—3
c.
1
mm
mm
mm
long,
long; pedicels filiform,
(?). Sepals
4 —5
I
by
anthers
3
3 —5
c. 2
mm,
mm
the
to
ones
ovate to ovate-
c.
—if
opposite
long filaments;
long; stigma
2
obovate-lanceolate,
base, margin irregularly
cm
somewhat spatulate,
connate
pinkish
long;
inner
stamens
by \—f
mm
Yao-shan, Lohsiang:
Reported
from 600—1000
Sin
m
8152
(A, Bf,
altitude.
3 —6
mm,
long. Ovary
minute. Fruit and seeds unknown.
China.
Mt.
Kwangsi,
Ecology:
cv,
mm
glandulose-ciliate.
mm,
spatulate,
1
with 3 —5 ± distinct, parallel nerves, the alternating
distinct
ovoid, i\ —3
c.
to
up
lamina
petiole;
long (?);bracts small, glandulose-ciliate. Flowers
staminodes
67
(1930) 889; Vidal,
10
5 —8
4—5
CHINA.
high.
with
sharply
white. Outer staminodes
i
Gart. Berlin
Inflorescences
oblong,
with
Bot.
alt., fl. 4-V-1929.
m
m
Leaves
cm,
China.
f. ib.— Type: Sin 8197 (B f holo, K, SYS), Kwangsi, Mt. Yao-
350,
to
glandulose-ciliate.
SE.
Diels, Notizbl.
Sinia rhodoleuca
Bot. Fr.
in
species
SYS), 8197 (Bf, K, SYS).
Flowering
in
April and May.
BLUMEA
68
Remarks:
specimens
are
Diels' original
in A and K
are
VOL.
Bull. Soc. Bot. Fr.
Bot.
521
Fr.
(1948)
95
31,
(1965)
in
Distephana
=
405.
Distephania
—
Mus. Paris
illeg.-, Vidal, ibid.
Indosinia involucrata (Gagnep.)
[«o«
(1805)
6
(1965)
111
Gagnep., Bull.
346 —350.
panicles. Flowers
corolla). Petals
soon
One
Distribution:
into
in
species
stamens
S.
Distephania involucrata Gagnep., Bull.
ibid.
(1950)
97
Vidal, ibid,
85, fig.; Duchaigne
(1965)
in
Trang, forest,
346, f.
1700
Shrubs
with
1 —3
5
Vidal, Bull.
up
to
c.
3
Flowers
1
in
erect.
to
acute
at
Sepals ovate-oblong,
—2\ by
2
i
long; style
long,
c.
6 mm
Distribution:
S. VIETNAM.
Ninh Hoa:
mm
stigma
1.
Bot.
(1948)
Fr.
31,
in
(1965)
405.
—
illeg.-, Du Chalard,
nom.
106—108, f.
i,
2;
Paratype: Poilane 6489 (K, L, P), Annam,
—
triangular,
±
at
tapering
14—18
1 —2
3 —4
to
by
c.
1
6|—8 by
c.
base,
the alternating
filaments, adnate
long,
acuminate. Leaves
linear-lanceolate,
by i-J
4
mm,
2\
ones
by
acute,
the
long,
cm
I—2
glandulose-ciliate.
acute,
mm,
glandulose-ciliate, distinctly
reflexed.
white, distinctly
mm, acute,
to
5 —15
margin serrulate, chartaceous,
stamens
6—6% by
c.
§
7
—8
mm
by
c.
with
mm,
distinct
1
tube of staminodes; anthers subtriangular,
to
long, stigma capitulate.
mm
to
long; pedicels filiform,
cm
up
Seeds obliquely obovoid,
0.
a
Fruit
3
c.
little
Ovary ovoid,
mucro.
2 —3
ovoid, cuspidate by the style,
by
I
—mm,
testa
c.
mm
12 mm
reticulately ribbed.
S. Vietnam.
Nha
Poilane
Ecology:
1.
the connective terminating in
mm,
3 —4
and
calyx
alternating
Lectotype: Poilane 3498 (P), Annam, Nha
purplish, those opposite
nerves,
Stamens with short
nerve.
pi.
±
apex,
Inflorescences
distinct, parallel
2
2
2-carpelled;
Chalard, ibid. 98 (1951)
—
fruit; bracts ovate-oblong,
Staminodes linear-lanceolate,
in
17-V-1923.
reflexed in fruit. Petals ovate-oblong,
with
Du
high. Stipules
m
apart.
mm
cm
2,
&
Soc.
95
long petiole; lamina obovate-lanceolate
mm
obtuse
cm,
nerves
2-£ (?)
to
up
—io
io,
winged.
not
Soc. Bot. Fr.
fl. 20-V-1922.
alt.,
m
ia,
Nha Trang, N. of Ninh Hoa, fl.
base. Ovary
at
—
Vietnam.
Indosinia involucrata (Gagnep.)
I.
connate
parts. Seeds
2
70
many-flowered
between
axis
in size. Staminodes
of
angle
at an
terminal,
(prolonged
anthophore
an
surpassing the sepals
and with the
whorls, mutually
Style of ripe fruits splitting
the margin. Inflorescences
near
with
bisexual,
in anthesis
species:
Type
—
2,
Soc.
Vidal.
slightly curved
or
are
Nomencl. ed.
Steud.,
396]
Shrubs with slender branchlets. Stipules lacerate. Nerves ascending
8o°, straight
specimens
presented here
INDOSINIA
Ann.
Juss.,
nam.
Cantonese
the data
obtain,
to
description.
6.
(1840)
in the Berlin herbarium. The
specimens have been burnt
available in P. Since good material is difficult
Vidal,
1968
i,
only fragments, but photographs of the
mainly compiled from Diels'
I
No.
XVI,
Trang:
Poilane
6489 (K,
Reported
3498 (P);
Cascade
W. of Nha
Trang:
Poilane
3656 (K,
L,
P);
N.
of
L, P).
twice
from
forest
at
1700
m
altitude.
Flowering
and
fruiting
in May.
Remarks:
a
The
genus
member of the
Distephania
was
wrongly
Saxifragaceae-Escalonieae.
He
because of the bract-like appearance of the sepals,
by
an
anthophore. Consequently,
staminodes
as
petals.
the corolla
described
by
was
separated from the
interpreted
as
(1948)
Gagnepain
interpreted the calyx
a
as
rest
an
as
involucre,
of the flower
calyx and
5
of the
Kanis :
A.
Van
as
presumed the
He
1951.
communication).
and later
(Vidal,
it
on
It
taxonomic
status
(1965) published
has
Gagnep.
be treated
to
Ned. Kruidk.
Korth.,
(1862)
as
2
7of; Ridl., J.
the
to
Distephania
since
under Art.
Juss.
of the
75
NECKIA
Walp.,
358;
(1873)
4
49
(1908)
n;
Inflorescences
Fl. Mai.
but only
the inner
3-carpelled; stigma
forming
ones
Kew Bull.
Shaw,
angle of
at an
the rachis
reduced;
much
tube
a
base;
at
Style of ripe fruit splitting
1.
(1922)
1
3,
6
134;
(1940)
bearing
70—8o°,
small
some
pedicels articulate. Flowers bisexual. Staminodes
terminal flower;
I
Pen.
Fam.
Korth.
serrata
axillary,
Nat. Pfl.
Gen.
H.,
&
67; B.
P.,
&
f. 33F; Airy
(1925) 81,
2, 21
(1852)
2
in E.
355;
Str. Br. R. As. Soc.
Ann.
Gilg
undershrubs. Stipules pectinate. Nerves ascending
or
types,
Pi.
Type species: Neckia
—
curved.
slightly
(1848)
1
Hist.
P., Nat. Pfl. Fam. ed.
&
in E.
(key).
Shrublets
of
Indosinia,
name
new
later homonym of Distephana
a
Arch.
Baill.,
120;
f.
(1895) 148,
bracts,
in 1959
Mme Lecompte
of the flower. He referred the genus
reinterpretation
a
7.
252
Luxemburgieae by
Code.
present
Gilg
early
as
be in the Ochnaceae (personal
to
the Ochnaceae by Hoogland
to
He also introduced the
Ochnaceae-Sauvagesieae.
1
69
interpretation of the flowerparts
within the
precisely placed
Area
Indo-Pacific
1965).
Vidal
Pi.
the
of
of the genus
also referred (in sched.)
was
more
was
Ochnaceae
doubted Gagnepain's
already
Steenis
the
of
A revision
stamens
adnate
Seeds
into 3 parts.
CV),
the tube. Ovary
to
winged.
not
Distribution: One species in Sumatra, S. Malay Peninsula, Borneo, and the Philippines.
Neckia
I.
Koord.,
Ic.
I, 2
(1859) 118;
Ann. Mus. Bot.
Bogor.
1, 4
(1901)
76; Ridl., J. Str. Br. R. As.
Singgalang
N.
Lobb
N.
Str. Br. R. As.
s.n.
(K
humilis
Ridl., J.
N.
f.
13.
Lake
—
&
fr.
—
R.
As.
Ridl., J.
Kelsall
s.n.
f.
major Airy Shaw,
Type: Korthals
—
fr.
Ridl., J.
P.
Soc.
(1908)
49
13,
&
14;
(L holo, U), Sumatra,
s.n.
Aim.
158; Walp.,
Kew Bull.
(1868)
7
(1940)
252.
—
221;
Type:
Soc.
(1908)
314
R. As.
(i860)
23
14.
(K),
—
N. Borneo,
Soc.
49
G.
Johore,
Walp.,
159;
Lectotype:
(1908)
S.
Ann.
s.n.
Labuan I.,
11;
&
fr.
20-X-1892;
(1868)
7
(K),
N.
221,
Borneo,
fr.
Fl. Mal.
fl.
Pelepak,
Janeng, fr.
Lobb
Pen.
1905.
1
(1922)
Hullett
134,
Paratypes:
—
s.n.
(SING),
Fl. Mai. Pen.
(1940)
252.
Str. Br. R. As.
1
(1922)
135.
Soc.
—
N.
49
(1908)
Type: Ridley 2269 (BM, K,
—
11.
—
N.
Hook, fl
lancifolia
SING
f.
halo),
1891.
Str. Br. R. As.
Kew Bull.
Soc.
49
(1940)
(1908)
250.
—
12;
Merr., J. Str. Br. R. As. Soc.
Type: Ridley
9034
(K,
86
SING
holo),
(K.
SING
Bongaya, fl. X-1887.
N. Borneo, Labuk Bay,
Ridl., J.
(1852) 67;
16-VII-1893.
Bull.
fr.
2
(1869) 218; Boerl.
4
1853.
angustifolia Ridl., J.
var.
Kew
13;
(i860)
Airy Shaw,
(SING), Johore,
s.n.
major Ridl.,
Tahan R.,
holo), Riau,
49
Br.
Str.
(1921) 388; Airy Shaw,
N.
Soc.
(SING),
ft.,
malayana Ridl.
N. klossii
(1908)
Paratype: Motley
200 —300
N. distans
49
N. Borneo, fl.
Lectotype: Kloss
malayana Ridl.
Pahang,
Soc.
Bat.
Lugd.
fr.
Hook, fl, Trans. Linn.
malayana
Lingga,
N.
holo),
Str. Br.
Labuan I.,
(1921) 388.
Hook, fl, Trans. Linn. Soc. 23
lancifolia
Ridl., J.
86
Malintang,
& G.
t.
1,
Ann.
(1848) 358; Walp.,
I
Bat.
Merr., J. Str. Br. R. As. Soc.
G.
Arch.
Korth., Ned. Kruidk.
serrata
Miq., Fl. Ind.
Str.
Br. R. As.
Soc.
49
(1908)
13.
—
Type:
Kloss
s.n.
Batam, fl. III-1906.
parviflora Ridl., J.
Str.
Br.
R. As. Soc. 49
(1908)
14;
Merr., J. Str. Br. R. As. Soc.
BLUMEA
70
388.
(1921)
86
Type: Ridley
—
(1921)
86
Soc.
fl.
Damoes,
fl.
Ajah,
holo),
klossii Ridl.
obovata
6248 (L,
Shrublet
i —3
mostly
1
to
up
1^—3
linear,
to,
by
creamy
—0.6
SUMATRA.
djanki:
L,
G.
Kloss
Chie
600
(K,
s.n.
L,
(BO,
S.
S.F.
Endau:
„
S.
—
Lake
Holttum
S.F.
(K, L);
Ulu
s.n.
mm
1
Batten
P.
(BO).
s.n.
15775
I
s.n.
by
0.5
(L);
s.n.
Taram:
Kloss
(BO);
0.3
Batu
(SING);
s.n.
S.
Nipa:
s.n.
Sinclair
mm,
mm
spatulate,
the free
filaments,
yellow. Ovary ovoid,
mm,
subtrigonous
in
cross-
areolate.
mm,
Is., P.
Corner
K,
S.
S.F. 3g62b
s.n.
(BO, L),
Brooks
K.F.N.
—
Kuala
W.
5003
S.F.
G.
Hullett
200
P.
Pahang,
Tahan:
(SING), Nong
s.n.
G.
Panti:
(K);
Posthumus
G. Tahan:
Seimund
Henderson
19995
(BO,
Battam:
(K).
98017 (K);
(K, L, SING);
Nur
7605
Labis:
Iboet
Arch.,
655 (BO); Bangko:
(SING).
SING);
Merton
(BO). —
Raap 567
Riau
(L);
4249
Posthumus
Gatong,
Susor:
Pini:
Meijer 6854, 6952, 6963 (L); Muarapa-
Meijer
Karing:
(BM,
Rotan
1—3
gland-like, J—I
terete
cm
pedicels
elliptic, 2|—5
14076 (BO, K, SING),
S.F.
Taluk:
19666 (SING),
S.F.
(SING);
\
c.
to
cm,
Borneo, and the Philippines.
(SING); Kluang F.R.: Ng
Pulai:
by
4—q\
or
clavate. Fruit
Sumatra, Bengkulu, Lebongtandai:
G.
Corner
Stamens with
—1| by
up
q\
I —j
± half connate, the free lobes
long,
Djambi,
S.
rachis,
„
longer.
R.: Ridley 2269
—
small stipules;
lanceolate,
Lingga: Biinnemeijer 6585
—
by f
3 —25
filiform
unequal, long spatulate
Siberut:
Is.,
Stipules compound,
base, biserrulate, membranous
with
Sepals
long; stigma
Pooll
at
Sulit
Type:
—
lacerate. Leaves with
long,
cm
2.
IV—V-1948.
often unbranched.
.
Malay Peninsula,
(BM, SING),
Segum:
Sarawak,
Matang:
s.n.
Bagacay,
ph
329,
fr.
(K holo),
1413
23-VIII-1932.
S.F.
958
Hatiijj &
(SING);
38253 (SING);
Blumut:
S.F.
Holttum
Ridley 4164 (K, SING),
Kota
Shah
Tinggi:
(BO, SING);
MS
Pelepah:
444
Kloss
(SING).
BORNEO.
G.
(1954)
82
fr. 27-VTII-1932.
Richards
fr.
&
Type: Richards
—
&
dark red. Petals
green, turning
15546 (K, SING). —Johore,
SING);
18090
stamens
Koorders
Tahan
& Kelsal
BO,
10619 (BM,
C.F.
cv),
mm
Trengganu, Kemaman,
(SING);
Mahmud
fl.
fl.
sometimes with
„
x—if
Mentawei
Segati:
Teijsmann
SING).
8093
G.Janeng:
c.
SING); Lingga Arch.,
Malay Peninsula.
Nur
Batu
(K holo), Sumatra,
251.
m,
Type:
—
Inflorescences
long,
mm
Malintang: Korthals
G.
6412 (K, L);
(SING),
s.n.
Str.
J.
Bt.
Type: Ridley 4164
—
long obovate-lanceolate,
to
Seeds ellipsoid, about
Sumatra, S.
Riau,
—
style
Sumatra, Sibolga:
N.
135.
700
c.
m,
300
Sc.
J.
anthers lanceolate,
mm.
Singgalang &
Buwalda
SING).
15 —25,
mm,
by 2\
5
Distribution:
Sumatra,
13; Merr.,
t.
397,
(1922)
1
251.
under
pinkish
or
,
long;
mm
0.5
c.
2
c.
lobes alternating with the
i
(1940)
Flowers pendulous.
long.
mm
the
section,
(BO), Borneo,
(BO holo, L), Borneo,
Long Kapa,
apart.
mm
inner staminodes
up
Hallier 638
Type: Brooks 7603
—
unequal, acicular, \—I
white. Outer staminodes
mm,
77.
near
lamina obovate
,
i—1.2
R. As.
Lectotype: Jalter i 433 (BO
—
(1910)
57
1626
Samar, Concord,
long;
part
IX-1905.
Br.
fl. VI-1922.
Quis., Philip.
&
parts
margins dentate,
to
(1925)
Bull.
Kew
holo?),
nerves
8—15
by 11—2
5
Fr.
acuminate, sometimes obtuse, tapering
acute to
filiform,
acute,
Bot.
high, often much smaller,
m
long petiole;
bracts
long;
Str.
fr. X-1892.
Long Kapa,
near
PNH
to
up
chartaceous,
to
fr.
Puak,
Merr., J.
13;
Paratype:
—
Type: J alteri
—
Mt. Dulit
Merr.
wide, the
mm
mm
15
fr. 1896—97.
&
ft.,
iooo
Airy Shaw,
philippinensis
P.N.H.
252.
t.
borneensis Airy Shaw, Kew Bull. (1940)
var.
Mt. Dulit
Sarawak,
N.
(1940)
G. Panti,
(K holo), Sarawak,
N.
Kew Bull.
Kew Bull.
grandifolia Ridl.,
N.
397,
f. minor Ridl., Fl. Mai. Pen.
Johore,
Bengkulu, Lebongtandai,
147Q
holo), Sarawak,
(1910)
1896—97.
SING
N.
SING
1968
I,
57
Bull. Soc.
Capit.,
(1921) 388.
86
Soc.
malayana Ridl.
N.
(K,
As.
No.
22—24-X-1893.
Sauvagesia jaheriana
Br. R.
fl.
Bloe-oe,
S.
(K,
12320
388; Airy Shaw,
holo, L), Borneo,
G.
XVI,
Soc. Bot. Fr.
ovalifolia Capit., Bull.
N.
VOL.
Anderson
(SING), Ridley
Purseglove
4877
1st
15
s.n.
(K, L);
Div.,
G.
(SING),
Pueh:
(BM, K, SING);
Telok
Purseglove 4807 (L,
Beccari P.B.
G.
1459
(Fl),
Miss
SING);
Brooke
Santubong: Mjoberg
Tajor: Purseglove
4935
161
(K, L); Tg.
Puak:
9512
(BM);
Po:
Ridley
(L),
Bako
Miss
Nat.
Brooke
(K, SING);
12320
Clemens
20921
(K),
Hullett
Park,
Telok Asam:
10610
(BM, G, L);
Kanis :
A.
Ulu
G,
Serait:
Beccari
P.B.
G.
Miss Brooke
Teneong:
Ra.,
F.
Ulu
R.,
Miss
Lobb
Lobb
Marai
Parai : Clemens
Kanis
SAN 53971
C.F.
Kanis &
314
Sandakan
Winkler
4897
9087,
9090
BO,
(A,
L),
Found
Ecology:
as
G.
E.
reported
Remarks:
F. R.:
to
Shaw
Airy
had
1200
Sulit
P.N.H.
on
key
leaves
Bornean
the whole
a
a
fine
normal,
from
survey
The
apex,
narrower
found
are
Neckia specimens
some
and
descriptions
as
col-
ovalifolia Capit.
a
species
except
was
not
for N.
able
with
be conspecific
to
a
he
the specimens
on
Hooker and Ridley,
to
one
fruiting specimens,
are
relatively broad, obovate
with
over
are
habit indeed;
1200
a
regarded
moss
by
still
Shaw
one
m
to
than
of all
100
taxa
an acute to
collections of Neckia from
described in the
It
appears
which
type), however,
(.Richards
acuminate
apex
and in
my
of N.
a
This
draw
fully
obovata
has leaves which
some
1479,
to
together
name
1381)
has
genus.
impossible
form
described under the
specimen (Richards
1413,
leaves that
1672)
show
opinion the
are
more
forms
closely related.
Neckia
must
primarily
find correlation of
and,
zones.
carpet
me
more
from Mt. Dulit
specimens
genus
not
and subalpine
in
study
abundance of specimens
as
I
shady places and will be capable
growing
forests
kerangas
tuff.
or
of the variability in Neckia.
leaves with
I could
to
the other ( Richards
The variability in the
not
Batu
S. Telen:
occasional inundation. Soils
to
of the Neckias collected
specimens Airy
Other
this mountain
montane
most
on
slopes, along brooklets
wet
laticifolia Hook. f. Though he
N.
Puhus
Sarawak in 1932. In 1940 he published
described by
which type material
uncommon
at
obtuse.
conditions.
is
to
(L);Bt.
Bt.
(BO);
Meijer 1856 (BO).
in
places
identify
to
leaf-characters and
within this
mass.
rather
rounded
distinctly
on
opportunity
among
demarcations
intergrading
arc
As
species.
I had the
area,
material gives
have
shady
subject
or
Landak:
1273
P. Nunukan: Kostermans
(BO, L);
Tarakan:
P.
Borneo,
453
Arsat:
Leila F.R.:
(L).
in moist,
the earlier
presumed
W.
Bahandoi:
Sandakan:
Winkler
Jaheri
L);
s.n.
Kinabalu,
newly added.
was
Fortunately
any
on
the species
he
mainly based
was
(BO, L);
K,
(BO);
—
12786 (L); Long
Kostermans
6248
with the difficulty
based
Bluhu:
S.
Mt.
1370
Hans
Malang:
boulders, cliffs, and
waterfalls
near
S.
(BM);
(K, SING);
9054
Barber
I.:
(A, BO, G,
s.n.
31018,
Div.,
N. Bt.
Sabah:
—
Labuan
500
Sinanggul SAN36613 (K).
2215a
altitude,
m
(K);
253
Dulit
SING); 5th
(L); Temburong,
s.n.
Darnton
Wain: Kostermans 4049
Ridl. which he united with N.
of the other
his
genus
original material,
the
Clemens
Belajan:
2215,
especially
met
hand. He accepted
at
malayana
see
of the
key
Kota Belud:
638 (BO);
lected by the Oxford University Expedition
tentative
Cox
sandstone, acid sand, sandy loam, loam,
as
Hotta
(L);
15341
(BM, K) ;
Belalong: Jacobs 5578 (K, L).
S.
Kidman
Upper Mahakam,
S.
BO, K,
1672 (A,
Teraja:
and
Bejangung:
Mujong:
9085 (G,L);
Hotta
Haviland & Hose 3155
(A, BM, BO, L, P), Topping
Hallier
on
Meijer 2022 (BO),
up
Ulu
Kana,
Richards
Padas:
Res.,
1122
Borneo,
G. Palimasan
in richer rainforests,
Bt.
Bt.
Penibukan:
China
Damus:
—
and small rivers, sometimes
are
Ramos
Concord, Bagacay:
Philippines. Samar,
well
Coast
K, L, SING); Balikpapan, S.
K,
(BO,
E.
Bantin:
1546 (K);
Ulu
(K);
(BM.
9613
Haviland
17612
Gilam Bakun: Miss Brooke
Baram Distr.:
S.
Miss Brooke
(K); Bg. Lupar,
745
Ashton S.
Res., Labuk, Bay, Bongaya: Ridley
Kebun
656 (L).
(K);
Bt. Dulit:
Res.,
71
(G, L); Bg. Saribas, Kalong:
Temburong
West
9498 (A, BO),
Ajah: Jaheri 1626, 1636a (BO);
Endert
S.
Collenette
Ulu Temalad:
Brunei, Seria,
—
Interior
(K);
(K, L);
(K);
1479
junction
(BM);
(BO, FI); Sambas,
s.n.
Hans
Mehipit:
near
53480 (K);
SAN
Meijer
Teijsmann
Mts.,
Area
Krusin:
(L); Serian,
Apeng:
Clemens 21821
10906 (K), 32489 (BO);
(K);
Clemens
835 (K),
1413,
10246 (L).
s.n.
Motley
(K),
s.n.
1381,
10006,
(K), Burbidge
10682
Hose
(L);
Indo-Pacific
the
of
19
G.
9187 (L); 4th Div., Bintulu,
13258 (L);
Hotta
s.n.
(K),
Brooke
.18, B
Div.,
Brooke
Miss
9012
Sinrok: Ashton S..
.18313
Richards
Long Kapa:
Bangar:
Antu:
2nd
21236 (K, L); Upper Rejang, Gat:
Similajau
Lawas:
Lubok
Ochnaceae
Kanis B
(K, L);
S. '53'5
Bah: Miss Brooke
3rd Div., Long
Ashton S.
Anderson
(FI);
499
17835 (K, L),
& Pate S.
Chai
G. Gaharu:
L);
the
of
A revision
over
a
am
to
rule,
be due
measurements
to
variation in the
ecological
with the altitude, but Neckia
the influence of the altitude is shown better in
convinced that,
germinate and
most
to
of all, the plants need moist,
produce
a
boulders. The smaller specimens
be dwarfed forms
by lack of
few flowers
I
nutrition.
have
seen
even
while
in the field
Shade, water-supply,
BLUMEA
72
occasional inundation,
VOL.
other ecological
or
No.
XVI,
1968
I,
circumstances
also
may
influence
exert
upon
the habit of the plants.
In
differences between local
distinguish infraspecific
in
Bornean
among
is
but
it
characters.
Generally
impracticable
seems
useful differences
no
to
to
me
geographical
found in the
are
be
There will probably
monospecific.
populations,
since
taxa,
distribution of morphological
is found
Korth.
genus Neckia
opinion the
my
genetic
the greatest variability
speaking,
specimens, whereas the plants with the largest leaves
found
are
Sumatra.
the Geneva Herbarium I
From
'}.
&
eastern
Malesia,
I
it
presume
the other herbaria
Korth.
serrata
Yoangen
to
only specimen of Neckia
erroneously labelled and belongs
was
of the Clemenses from Borneo made in 1933.
in
of Neckia
specimen
one
As this would be the
ft., 18-VI-1936'.
4500—5500
in
saw
M. S. Clemens 3420, Flora of New Guinea, Morobe Distr.,
to
found
ever
former collections
duplicate of this number
No
labelled
Yunzaing,
found
was
consulted.
8. INDOVETHIA
Boerl., Feestbundel
Pfl. Fam. Nachtr.
Pfl.
Fam. ed.
Shrublets.
21
2,
J. Veth (1894)
P.
2
(1900)
45;
(1925)
80.
Stipules pectinate.
Ic. Bogor.
89;
Bartell., Malpighia
Nerves
ascending
the other
developed,
bisexual. Staminodes
base.
at
not
io
(1921)
388.
t.
388.
Paratype:
&
2—4
Style of ripe
&
P., Nat.
in E.
&
P., Nat.
Boerl.
calophylla
of 60 —70°,
slightly curved.
some
lower branches
flowers; bracts small. Flowers
and with the
fruits splitting
stamens
into
3
connate
Seeds
parts.
by 2-J
io—35
branous,
10893
P.B.
1
(1901)
cm,
3
acuminate
mm
—
apart.
the alternating
flattened filaments,
mm
0
0,
;
style
to
c.
papillate.
I
Ic. Bogor.
R. As.
Soc.
holo),
L
K,
1
86
Borneo,
ro
12
mm
mm
E.
c.
by
long;
if
c.
by
8
to
up
Sepals
mm,
anthers
connective with
at
c.
with
|
mm
by
0.2
mm,
areolate.
Sumatra and NW. Borneo.
c.
3
mem2
mm
distinct, parallel
mm,
Fruit
smaller,
3^ —5 by
white. Staminodes
±
nerve.
mucro.
ones
obovate,
mm,
distinct
long
higher
to
with
long; stigma minute, trigonous.
0.8
the
2—3
mm,
1
5—10
oblanceolate,
long; rachis
IJ—if by 0.5—0.8
c.
86
—
when fruiting; bracts with
mm,
by
§
lamina
base, biserrulate,
cm
suborbicular
3 —4
2\ by
—
%
long
2
Soc.
X-1865.
unequal, lanceolate,
5 —20
mm
fl.
IV-1866.
the segments
apex, tapering
at
R. As.
Kuching,
hardly distinct petiole;
erect.
stamens
i|—2
cuneate
Seeds
Distribution: Central
the
ones
0.2—0.4
basally separated by the
mm
plate;
Br.
Merr., J. Str. Br.
Singhi, fl.
margins denticulate. Petals obovate,
mm,
FI,
Sarawak,
Inflorescences
up
ones
underneath. Flowers
spatulate, those opposite
nerves,
the lower
lanceolate,
shortly pubescent
90,
Str.
J.
(BO,
11;
(FI),
G.
Leaves with
acute to
t.
172,
763
P.B.
(FI), Sarawak,
1413
stouter nerves
small stipules,
1610
H.B.
—
Merr.,
173;
high. Stipules compound,
m
margins lacerate.
—
15
Beccari
pedicels filiform, mostly short,
across;
—
Teijsmann
J. Veth (1894)
P.
(1901)
15
and NW. Borneo.
Sumatra
1874?.
Lectotype:
Beccari
mm,
fr.
Bartell., Malpighia
—
Shrublet about
12
1.
in Central E.
species
Type:
—
I. beccariana
(1921)
i
stigma
Bartell., Malpighia
1;
fl.
Montrado,
c.
few
alternating whorls, mutually
calophylla Boerl., Feestbundel
Indovethia
10,
3
2
a
in E.
Gilg
mostly
racemes;
much shortened with
Engl,
winged.
(1897)
by
in
3-carpelled;
Ovary
Distribution: One
I.
ones
9;
170;
angle
at an
Inflorescences terminal, many-flowered, compound
well
(1901)
Indovethia
Type species:
—
(1897)
1
15
Stamens with
the anther cells
Ovary subglobose,
subglobose,
up
to
Kanis :
A.
SUMATRA.
Kuantan,
Sebako:
(K), 3609B
Krusin:
Serian,
10898
B
ist
S.
Brunig
=
H.B.
(FI);
(GAL),
270
Ridley
Area
Taluk:
Native coll.
(SING),
300
G.
(K, SING);
12324
ggi8 (L);
Ampat:
gsg6 (SING);?
73
Meijer 4162
(L, SING);
Haviland
S.
(K);
75g
Mt. Koum: Haviland
Samarahan:
(K).—W.
I78g
g4$5
763 (FI),
Brooke
Sematan,
(A, K);
2401
Brooke
Beccari P.B.
Kuching:
near
(E),
54
Matang:
G.
(BM, L);
Haviland & Hose
g67i (BM, L, P);
Borneo, Monterado:
H.B. 1468 (BO, L);
FI, K, L); Landak, Ngebang: Teijsmann
(BÓ,
1610
Indo-Pacific
(K, SING); Indragiri,
goog
Bau: Brooke
Pankalan
(BM);
Brooke
the
of
Ochnaceae
(L).
Puak:
(K);
the
of
Ridley
6256
Div.: Hullett
4653
Beccari P.B. 1413
Singhi:
3609
Buwalda
Sarawak,
Borneo.
S.
revision
Kelantan:
Riau, Siak,
Pangian:
A
Teijsmann
Hallier
G. Kelam:
(BO).
2500
Ecology:
forest;
Probably
forest
carpaceous
of
the edge
a
lowland
a
in
generally
places:
'on bank above
lake',
'young
in
a
in
(Haviland),
'Diptero-
old
forest', 'under
in
trees
and 'under rubber'
in the forest'
torrent
altitude and lowland
m
150
jungle'
(Meijer), 'on ground
hills'
loamy
on
from
reported
species,
shady
moist,
at
(Miss
Brooke).
Remark: The second specimen cited
by Boerlage,
named afterwards Schuurmansia theophrasta
346),
a
regarded by
species
me
Hall. /, Rec. Trav. Bot. Neerl.
80.
Type
—
Inflorescences
8o°.
terminal,
Flowers bisexual. Petals
One
species
Schuurmansiella
345, t.
/,
fl.
Sarawak,
to
up
m
7
fr.
&
157;
base,
long;
12
chartaceous,
rachis
mm
c.
1
32
—
4-è by
if—2J
00,
flattened filaments,
Ovary
ovoid,
ellipsoid,
c.
8
by
subtrigonous
at
much
branch conferted.
one
Seeds
parts.
3
base.
of
angle
an
branches
racemes;
of
21
2,
Hall. /
ascending
connate at
(1921) 387.
Ann.
of
co,
Ovary 3-carpelled;
winged.
not
Rec. Trav. Bot. Neerl.
—
(1868)
7
10
(1913)
Schuurmansia angustifolia Hook.
22c.
to
12
by
or
25
1
c.
—Type: Lobbs.n. (Kholo),
nerves
acicular,
\—1
up
5
linear,
mm.
at
base,
mm
by
—
style
in cross-section,
c. 0.3
mm
8| by
c.
long, purplish;
3
mm
±
long
tapering
15
c.
mm.
Petals
distinctly reflexed.
long, purplish.
long, purplish; anthers lanceolate, i-|
mm;
—io
cm
up
long under branches, smaller
—ij by f—1
2—3
5
apex,
Inflorescences
apart.
I
pinkish, purplish
—30,
at
flowers; pedicels filiform,
mm
linguiform,
±
acuminate
mm
to
2—3
Leaves with
mm.
by f—cm, long
Sepals
ones
mm
0.7
up
—17
branches with
white
inner
|—I
1.2
Walp.,
bracts
erect.
mm,
small;
Schum.
K.
P., Nat. Pfl. Fam. ed.
&
small, those
stamens
into
86
serrulate,
margin
long when fruiting;
staminodes
was
1913,
1857.
mm across;
base of pedicels. Flowers
10,
Borneo.
Soc.
high. Stipules
petiole; lamina linear-oblong,
at
henningsii
(Hook, /)
Nerves
(Hook./) Hall./,
R. As.
Br.
Ambon,
Néerl.
Bot.
surpassing the sepals in size. Staminodes
splitting
in NW.
(i860)
Trans. Linn. Soc. 23
Shrub
in E.
compound
bracts
age;
soon
angustifolia
Merr., J. Str.
7;
Borneo,
fruits also
Style of ripe
I.
Distribution:
1.
in anthesis
mutually and with the
types, the inner ones
stigma
Gilg
344;
many-flowered,
flowers of successive
shortened, with
2
(1913)
10
Schuurmansiella angustifolia
species:
from
s.ti.
Trav.
SCHUURMANSIELLA
slender branchlets. Stipules acicular.
Shrubs with
c.
De Fretes
(Ree.
with Schuurmansia
conspecific
as
9.
(1925)
Hall. ƒ
—
i
by
at
ovate,
Outer
Stamens with
0.3 —0.6
stigma capitate.
Seeds O.J
to
0.2—0.5
mm.
Fruit
mm,
tomentose.
Distribution: NW.
BORNEO.
G.
Sarawak,
Matang:
(K);
Bidi:
Clemens
1st
Borneo.
Div.:
22352
(K),
Beccari P.B.
Collenette
Ridley 11765 (BM, SING);
S.
1606
(FI, K, L, P),
697 (L),
Haviland
Sarawak:
Lobb
s.n.
Haviland 33
s.tt.
(K),
Native
(K, SING);
coll.
2580
(A, L, P);
Smythies
S.•
15332
Tg. Sipang: Brunig
S.
12058
(SING); Tegora
Mine:
BLUMEA
74
8. Distributional
Fig.
from
Santubong:
G.
JC
■1371 (L),
S.
Brunig
Anderson
Telok Asam:
Park,
Enoch
of Schuurmansia.
species
8368 (K, L), Bujang
Muas
Abang
Kanis B
(K, L),
3139
1968
I,
Outlined
(L), Purseglove
li
kerangas forest,
areas
are
only slightly generalised
soils,
on
Mus. Bot.
Ind. Bat.
(1871)
147;
345;
1,
Baill.,
66;
Lugd.
(1859)
2
Bat.
117;
Pi.
Hist.
Bartell., Malpighia
Gilg
(1941)
in E. & P.,
524;
B.
15
1
&
reported from
(1850)
177,
t.
H., Gen. Pi.
(1901)
Nat. Pfl.
Bt. Gondol:
Hose
Irekan:
sandstone, and
(1873)
4
Mt.
Carrick
(K, L),
17912
1
to
up
cliffs
on
(K, L);
21431
(5)
Anderson
Bako
C.arrick
(K),
530
12486 (A,
K,
&
L),
(BM, K, L, SING).
600
altitude, especially
m
near
the
sea.
SCHUURMANSIA
10.
Blume,
Othman b. Haroti S.
(K),
(K, L, SING);
4911
lowland species,
a
on poor
12990
Ashton S.
7690 (K, L, SING); 4th Div., Bg. Baram,
Ecology: Probably
in
No.
XVI,
cited localities.
(K, L);
Nat.
of the
map
VOL.
Kanis, Nova Guinea, Bot. 6
2,
in E.
(1925)
21
Ann.
120;
&
P.,
2
(1852) 68; Miq., Fl.
Miq., 111. Fl.
Nat.
(1961) 63.
—
79; A. C.
Arch.
Pfl. Fam.
Rec. Trav. Bot. Neerl.
Hall./,
Fam. ed.
Walp.,
(1862)
Gilg
355;
171;
32;
1
10
Smith, J.
6
3,
Ind.
1
(1895)
(1913)
340,
Arn. Arb.
22
Type species: Schuurmansia elegans
Blume.
Trees
at
or
of 60—70°.
at
treelets with
stout,
often hollow branches.
intervals; glandular along the margins,
base;
Inflorescences
peduncle
Flowers bisexual
and with the
or
polygamous.
connate at
with
3
longitudinal slits,
with
2
blades.
Distribution: Three
Guinea, the Bismarck
Stipules
the
species
distinctly
Staminodes
base.
style
ribbed,
co,
I
often
or
2
often with
with
con-
usually
not
and the
many
splitting.
Seeds
Philippines,
Solomon Is.
I
or
winged
Celebes,
at
an
angle
hypsophylls
recaulescences.
or
whorls, the inner
Ovary 3-carpelled; stigmas
in Borneo, the
Archipelago,
in
entire. Leaves conferted
almost straightly ascending
terminal, many-flowered panicles,
and branches
stamens
nerves
ones
mutually
3. Fruit
like
a
opening
propeller
the Moluccas,
New
A.
Ecology
Pioneer
:
Kanis:
secondary
long
Style
as
I.
Style
half
2.
Filaments
;L
2.
Filaments
as
the
twice
long
Ann.
anthers
Capit., Bull.
29;
Fedde,
Rep.
S.
K),
(1951)
54
Guinea, Bot.
New
Braang,
Treelet
long
nerves
I
—1|-
lamina
the
3—6
bracts
long,
mm
Flowers bisexual,
by
15 —25,
long
2—5
linear
subovoid,
capitate
with
c.
or
3
mm
io—2j
fl.
m,
lanceolate,
vidalii
Walp.,
32;
(1871) 66,
I
(1913)
10
Heine in
68;
Kloss
1912
Nova
fl.
(BM holo,
s.n.
—'13.
(K hoio), Sarawak,
500
up
to
mm,
\
mm,
papillate,
long, growing
in
fusiform,
up
Fruit
linear,
eva,
long, with
c.
-
„
1
long,
by
I—mm
distinct
connective
2\ by \
cm,
striation parallel
mm.
shortly ciliate.
Petals
obovate,
,
inner staminodes
long;
Stamens with
nerve.
i
a
few
into the
mm
—2
distinctly protruding.
sometimes with
fruit, widening
to
2
involute,
pedicels filiform,
;
sometimes
i|—4
-
cm
rounded
cm,
somewhat
0
mm
Stipules
ï\ —6
finely reticulate by
very
impression of
2—5
3 —6
,
o—
mm
an
mm
2
obovate,
Leaves with
base, margins
at
long; peduncle
to
low stilt-roots.
by z\ —10
10—30
surface of dried leaves
cm
by
ij—2
11—2
mm
punctiform.
Distribution:
t.
Ind.
(1923)
3
Type:
—
45
shortly ciliate.
epidermis cells, giving
Outer staminodes
by
i|—3
18.
sometimes with
slightly tapering
apex,
Sepals elliptic
anthers
ig—3
hairs; style
high,
to
chartaceous;
spatulate, 2-|—5
to
filaments;
S.
(1953) 63; Kanis,
Type: Haviland
—
178,
Arch.
(L holo), Amboina,
s.n.
(1916)
,
mm.
(1850)
I
Philip. Fl. Pi.
En.
sometimes
broadly linguiform,
erect.
,
4 —8
at
Inflorescences
nerves.
77.
m
mm,
walls of
protruding intercellular
to
(1930)
elegans
henningsii
S.
Ree. Trav. Bot. Néerl.
Hall./!,
397;
19;
Bat.
118; 111. Fl.
(1859)
Mt. Carstensz,
obovate-oblong
apart,
mm
Lugd.
Bot.
Type: Zippelius
to
S.
I.
2.
Samml. Clemens Kinabalu
15(—30?)
to
up
l|—4 by i|—2-J
petioles;
punctiform
or
3-lobed.
3.
1, 2
Bot.
—
Kew Bull.
somewhat" acuminate
or
Pfl.
ia.
capitate
fr.
m,
tree,
or
linguiform,
from natural
especially
m,
SPECIES
Trans. Linn. Soc. Bot.
9
Ridl.,
420
3000
75
anthers
(1910)
57
Guinea, Utakwa R.,
S. borneensis
Mt.
240;
to
stigma
Mus.
(1916)
11
(1961) 64, f.
6
parviflora Ridl.,
W.
Sc.
J.
level up
Area
Indo-Pacific
shorter
Ind. Bat.
Soc. Bot.Fr.
Merr., Philip.
346;
the
or
Blume,
(1852) 68; Miq., Fl.
2
of
length
the
the
THE
shorter;
or
ovary
the
as
TO
longer; stigma
or
ovary
as
Schuurtnansia elegans
I.
t.
the
as
long
as
sea
of
vegetation.
KEY
I.
the Ochnaceae
of
from
plants reported
anthropogenous
or
revision
A
Ovary
glandular capitate
acuminate. Seeds
small,
stigma
ovary;
i by
c.
Ï mm,
slender wings.
long,
Borneo,
the
Philippines
(Mindanao?),
the
Celebes,
New
Moluccas,
Guinea.
BORNEO.
Sarawak,
Mt. Kinabalu,
(A,
BO, K,
Kalangan:
L,
Celebes.
bb.
24127
SING),
Wae,
(A, BO, K,
G.
NEW
(BO,
de
L,
Halmahera,
Kornassi
798
(A,
Mindanao, Agusan:
L,P), Teijsmann
Rant
40423
G.
H.B.,
K,
GUINEA. W.
de
BM,
—
E.
K,
130
Gurulau:
L);
Borneo,
s.n.
L),
Carr
500
s.n.
S.F.
W. Kutai, L.
Cortez & Fernandez
Teijsmann
(SING),
BO, K,
—
Sabah,
Endert
W.
Penibukan:
3112
(PNH t ?),.fide
(L, U); Makale-Rantepao, Tg.
Coast
(A, BO,
Merrill
Kesu:
Res.,
Clemens 30479
Sandakan
27078 (SING);
Petah:
F.B. 24333
(K).
(SING);
Res.,
K,
Kg.
L).
(1923).
Thung
N.I.F.S.
SING).
G.
Ema,
Sembilan:
U);
Pleyte
Amboina:
G. Hori:
Wiljes-Hissink
New
Haviland
(A), 33063 (A,
iq6I (BO, L, U),
L);
Salahutu:
Vriese &
(BO, L,
322
Braang:
10309
3636 (K, L).
Minahassa:
MOLUCCAS.
Roho:
Clemens
Alabazo A
Philippines.
Div.,
ist
Kiau:
412
de Vriese &
Teijsmann
33
(A,
(BO, L); Ceram, Riring:
Binnendijk 18498 (BO),
Rutten
Robinson
2180
2036 (A,
(BO,
BM,
U);
L,
BO,
K,
Teijsmann s.n. (L), Zippel s.n. (L); Galala, G. Malintang:
s.n.
(BO, L);
Hutuinuri:
Teijsmann
s.n.
(BO,
K,
L,
U);
L).
Guinea, Vogelkop Peninsula,
SE.
of
Sorong: Versteegh
BW
4681
(L);
Klabala
BLUMEA
76
R.
waterfall:
near
Sleumer & Vink
Aet
BW
K,
706 (A, BO,
(BO); Japen
12709
(BM, K).
Royen
van
Kamundan
(L); Upper
bb. 22347
Guinea
clay
on
level land
on
or
Gulf
slopes,
identified
lopu;
wat
this
see
collection;
under the
2000
riverbanks
S.
in Ceron,
Herb.
Pi.
Cat.
f?), Luzon,
parvifolia Merr., Philip. J.
or
Stipules
tree?
obovate-lanceolate,
Cortez
Fernandez F.B.
&
Sc.
Manila
(1916)
n
by i\—2
(1916)
11
(1961) 69, f.
(1892)
mm.
Petals
Bot.
19;
mm
subglobular,
Fruit and seeds
Distribution:
PHILIPPINES.
Ramos
B.S.
Ecology:
Remarks:
Luzon,
23648
BM,
3,
Pulle,
&
Merrill
two
f.
R.
Isarog:
K, L,
(1923)
incorrectly
collections
henningsii
(1889)
75;
Guinea
Proc.
to
Sepals
long,
mm
f—1 by
glabrous;
chartaceous.
obovate,
c.
|
—2f
2
10
—
mm connate
0.4—0.5
stigma
bracts
long;
mm
Ovary
mm.
3-lobed,
subsessile.
P,
in
Vidal
2134
(A, K);
mossy
forest from
described the ovary
of this
identity
Sorsogon
Prov., Lake
Polog:
SING).
to
1000
from Luzon
species
m
upward.
contain
are
from Mindanao could also
1
erect
known
be this
ovule.
to
me.
species:
Blume.
8
K.
Soc.
K.
Schum.,
Warb.,
50;
Schum.
(1912)
P., Nat. Pfl. Fam. ed.
Francis,
anthers
lamina
tapering
at apex,
Outer staminodes
pink.
or
I—2
erect.
linear, i|—2
0,
mm
Prov., Mt.
of uncertain
elegans
(,1895)
Nova
25,
acuminate
Luzon).
BO, BRI,
to
only
K. Wilhelmsland
6
(SE.
Fernandez-Villar
3. Schuurmansia
Fam.
c.
white
long filaments;
0.4—0.5
Camarines Sur
(A,
According
under S.
mm,
bisexual,
fl. VIII-1915.
long petiole;
cm
pedicels
;
2134
23648 (A, BM,
apart, papyraceous
0
vidalii
Vidal
Type:
—
unknown.
Another collection
Fl.
mm
3-lobed,
Philippines
Unfortunately
see
if
c.
mm
mm
Flowers
3), by
c.
\ —1
i\—2
inner staminodes
long;
slightly
nerves
\—2
obtusely
to
Philip.
En.
Calophyllum
—
Ramos B.S.
Type:
Leaves with
obtuse
cm,
triangular.
to
obovate,
base. Stamens with
at
not
the remark
(see
fl. Ill— 1886.
Isarog,
—
ic.
plate.
+
229
Mt.
Bot. 19.
mm.
long; peduncle
cm
7—15
filiform, I—I
E.
by ij
.
c.
by \\ —4
5 —11
small, semi-annular
very
did
I
24353.
from the Philippines
6
Camarines Sur,
base, margins somewhat involute,
at
Inflorescences
labo,
hikselah (Tehid).
BO, BRI, K, L, P, PNH holo f?, SING), Luzon, Sorsogon, Lake Polog,
Treelet
localities,
swampy
species).
next
PNH holo
(A, K,
s.n.
sterile specimen from Agusan, Mindanao, which
a
elegans:
Fl. Pi. 3 (1923) 69; Kanis, Nova Guinea, Bot.
F.-Villar
R.: Kloss
6433 (L).
(1923); Celebes:
Penins.:
Schuurmansia vidalii (F.-Villar) Merr., Philip. J. Sc.
2.
Utakwa
Pullen
in
or
Merrill
fide
Vogelkop
would be the only record
it
Ihu:
R.,
Babo:
& Hatusima
altitude in primary and secondary
m
near
Guinea,
New
Schuurmansia cf.
as
to
up
Kanehira
4896
Misjnuk:
Mt.
(BO, L); Bomberai,
Bumi R.:
Mimika,
L);
of Vailala
BW
(L), Versteegh
rocky soils.
or
Remark: Merrill (1923) reported
was
BO, K,
estuary
6044
13153
Bay, Nabire,
Idjan 428 (A,
sometimes
BW
7643 (K, L); Anggi Lakes,
& Hatusima
Geelvink
Distr.,
1968
i,
Schram
Sleumer
Mindanao: tanang (Manobo),
names:
Amboina:
(Toradja);
Aet &
&
Kanehira
SING);
level
sea
steep
sandy
more
Vernacular
to
Waren:
(Papua),
from
Reported
forest,
Royen
van
L,
(BO,
Wasabori:
Serui,
I.,
SE. New
—
R.:
Ransiki,
(L);
14033
L),
No.
XVI,
(L); Teminabuan, Beriat:
32 og
Basin, Ije
R.
VOL.
2, 21
667;
&
Jahrb.
Jahrb.
Laut., Fl.
Hall, f.,
(1925) f.
Queensl.
Bot.
Bot.
41;
38 (1927)
9
(1888)
13
Deut.
Rec.
210;
K. Schum.
(1891) 283;
Schutzgeb.
Trav.
Lane
Poole,
247;
White,
Bot.
E.
J. Arn.
&
Hollr.,
Nat.
Siidsee (1901)
Neerl.
For. Res.
P.,
&
10
(1925)
Arb.
(1913)
Pfl.
448;
346;
116; White &
10
(1929)
241;
Kanis :
A.
Kanis, Nova Guinea, Bot.
S. bamleri K. Schum.
Trav.
Néerl.
Bot.
Guinea, Sattelberg,
S. bamleri K.
200—400
&
fl.
m,
Laut.
longifolia Gilg
Arb.
500
22
(1941)
fl.
m,
S.
319,
in
E.
524.
Area
holo
(B
216
holo
(B
f. BRSL),
longifolia
var.
Lain,
Schum.
in K.
gilgiana
Laut. in K. Schum.
Ree. Trav. Bot. Néerl.
NE.
Laut., Fl.
&
(1925)
21
2,
80;
(1913)
10
61
(1923)
fl.
1885—86.
S.
fr.
Suppl.
4.
213.
IX-1876.
s.n.
Südsee Nachtr.
Jahrb.
n,
12,
Baker, J.
17;
Obree Ra.,
Mt.
(1913) 346;
(BO, K,
Holth.
1857—61.
(1913)
10
347.
&
far.
Hutumuri,
sterile.
Lectotype: de Vriese
—
de
Paratype:
—
fl.
Lam, Blumea
&
nolo), Ambon,
L
(BO, CAL, L, U), Ambon,
5524
Bot.
region,
Teijsmann
Vriese &
s.n.
211.
Ree. Trav.
Néerl.
Bot.
(1913)
10
349.
S.
—
sp.
Type: Forbes 677 (BM, K, L holo), Papua,
—
Schum.,
K.
Sogeri region,
1885—86.
?
tophiroides Gilg
S.
indicated
oreophila
? S.
Gilg
indicated (B f?).
Mts.,
?
500
BO,
E. &
in
or
at
Mt.
Smith, J.
A. C.
tree,
lamina
Smith, J.
Owen
Schlechter
ed.
2,
Fam. ed.
2,
Fam.
21
21
(1925)
80.
(1923)
80.
(1925)
80.
Kaiser
16639 (A, K, L),
&
Nat. Pfl.
P.,
up
—3
to
long;
Tafa,
Fam. ed.
Arn. Arb.
fl.
m,
—
Type:
not
—
Type:
not
—
Type:
Wilhelmsland,
not
Kani
21
2,
(1925)
80.
—
Type:
not
2840
Arn. Arb.
22
high,
m
often up
to
3
(1941)
7 —65
usually
very
or
Type: Brass 5076 (A holo,
326.
m,
—
Type:
Brass
fl. 6-IX-1933.
—
4743
(A holo,
Paratype:
Brass
long ciliate.
\\—15
cm,
nerves
long, peduncle
small,
±
—
Type:
sometimes with
mm
6—85 by
cm
527.
Lane Poole 371
(A holo,
fl. II-1923.
m,
margins somewhat involute,
bracts
—
fr. 6-IX-1933.
I5(—20?)
leaves. Flowers polygamous
525.
5-VIII-1933.
(1941)
22
1800
mm,
(1941)
22
2400
Murray Pass,
obovate-lanceolate,
base,
Arb.
Arn.
Stanley Ra.,
subcoriaceous. Inflorescences
mm
Pfl.
Nat. Pfl.
P.,
BO, BRI), idem,
Stipules | —5 by
2 —5
E.
Wharton Ra.,
K), Papua,
tapering
Nat.
P.,
&
Smith, J.
A. C.
grandiflora
petiole;
in
Gilg
A. C.
montana
Treelet
E.
in
BRI), Papua,
4706 (A, BM,
S.
21
2,
(B f?).
BRI), Papua,
BRI,
Fam. ed.
fl. 5-X-1907.
m,
S. coriacea
S.
Pfl.
Nat.
P.,
Specimen:
—
S. crassinervia
indicated
BM,
&
(B f?).
schlechteri Gilg
? S.
in E.
(B f?).
indicated
to
t.
Yule, fr.
10
Rec. Trav. Bot. Neerl.
Hall./,
(1888)
9
IV-1902.
398,
(BO), Papua,
291
(1905)
(Bholof,
.
sterile, 1839—60.
rauwolfioides
S.
fl.
m,
Mt.
Néerl.
H.B.
Teijsmann
(L holo), Ternate, sterile
47
(L), Halmaheira,
Bot.
—
Arn.
Ireland, Punam,
Schlechter 14583
Type:
—
(1910)
MEL
ex
s.n.
Paratype: de Fretes H.B.
—
ioo
57
(BO), Papua,
Ree. Trav. Bot.
,
Lectotype:
—
Bot. Fr.
Sayer
MEL
ex
pseudopalma Hall./,
S.
Deut.
351.
Lectotype: Forbes 613 (BM, BO nolo, L), Papua, Sogeri
—
theophrasta Hall. f.
H.L.B.
fl.
Bull. Soc.
Paratypes:
—
Macgregor
(1942)
5
New
Smith, J.
A. C.
New
Schutzgeb.
Deut.
(1913) 348.
10
Torricelli Mts.,
New Guinea,
microcarpa Capit.,
1889; ?
NE.
VII-1902.
Hall./,
K),
S.
NE.
Laut., Fl.
&
Ree. Trav. Bot. Néerl.
„
BO,
f, K),
Südsee (1901) 448; Hall./, Ree.
20
Type: Schlechter 14646 (B holo f, BRSL),
—
77
18-XII-1898.
P., Nat. Pfl. Fam. ed.
&
Indo-Pacific
VIl-1886.
Type: Bamler
—
350.
Schutzgeb. Siidsee Nachtr. (1905) 318; Hall./,
S
fl.
Laut., Fl. Deut. Schutzgeb.
&
Schum.
the
of
Type: Hollrung
—
Sattelberg, Kakulu,
(1913)
10
Ochnaceae
(1961) 65, £ ib.
6
Guinea, Finschhafen,
New
of the
revision
A
triangular,
stiltroots,
Leaves
obtuse
2—7
nun
i|—8
mm
sometimes
functionally unisexual,
erect.
up
with up
to
to
1
;
m
4 cm
acuminate
apait,
0
to
at
high.
long
apex,
chartaceous
or
pedicels filiform,
larger and transitional
Sepals
obovate
to
elliptic,
BLUMEA
78
by
3 —5
i\—4
c.
i
white,
in (J flowers with
with
I—I
f—2\
filaments; anthers
long
mm
slightly 3-lobed, glabrous,
style cylindric,
Distribution:
c.
1
Foramadiahi:
Beguin
with
by
I
|
G. Duata:
K,
(BO,
L,
Lam
in
subglobular
flowers
$
fusiform,
Fruit
c.
Stamens
$ flowers
ovoid,
to
by
3
2 mm;
i\ by f
to
up
filiform,
cm,
Archipelago, Solomon Islands.
Halmahera:
2946 (BO, L);
Temate:
U);
Ovary
mm,
—
in
mm;
by
4
oo,
long.
mm
i\—2\ by | —£
mm.
o—
long, slender wings.
mm
Guinea, Bismarck
New
1519
c.
staminodes
linear, i| —2
to
|—I by |—§
2.\
c.
Petals obovate-oblong,
Outer
anthers
long; stigma 3-lobed.
mm
5
Moluccas,
flowers
(J
mm
by
1
in
Talaud, Karakelong,
MOLUCCAS.
(L);
to
up
acuminate. Seeds
filiform
5 —30,
long filaments;
mm
1968
i,
red.
purplish
or
No.
XVI,
purplish.
sometimes
pink,
creamy,
inner staminodes
long;
mm
greenish,
mm,
—3
mm,
VOL.
H.B.■
Teijsmann
de Vriese
5169
&
Teijsmann
de
(BO, CAL, L),
s.tt.
Vriese
_
H.L.B.
Tidore:
(L);
47
H.B.
Fretes
L); Wae,'
"
H.B.
(BO, K,
291
NEW GUINEA. W.
BW
Versteegh
BW
Vink
Vink B W
Brass
Camp:
Ormu:
(Sterren)
NE.
Say ers
Brass
(K,
__,
:
L);
(L);
van
(K, L);
E.
(K, L);
of Mt.
NE.
Pullen 5432
of Mt.
Michael:
Brass
958i
Dunstone
(BO,
Lane Poole
Joangey:
216
(K);
(A,
BO, K,
Bamler
Malolo:
L);
20
Clemens
Selileo:
Mt.
R. :
(K, L).
142
Brass
Ne: Kalkman
7153
4871
(K, L); Anga Valley,
(K, L);
Mt.
R.:
Yule:
(A, BM, BO, BRI);
Boridi:
14947
Can
(A,
K,
13572,
L,
Sayer
s.n.
Mts.,
139,
Hartley
Mt.
—
(BM,
SE.
(L);
Lake
Ebenda:
1340
New
Floyd
Kutub,
s.n.
Neneba:
?
613
(Papua),
Imu:
(BO);
11337
R.
near
SING);
K, L,
of Mt. Wilhelm:
E.
Asaro-Mairi
Mafulu:
Isuarava:
16472 (K, L);
Corner &
Gray
N.G.F.
Divide,
Brass
30964
L); Okapa:
White
_
s.n.
20057
(A);
3130
Distr.,
2232
Central
Brass
5291
Distr.,
Can
15382,
15750
Bisiatabu:
N.G.F. 12909
(A,
Hartley
(K, L);
Valley,
(K),
13340
?Andarora
junction
2035
4786
Schodde
(K, L);
: Darbyshire
Maipa:
Mt. Tafa:
K, L,
T.
5458
Ooinsis:
Kiburu:
Schodde
L);
Brass
5076
13789 (A,K,L, SING);
10688
C.
N.G.F.
R. below
Akaifu R.,
Can
(K,
Kerewa: Kalkman
(A, BO, BRI);
4738,
Hollrung
of Lae,
N.G.F.
Palmer
Mt.
Giluwe, Klareg:
(K); Gap region:
Owen's Corner:
W.
(A.BRI.K);
Mendi
(K, L);
16112
Kakulu:
L, SING),
K,
Womersley
N.G.F.
Clemens
L); Mongi Valley,
K,
N.G.F.
(BM, K);
Sassaura:
(L);
Clemens 3371 (A);
Royen
(A, BO,
Western
W. of Mt.
s.n.
1035
Ogeramnang:
Highlands Distr.,
Schodde
Giulianetti
van
Creek:
Womersley
Southern
Smith
Warburg
Karnbach
(BM, BO, L), 677 (BM, K, L);
Woitape:
(A, BO,
K,
Hoogland 9560 (BM,
Wau, Edie
1623 (K, L);
Scratchley,
Forbes
N.G.F.
of Mt. Otto:
, . (BO,
11428
Sarawaket,
N.G.F. 5457
(A, BRI, K, P, SING);
Can
S.
Kaulo:
Butemu:
(BM, BO, K, L);
(A); Yunzaing &Joangey:
L); Hopi:
(L);
Guinea
14798 (A, K, L, SING);
(BO);
(A,
435
10447
IMacgregor
Piati:
6128
(L);
Star
(13M,
4338
Mts.
Womersley
%
Sambui:
Ra.,
K, L); Morobe:
Schodde
N.G.F.
Mt.
Mannasat:
Rawlinson
(A, BO, BRI, L);
Brass
SING);
L); Sogeri region:
Obree :
Hill:
•
Blockwood
Black
Red
Kalkman
Nondugl, Warranga
(L);
mouth
968 (BO, L);
Madang Distr.,
& Pulleti
908
towards
(A);
143
R.:
Bernhard
(K, L);
Finisterre
L);
N.G.F.
. 23721
Aiyura:
Clemens 3905
Mararuo:
(A);
4411
R.:
5356 (BM, BO, K, L);
13165 (L);
Hellwig 556 (BO, K); Simbang:
Sleumer N.G.F.
13949
Gulf Distr., Murua
1015
Cromwell
(L);
K,
(L);
255
M
Peninsula,
Huon
Clemens
(BRSL),
(A, K, L);
29335
Womersley &
1422
(BRI);
Rdmer
von
Merimanta:
Saunders
,,
(A); Yunzaing:
Hoogland 8856 (K, L), 8954
Sattelberg:
(L);
580
,
(A,
3661
Sigafoos
4623
Otken
Bivouac,
&
Tributary
-
Hartley
Distr.,
Clemens 41199
(L),
439
Sleumer
& al. N.G.F. 5159
Sayers
(K, L), Womersley
v
-
Wanatabi:
-
L);
Morobe
(L);
12053
4842 (A); Matap:
Pindiu :
K,
-
Pullen
Womersley
& Pullen
&
14583 (BO, K);
17490
Wabag,
Millar &
Pullen
(BO),
31197
..
N.G.F.
village:
R.:
Kerigomna: Hoogland
—
with
A1
(K, L);
18272
of
R.,
(BO, L); Idenburg R.,
(L),
980
Hagen: Hoogland
Divide:
(L); Minj-Nona
423
N.G.F.
Camp : Hoogland &
L);
Brass
of Mt.
S.
834
Bon R.—Minam
16639,
N.
Highlands Distr.,
2028
Rouffaer
Pionier
Valley,
of Simboro:
N.
Schlechter
Schlechter
Versteegh
Son:
Kg.
Sleumer
Sleumer &
Wasior: Koster
Bay,
I.,
(L);
'"97
Royen
van
Lake,
Mts.: Pulle
Mts.:
Torricelli
BW'
Gjellerup
(BM, L); junction
Kani Mts.:
Western
Robins
Royen
Sentani
Biak
Valley:
&
Gwainongga:
Geelvink
(FI);
s.n.
Roven
van
Lakes, Ginambarai-Djembodini: Eyma
Ifar:
Cycloop Mts.,
4293
de
Teijsmann
Kebar
(L);
4987
Tobi:
Mt.
12089 (BO);
Mamberamo
Gautier Mts.:
(BO, K, L); Hellwig
Kalkman
Flenley
Wissel
L);
K,
Bergman 22 (L);
Highlands Distr., Chimbu, Pengagl Creek:
30354
Daulo
21266
BW
Mt.
to
14164 (BO);
Aisau: Vink
(FI);
Hutumuri:
(BO, K, L).
274
Nertoi
Beccari
(Kapaor):
s.n.
(BO, L);
BRI,
(L);
& Hatusima
6007, 6138 (K, L);
(A, K, L);
75
Giluwe:
R.
661
3395
Valley:
Lufamunda:
of Mt.
Eastern
1
>
Warapuri
BW
Beccari
Guinea, Sepik Distr.,
16754
N.G.F.
(K, L);
x
New
—
Schlechter
Supadmo
Kanehira & Hatusima
10461 (BO,
& Sleumer
Sibil
Fakfak
Amboina:
(BO, L);
1227
"
&
BW 68
51
Kanehira
Kadubaka:
12766 (A, BO,
Mts.,
Kornassi
Boerlage 65 (BO, L),
~
Kuswata
Koster
Dalman:
Thomson
Kalkman
Andjai:
Ansus:
Valley,
L),
Royen
van
of Tami R.:
E.
K,
481 (BO,
Salahutu:
Peninsula,
Nabire,
Leeuwen
van
Swart
(A, BO, K, L);
Lam
of
82q6 (L); Japen I.,
Docters
Camp:
L).
S.
"*
~
from Momi:
Onin
1426j (L);
of
S.
10362 (L);
13760 (L);
BW
~
G.
G. Horiel:
(BO);
303
Guinea, Vogelkop Peninsula, Ajamaru: Versteegh
(K, L) ; Anggi Lakes,
6844
Rant
""
New
Wai Kahula:
3768 (BO, L); Ceram,
Lam
(BO, L, U),
5524
SING); Kagi:
(L),
White
Northern
Schodde
356 (BRI);
Distr.,
Carr
2961
Wharton
Mt.
Ra.,
A.
Murray
BRI,
land
Pass:
4429
(BO);
Hoogland
K,
(A,
Gumini
(K, L);
4706 (A,
Brass
Saiho:
K);
Misima
Kanis:
Milne
L);
N.
(A,
Plateau:
of Mt.
Kere
Paehena
1024
Ecology:
San
Mt.
White N.G.F.
Lollo:
Star Harbour:
from
Brass
Arch.:
(A,
371
Oi-ai:
Hoog-
Crutwell
(A, L); Agupon:
Louisiades
630
Macgregor
Ireland,
New
(K, L);
10851
in
or
was
localities,
in
habitats.
open
s.n.
Punam:
W.
New
Wabag:
opaga,
or
more
Kairuku:
engefukenge;
sa’iabura;
Malaita: abekweto.
kembusa;
Buna:
batsjevak
distinguished
vegetative
characters is
differences in
New Guinea
mainly
on
Biak
but it is
material, arranged according
decrease in leaf size, especially between
became
more
coriaceous and
of collections
from
in
larger
more
areas.
and
iooo
names
age
of the
are
no
in
greater part
the altitude.
a
regular
very
time the leaves
gaps
plants play
of species
by
by
same
op;
du’ugwau
variation
The
exercised
At the
There
distinctly petioled.
Exposition and
haiwinge
S. Isabel:
caused for the
m.
3000
or
rambuan;
karadewa; Mendi:
increasing altitude, showed
to
near
Amboina:
I.:
Andarora:
leaf-characters.
certainly
forests,
arebi; Budemu:
Aiyura:
(m)beli;
conditions. The greatest influence is
ecological
3018
popai, porkai, pupai; Minj:
Solomon Is.,
werawera;
differences
striking indeed,
Brass
Scattered
maletope;
(Maibrat);
Kutubu:
;
soils.
rocky
or
Remarks: In Schuurmansia henningsii K. Schum. I have united several
were
of
B.S.I.P.
sometimes
slopes,
Tidore:
urune;
Sattelberg:
Bausa:
SW.
Womersley
Hinuahaoro:
L);
steep
sandy
SE. New Guinea, Tari: obbo
(Nauti), yatsiga (Manki);
to
Hagen: pappai,
pelip;
(L);
primary and secondary
in
akesse, a’ulareh, hahessa, menmeh;
Sambui:
Peninsula,
m
arisusu
orpach;
K,
(A, BRI,
level land
Guinea, Ajamaru:
bubar, bubus; Chimbu: akessa,
Huon
Is.:
Whitmore &
Jonapau:
Kalena
Georgia,
B.S.I.P. 6810
L).
3000
on
clay
on
Moluccas, Talaud
names:
lapu;
to
up
grasslands,
swampy
New Guinea,
sipulund;
259
BRI,
(A,
Beer's coll.
R.:
Walker B.S.I.P
>.
3112
SE. New
Kajewski 1692 (A, BRI);
Kolokofa
Mt.
level
sea
Field:
Guadalcanal,
7756 (L);
Cristoval, Anganiwai:
common
lapu,
that
Poole
(BRI); Tufi,
237
22804
(K, L);
19309
Lane
Stanley Ra.:
Lane Poole
Brass
Dayman:
N.G.F.
Isabel,
NW. Santa
B.S.I.P.
■
Reported
Vernacular
NE.
Owen
79
27484 (A, L).
Britain,
landslides and in artificial
riverbanks
ut
(K, L);
5937
Beer's coll.
Pt.:
(K, L);
locally
of Mt.
Bougainville,Kupai Gold
ISLANDS.
B.S.I.P.■
(A, BO, BRI, L);
on
R.:
Area
Indo-Pacific
14646 (BRSL).
SOLOMON
Bay:
Brass
Sisa:
BISMARCK ARCHIPELAGO. New
Schlechter
N.
Womersley
the
of
(A, BRI);
4743
L); Hydrographers
K,
Bay Distr.,
R., Cameron
I.,
BO, BRI),
BM,
3307
the Ochnaceae
of
revision
A
a
in series made
less
important
role in the determination of the leaf size.
The dimensions and colours
of flower
are
parts
rather variable
correlate with other data. These variations might be expressions
between local
whether
any
populations.
infraspecific
Some authors
unisexual. In
their
a
described
previous
and has been described
as
as
is
in
certain,
likely
S.
a
(1961)
as
dioecious
I have
are
to
however,
occur,
crassinervia,
key
and
no
and the flowers
doubt
expressed
complete
and
as
as
the
to
'pistillate
Further examinations have convinced
male. A
as
S.
to
decide
(functionally)
correctness
flowers'
that
me
a
are
minority
has
relatively small
anthers
and
of
rare
majority
big
I have
pistils
female. This is in accordance with field observations by Dr. P.
anthers and did
that
the
flowers
not
are
the reduction of either
lophiroides,
types
to
be distinguished.
Royen, formerly in Lae, who fotmd that
stand had relatively big
not
easy
has relatively large anthers and small pistils, and consequently
described these above
van
detailed local studies would be necessary
the species
paper
herbarium materials.
of the specimens
can
because all flowers
observations,
among
More
taxa
and less
too
of genetical differences
were
S.
oreophila,
most
set
specimens in
fruit
always
stamens
and S.
a
profusely flowering
(personal communication).
functionally
or
ovary
schlechteri
is
unisexual.
never
were
I
am
Polygamy
complete.
published
by Gilg
indicated. The brief key hardly permits identification since
BLUMEA
80
it is
based
mainly
This
leaf-characters, but
on
supported
view is
VOL.
by
which
L, labelled S. schlechteri,
1968
I,
I consider all of them identical with S.
Berlin
some
No.
XVI,
duplicates of Schlechter 16639
undoubtedly
belong
S.
to
henningsii.
A, K, and
in
henningsii.
EXCLUDED
Bull.
Capusia Lecomte,
Merr.,
Arb.
Arn.
J.
Mus.
Hist.
Gomphia magnoliaefolia Zipp.
Ochna
Ann.
decaisnei
Sc.
ex
Nat.
Tiegh.,
v.
Bot. VIII,
Paris
(1926)
32
Linnaea
Span.,
(Menispermaceae), fide
Becc.
(Decne.)
Nat.
Diels
in
Siphonodon
=
(1902) 356.
8
nud.
nom.
Heft
(1902)
RiedU
Type:
—
186,
(1841)
15
Pflanzenreich
Bull. Mus. Hist. Nat. Paris
16
95
Griff.
( Celastraceae), fide
108.
(1940)
21
47—49.
51.
decaisnei
Diporidium
—
(P holo),
s.n.
Pycnarrhena longifolia
=
(1910)
46
Timor?,
1803
v.
Tiegh.,
O. mauritiana
=
Lamk.
The
must
description
have
been
Timor. It
to
2
have
must
Sauvagesia
Java
based
was
collected
Erroneously recorded
Later
authors
Miquel's
It is true
Indo-Pacific
collected
in
d'Histoire
Ges.
(1893)
ii
by
are
a
Fl. Ind.
Miq.,
(1963)
collection
The
(Mauritius)
specimen
on
the way
Bat.
118;
(1857)
2
1,
Koord.,
Exk.
Fl.
327.
reference
same
voyage.
drew
kindly
Bat.
of
material
to
that
the
my
of
other
or
of information.
source
Sauvagesia
concerned
534,
to
this
particular
is
year.
known
not
'Perrottet,
However,
otherwise
and in
mislabelled
was
Mr. C. Sastre
supported by
attention
which is labelled:
G,
in that
Java
and
specimen
This view is
Suppl. (i860)
erecta L. in
Sauvagesia
the eastern part
from that island
known
probable
very
the
Fl. Ind.
&
203;
1
France'
de
flower.
fruiting
one
'lie
on
on.
Java by Miquel without
caused
who
Miq.,
22,
Java
expedition
Miquel.
to
It is
on
Naturelle,
Tetramerista
Fl.
that Perrottet visited
area.
Cayenne
later
(1753)
1
Bakh./.,
of Ochnaceae
other collections
the
for
might be
error
Java, 1819'.
&
refer
only
PL
Linne, Sp.
erecta
Baudin's
mislabelled
been
607; Back.
(1912)
branchlet, originally bearing
one
on
during Capt.
no
from
fact
was
of the Museum National
collection.
belongs
the
to
Ber.
Theaceae, fide Gilg,
Deut. Bot.
div.
auct.
Index
New
species
names
or
in bold
are
Brackenridgea
Gray
A.
sect.
Brackenridgea
sect.
Notochnella
F.
australiana
Furt.
denticulata
ssp.
ssp.
fascicularis
foxworthyi (Elm.)
hookeri
var.
kingii
nitida
Tiegh.
v.
A.
Gray
2:
australiana
ssp.
nitida
(Merr.)
2:
2:
Tiegh.
Tiegh.
2:
5a
v.
australiana
kingii
2:
2:
3c
2:
3b
(F.
2
3a
3a
v.
v.
retinerve
rheedei
strictum
v.
v.
v.
3:
Tiegh.
3:
3:
1
1
1
v.
Tiegh.
3:
1
Tiegh.
1
3:
3:
1
1
1
Tiegh.
3:
v.
3:
3:
1
Tiegh.
Tiegh.
1
3:
Tiegh.
1
Tiegh.
5a
1
Tiegh.
3:
3:
(DC.)
2:
3
Tiegh.
v.
v.
Tiegh.
3:
v.
v.
Tiegh.
v.
perakense
v.
Tiegh.
Tiegh.
v.
1
Tiegh.
Tiegh.
1
2B
M.)
v.
3:
1
1
3:
(Bartell.)
v.
nodosum
3:
Tiegh.
v.
1
Tiegh.
v.
3:
Tiegh.
2
10:
3
3:
Tiegh.
(Bartell.)
v.
plicatum
and
1
3:
Tiegh.
v.
malabaricum
2:
2:
Tiegh.
striatum
leschenaultii
Kanis
Bartell.
Kanis
Kanis
Tiegh.
v.
beccarianum
a
3
3a
serrulata
3
M.)
palustris
v.
v.
angustifolium (Vahl)
2
of genus
vidalii F.-Vill.
v.
abbreviatum
2:
number
paucifolium
cumingii
v.
kjellbergii
v.
ib
2:
5
(F.
to
Tiegh.
v.
borneense
ssp.
rubescens
Kanis
3b
Gray
foxworthyi (Elm.)
perakensis
hohenackeri
zollingeri
1
3a
2:
2:
refer
end.
Campylospermum
ssp.
ssp.
6
2:
2:
5b
2:
Bartell.
palustris
the
Campylopora
Scheflf.
2:
at
neriifolium (Bartell.)
3a
4
A.
ssp.
2:
Kanis
Furt.
(Planch.)
leucocarpa
2:
listed
metzii
ia
2:
Tiegh.
v.
2A
5a
F.-Vill.
mindanaensis
forbesii
Kanis
Tiegh.)
2
(Blanco)
italics. Numbers
Calophyllum
Tiegh.
v.
in
names
Campylocercum
2:
elegantissima (Wall.)
fascicularis
excluded
2
(v.
2:
to
2B
M.
v.
(King)
corymbosa
type, synonyms
'Excl.' refers
variety.
3:
1
1
accepted
vahlianum
v.
walkeri
Tiegh.
zeylanicum
annamensis
brevis
doumerii
v.
elongata
thorelii
wightii
v.
3:
3:
3:
1
3:
1
Diporidium Wendl./.
decaisnei
(v. Tiegh.)
(Blanco)
poulocondorense
pumilum (DC.)
(L.)
squarrosum
walkeri
1:
Excl.
2:
ia
3c
2
1:
2
Tiegh.
1:
1:
O. Kuntze
Kuntze
1:
4
O.
Kuntze
1:
4
normale O. Kuntze
dalzellii
Tiegh.
v.
gaudichaudii
harmandii
koenigii
Tiegh.
(Lamk.)
obtusatum
wallichii
1:
(Planch.)
1:
non
6:
Gagnep.
Elaeocarpus integerrimus
Euthemis
ciliata
Jack
engleri Gilg
4:
Hook.
var.
robusta
Gomphia
sect.
2:
4:
4:
Vahl
v.
4
4:
Planch.
Thw.
v.
6
2:
4:
3:
M.
var.
Planch.
2:
3
var.
5a
var.
3
1
b
4
3a
Kurz
ex
1:
Excl.
2:
1:
4
Brandis
1:
1:
3b
3a
4
(v. Tiegh.)
annamensis
cambodiana
1:
ia
3b
2:
Moon
Lecomte
1
3
3c
Tiegh.
Ridl.
ia
4
Elm.
harmandii
3
1:
foxworthyi
grandis
1
typus
O. Kuntze 3:
1:
Blanco
grandijlora
4:2
2:
4
1:
fascicularis
gamblei King
/.
1:
1:
1:
Tiegh.
1
2A
v.
excl.
(Vahl)
Griff.
7:
1
p.p.
Edgew.
Quis.
&
1
7:
Gamble
fruticulosa
1
2
1
7:
angustifolia
decaisnei
1
Merr.
Kurz
beddomei
1
7:
7:
p.p.
Linne
1
7:
1
7:
andamanica
cordata
1
Gomphiastrum
F.
6
4:1
Benn.
Schreb.
australiana
1:
2
4:
/
Hook./.
angustifolia
1
Lour.
Ochna
crocea
Hook.
pulcherrima
4
6
1
7:
1
(Blanco)
Linne
collina
latifolia Miq.
Jack
obtusifolia
1:
Steud.
2
leucocarpa Jack
minor
Ochna
2
1
7:
Shaw
Ridl.
Tiegh.
v.
Shaw
1
Airy
7:
Ridl.
Notochnella
syn.
1
Shaw
Korth.
brevipes
Diels
hackenbergii
1:
2
4:
Wall.
elegantissima
3a
syn.
1
7:
7:
Ridl.
fascicularis
4
Pearson
jackiana
1:
2
excl.
1
Airy
/.
Ridl.
philippinensis
quoad
p.p.
1
(Ridl.)
Airy
serrata
1
7:
angustifolia
minor
1
1
7:
7:
Ridl.
f.
6:
1
7:
Hook.
major
2
1
Adans.
borneensis
parviflora
3
8:
3:
Hook./.
f.
Vidal
Adans. p.p.
Ridl.
major
2:
1
7
Ridl.
1:3a
Tiegh.
v.
Distephania Gagnep.
Gaertn.
2
1:
Tiegh.
v.
f.
3:
3
ovalifolia Capit.
3a
Tiegh.
v.
ex
obovata
Tiegh.
v.
(L.)
involucrata
4
2
Tiegh.
v.
3a
Tiegh.
v.
(DC.)
squarrosum
1:
1:
1
Benn.
ex
Boerl. 8:
ex
var.
1:
Tiegh. 1:
v.
(DC.)
planchonii
3a
Tiegh.
v.
microphyllum
nitidum
1:
Tiegh.
1:
1
3:
DC.
Bartell.
Plum,
malayana
1
1
8
Plum,
lancifolia
4
O. Kuntze
A.)
Tiegh.
v.
v.
v.
leschenaultii
lucidum
&
1:
Excl.
1
/.
Boerl.
Ridl.
3a
Span.
Kanis
3:
Jabotapita
var.
2
1
3:
Jabotapita
klossii
2:
1
(Gagnep.)
Korth.
typus
6
Vidal
humilis
4
O.
Tiegh.
Hook.
distans
longifolium
v.
umbellata
grandifolia
1
brevifolium
wightianum (W.
Jack
2:
ex
3:
(Gaertn.)
sumatrana
Neckia
var.
Discladium
Ridl.
serrata
1:
var.
var.
Ridl.
Meesia Gaertn.
4
1:
Tiegh.
O. Kuntze
(Planch.)
microphylla
3a
excl.
2
3:
oblongifolia
calophylla
3a
Tiegh.
v.
v.
DC.
2:
p.p.
corymbosa King
beccariana
4
1:
O. Kuntze
v.
(Planch.)
wallichii
1:
O. Kuntze
rufescens (Thw.)
2:
81
p.p.
1
Indovethia
Tiegh.
v.
Planch.
involucrata
Tiegh.
v.
Ridl.
Ridl.
Indosinia
1
Area
corymbosa (King)
zeylanica (Lamk.)
Kuntze
brevipes (Planch.)
Indo-Pacific
corymbosa (King)
serrata
1
Tiegh.
(Thw.)
1
1
Tiegh.
the
of
malabarica
1
3:
1
v.
fasciculare
Tiegh.
3:
Tiegh.
Ochnaceae
magnoliaefolia Zipp.
1
3
Tiegh.
the
var.
v.
3:
of
hookeri
1
1
3:
v.
cordatum
1
1
3:
Tiegh.
v.
revision
Excl.
Tiegh.
v.
3:
(Lamk.)
Tiegh.
v.
Tiegh.
3:
Tiegh. 3:
v.
Lecomte
Cercinia
v.
Tiegh.
wallichianum
Capusia
A
Tiegh.
v.
v.
Kanis :
(Jack)
sumatranum
thwaitesii
A.
Lecomte
Lecomte
(v.
1:
1:
4
4
Tiegh.)
4
latifolia (v. Tiegh.)
Lecomte
1:
4
Linn.
1
Linn.
3
VOL.
BLUMEA
82
Lecomte
retusa
var.
W.
heyneana
&
A.
(Planch.)
humilis
Buch.-Ham.
Spreng.
Lamk. p.p.
Lamk.
lucida
(non Lamk.)
moonii Thw.
Thw.
/?
var.
nana
1:
(non Swartz)
{non Swartz)
1:
DC.
DC.
pumila
1:
Kanis
cordata
1:
rufescens
Benn.
Kurz
Rottb.
O.
1:
Planch.
1:
wightiana
W.
var.
moonii
var.
moonii
Ouratea
& A.:
{non
Benn.
3:
Trim.
3:
Bartell.
3:
(Griff)
ex
3:
1
1
3:
var.
v.
typus 3
:
Craib
3:
parviflora
schlechteri
1
10:
vidalii
3:
3:
3:
3:
Sinia
Alst. 3:
Walkera
1
serrata
v.
Tiegh.
1:
3a
3
10:
3
3
10:
3
3
3
10:
3
3
10:
10:
1
2
10:
/
10:
3
10:
3
3
/
10:
3
Merr.
Hall,
f
10:
2
9
Hall.
/
5
Tetramerista
1
1
gaudichaudii (v. Tiegh.)
Diels
rhodoleuca
1
3:
10:
10:
10:
Hall.
Diels
Miq.
Schreb.
5:
1
Excl.
3
(Gaertn.)
3
3
3
angustifolia (Hook./.)
1
1
2
1
10:
(F.-Vill.)
10:
10:
Smith
Hall.
Schuurmansiella
1
1:
3a
Smith
Ridl.
Gilg
1:
3c
1
10:
C.
1:
3
Sch.
Merr.
theophrasta
ib
Lecomte
Tiegh.
1
parvifolia
2
Lecomte
Gilg
zeylanica (Lamk.)
A.
Tiegh.
9:1
10:
C.
K.
4
Laut.
10:
A.
rauwolfioides
Robs.
(v. Tiegh.)
Pleopetalum
Laut.
oreophila Gilg
4
1
3:
(v. Tiegh.)
nervosa
Blume
grandiflora
/.
10:
Gilg
1:
3c
1
pseudopalma Hall./.
2:
(Jack)
3
1:
Smith
elegans
1:
4
longifolia (Laut.) Gilg
1
3:
1
mindanaensis Merr.
thorelii
Laness.
excl.
Craib
Bartell.
:
2:
microphylla (Ridl.)
striata
C.
gilgiana
montana
Burk.
megacarpa Ridl.
sumatrana
A.
coriacea
microcarpa Capit.
Burk. p.p.
(Gaertn.)
1
henningsii
1
lobopetala Gagnep.
serrata
3
Gilg
3
1
Burk. p.p.
(Planch.)
neriifolia
1:
lophiroides Gilg
Bartell.
crocea
Gilg
Reticulatae
Engl.
Baill.
borneensis
(Griff.)
3
Reticulatae
subsect.
beccariana
crocea
typus
1
3:
Ridl.
borneensis
3a
excl.
Tiegh.
v.
v.
Laut.
longifolia
4
3a
1:
Excl.
&
Sch.
crassinervia
Reticulatae"
Craib
1:
p.p.
1
sect.
angustifolia (Vahl)
hookeri
var.
Trim. p.p.
K.
bamleri
4
1
Baill.
Aubl.)
1:
1:
10
Hook.
angustifolia
4
Tiegh.
v.
Tiegh.
7:
Bl.
3a
4
1:
L.
jaheriana Capit.
1
Schuurmansia
(Thw.)
Palaeouratea
"Gruppe
3:
4
4
1:
Tiegh.
erecta
4
4
v.
Tiegh.
v.
1:
Tiegh.
v.
(Thw.)
Sauvagesia
3a
4
(v. Tiegh.)
v.
thwaitesii
4
Kuntze
(Thw.)
Palaeouratea
subg.
thorelii
3a
1
1:
Lamk.
zeylanica
1:
i:
parviflora (Griff.)
var.
Tiegh. 1:
pruinosum
4
2
(inoti L.)
(Jack)
1:
1:
Tiegh.
Tiegh.
v.
Tiegh.
DC.)
pellucidum
v.
4
Tiegh.
ex
v.
pumilum (DC.)
(Thw.)
Planch.
Chit
&
2
(non L.)
wallichii
4
1:
4
1:
v.
4
3a
pedunculatum
pierrei
squarrosa
walkeri
1:
Tiegh.
1:
1:
(Thunb.
nitidum
3c
Lecomte
squarrosa
sumatrana
1:
Hundley
1:
Linne
squarrosa
Kanis
1:
Tiegh. 1:
4
Tiegh.)
(Thw.)
moonii
a
1:
Tiegh.
v.
Tiegh.
v.
3a
Tiegh.
v.
3a
Tiegh. 1:
1:
Tiegh.
3a
1
1:
Tiegh.
v.
lefevrei
1:3b
4
DC.)
Thw.
rufescens
kingii
2
v.
Tiegh.
v.
v.
inaequale
3
1:
3c
1:
pumila (non
var.
3
(v. Tiegh.)
pruinosa
helferi
3c
latifolium (v.
1:
Griff.
parviflora
1:
3
gamblei (Brandis)
obtusata
arcta
1:
Planch.
pumila (DC.)
sect.
& A.
v.
v.
Tiegh.
1:
v.
Tiegh.
v.
(Thw.)
i:
1:3a
1
(v. Tiegh.)
Tiegh.
v.
v.
griffithii
var.
Tiegh.
(v. Tiegh.)
cochinchinense
4
cordatum
W.
var.
v.
(v. Tiegh.)
Polythecium
3a
1
nitida
DC.
1:
thorelii
2
1:
discolor
ex
nitida
var.
Linn.
Excl.
Buch.-Ham.
obtusata
1:
Linn.
syti.
syn.
3a
1:
Tiegh.
lefevrei (v. Tiegh.)
Griff.
mauritiana Lamk.
cambodianum
4
1
excl.
p.p.
1:
Tiegh.
v.
v.
Tiegh.
v.
cochinchinense
quoad
lucida
(DC.)
Polythecanthum
2
1:
lucida
(v. Tiegh.)
(Lamk.)
obtusatum
2
2:
3c
Merr.
1:
1968
I,
lucidum
i:
integerrima (Lour.)
jabotapita Linne
No.
leschenaultii
4
1
O. Kuntze
hookeri
lanceolata
1:
1:
XVI,
Willd.
3:
1
9:
1
4
3a