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1/7/2016 Organisms capture and store free energy for use in biological processes Introduction to Photosynthesis • Autotrophs capture free energy from physical sources in the environment • Photosynthesis versus chemosynthesis • Heterotrophs capture free energy present in carbon compounds produced by other organisms • Metabolism of carbohydrates, lipids, and proteins • Fermentation produces organic compounds (alcohol and lactic acid) and occurs in the absence of oxygen • Different energy-capturing processes use different types of electron acceptors • NADP+ in photosynthesis • Oxygen in cellular respiration Photosynthesis: light-dependent reactions + the Calvin cycle 1. 2. 3. 4. 5. Chlorophylls absorb free energy from light, boosting electrons to a higher energy level in photosystems I and II These photosystems are embedded in the internal membranes of chloroplasts (thylakoids) and are connected by the transfer of higher free energy electrons through an electron transport chain (ETC) As electrons pass through the ETC an electrochemical gradient of hydrogen ions (protons) across the thylakoid membrane is established This proton gradient is linked to the synthesis of ATP from ADP and inorganic phosphate via ATP synthase The energy captured in the light reactions as ATP and NADPH powers the production of carbohydrates from CO2 in the Calvin cycle, which occurs in the stroma of the chloroplast Take a mental journey….. Photosynthesis • First evolved in prokaryotic organisms • Evidence supports that prokaryotic (bacterial) photosynthesis was responsible for the production of an oxygenated atmosphere • Prokaryotic photosynthetic pathways were the foundation of eukaryotic photosynthesis Biological Hierarchy of a Plant Using the structures involved in photosynthesis, put in order from largest (organism) to smallest (molecule). 1 1/7/2016 Biological Hierarchy of a Plant • Organism – plant • Organ system – shoot system (leaves, buds, stems, flowers) • Organs – leaves • Tissues - vascular (xylem & phloem), dermal, ground • Cells – Parenchyma cells mesophyll cells are photosynthetic palisade & spongy mesophyll • Organelles – chloroplasts (thylakoid membranes & stroma) • Molecules – chlorophylls & proteins embedded in thylakoid membranes move around photons (particle of light energy) Use this diagram and page 461 to label the cross section of a leaf. Define – pgs. 101-107 • Autotroph • Pigment • Chlorophyll a • Chloroplast • Thylakoid membrane • Stroma • Photosystems • Photolysis • The energy captured photosynthesis forms the Photosynthesis &through Ecology • Photosynthesis is the process which Photosynthesis: ThebyBIG Picture PHOTOAUTOTROPHS convert the energy in SUNLIGHT into the energy stored in ORGANIC COMPOUNDS. basis of the ecological pyramid. • The biomass that is generated by producers supports nearly all the living organisms on the planet. • Autotrophs capture energy and “fix” it into organic compounds; heterotrophs consume compounds produced by other organisms to obtain energy. Don’t forget the 1st Law of Thermodynamics, which says… What is an autotroph? What is an organic compound? light 11 12 2 1/7/2016 Leaves: The Photosynthetic Organs of Plants • Leaves perform most of the photosynthesis in plants. • In eukaryotes,Structure: photosynthesis place inside Chloroplast Atakes Review chloroplasts inside cells (inside leaves). • Chloroplasts have 3 membranes: cuticle • Outer membrane • Inner membrane • Thylakoid membrane, folded to form thylakoids • Thylakoids are arranged in stacks called grana. upper epidermis palisade mesophyll spongy mesophyll lower epidermis stoma Bundlesheath cells Xylem & phloem 13 Light & Pigments Granum • Chlorophyll and other pigments involved in photosynthesis are embedded in the thylakoid membrane. Light & Pigments • Visible light is made up of different colors of light with different wavelengths. • Pigments are molecules that absorb light energy. • Different pigments absorb light of different wavelengths. • Light has a dual nature. • Accessory pigments: Chlorophyll B Carotenoids • Light exhibits properties of both waves and particles (photons). • Xanthophyll • Carotenes • Major photosynthetic pigment: Chlorophyll A 15 Light & Pigments What is the advantage 16 Light & Pigments What to having multiple pigments? information does this ABSORPTION SPECTRUM tell you? Which colors of light are absorbed well? Not well? What information does this ACTION SPECTRUM tell you? 17 18 3 1/7/2016 Photosynthesis: An Overview • The net overall equation for photosynthesis is: 6 CO2 + 6 H2O light C6H12O6 + 6 O2 • Photosynthesis occurs in 2 “stages”: 1. The Light Reactions (or Light-Dependent Reactions) 2. The Calvin Cycle (or Calvin-Benson Cycle or Dark Reactions or Light-Independent Reactions) Is photosynthesis an ENDERGONIC or EXERGONIC reaction? Photosynthesis: An Overview • Follow the energy in photosynthesis, Summary of Photosynthesis Stage Location Starting Ending 1st Thylakoid membranes H2O ADP NADP+ O2 ATP NADPH – Lightdependent Reactions 2nd – Calvin cycle (Lightindependent reactions) Stroma CO2 ATP NADPH 19 Light Reactions light thylakoids light ATP NADPH Calvin Cycle stroma Organic compounds (carbs) C6H12O6 ADP NADP+ 22 4