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Transcript
CHAPTER 8: DEVELOPING A THEORY OF
EVOLUTION
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Scientific knowledge develops as people observe the world around them, ask questions, and seek answers to their
questions.
A scientific hypothesis is a statement that provides one possible answer to a question, or one possible explanation
for an observation. Hypotheses are tested to determine their validity, mainly through experiments, observation,
developing models from data, or a combination of these activities. Hypotheses that lead to successful predictions
and explanations are sometimes synthesized into a general statement that explains and makes successful
predictions about a broad range of observations. Such a statement is called a scientific theory.
The Science of Paleontology
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Fossils are important to the study of evolution. A fossil is the preserved remains of a once-living organism.
Fossils include specimens preserved in amber, permafrost, dry caves, and the more common fossils are preserved
in rock.
George Cuvier (1769-1832) is largely credited with developing the science of paleontology.
PALEONTOLOGY is the study of ancient life through the examination of fossils.
Cuvier found that each stratum (layer of rock) is characterized by a unique group of fossil species. He also found
that the deeper (older) the stratum, the more dissimilar the species are form modern life. This evidence showed
that species could become extinct.
To explain his observations, Cuvier proposed the idea that Earth experience many destructive natural events in the
past, such as floods and volcanic eruptions. These events, which he called revolutions, were violent enough to
have killed numerous species each time they occurred. This idea is now known as CATASTROPHISM.
Other scientists had other ideas that differed from Cuvier. Scottish geologist Charles Lyell rejected
catastrophism. He proposed (based on the work of James Hutton) that geological processes operate at the same
rates in the past as they do today in a process called UNIFORMITARIANISM. Lyell reasoned that, if
geological changes are slow and continuous rather than catastrophic, then Earth might be more than 6000 years
old. He also theorized that slow, subtle processes could happen over a long period of time and could result in
substantial changes. The forces that build and erode mountains and the rate, at which such geological change
happens, are no different today than they were in the past. This idea inspired naturalist Charles Darwin and
others.
French naturalist Jean-Baptiste Lamarck (1744-1829) observed a “line of descent” or progression, in which a
series of fossils (from older to more recent) led to a modern species. He thought that species increased in
complexity over time, until they reached a level of perfection. He hypothesized that the organisms would become
progressively better adapted to their environments. He believed that body parts that were used extensively to
cope with conditions in the environment would become larger and stronger. For example, giraffes stretched their
necks to eat foliage from tall trees. Over time, they would pass on this stretched neck condition to their offspring.
Lamarck called his idea the INHERITANCE OF ACQUIRED CHARACTERISTICS. He
also suggested that body parts not used would eventually disappear. This idea is called use and disuse.
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Charles Darwin (1809-1882) created his famous THEORY
OF EVOLUTION BY NATURAL
SELECTION which is a theory is explaining how life has changed, and continues to change, during Earth’s
history.
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Darwin’s observations from his expeditions to the coast of South America included:
a) The flora and fauna of the different regions were distinct from those on other continents.
b) He observed fossils of extinct animals that looked very similar to living animals. For example, the armadillolike glyptodont.
c) The finches and other animals Darwin saw on the Galapagos Islands closely resembled animals he had
observed on the west coast of South America.
d) Galapagos species looked identical at first, but they actually varied slightly between islands. Each type of
Galapagos finch, for example, was adapted to eating a different type of food based on the size and shape of its
beak.
e) Through his experience with artificial selection, Darwin knew that it was possible for traits to be passed on
from parent to offspring, and that sexual reproduction resulted in many variations within a species.
Alfred Russel Wallace (1823-1913) reached conclusions that were similar to Darwin’s. They both accepted that
populations changed as time passed, but they were unclear how populations changed. According to Darwin and
Wallace, individuals with traits that helped them survive in their local environments were more likely to survive
and pass on these traits to offspring. They reasoned that competition for limited resources between individuals of
the same species would select for individuals with favourable traits – traits that increased their chances of
surviving and reproducing. Therefore, a growing proportion of the population would have these traits in later
generations and, as time passed, the population as a whole would have them. This is the idea of SURVIVAL
OF THE FITTEST, and Darwin called this process natural selection.
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Darwin proposed in his book, The Origin of Species, that all life descended from some unknown organism. As
descendants of this organism spread out over different habitats during the millennia, they developed adaptations
that helped them better survive in their local environments. His ideas are summarized as follows:
a) Organisms produce more offspring that can survive. Therefore, organisms compete for limited resources.
b) Individuals of a population vary extensively, and much of this variation is heritable.
c) Individuals that are better suited to local conditions survive to produce more offspring.
d) Processes for change are slow and gradual.
Sources of Evidence for Evolution
1) Sedimentary rock with fossils provides a FOSSIL RECORD of the history of life by showing the kinds
of species that were alive in the past. For instance, when people examined the Burgess Shale fossil beds in British
Columbia, they found fossils of animals that lived in an ancient ocean during the Cambrian period, over 500
million years ago. Some of the fossilized animals found in the Burgess Shale are ancestors of animals that are
common today.
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Evidence from the fossil record provides the following:
a) Fossils found in young layers of rock (from recent geological periods and usually closer to the surface) are
much more similar to species alive today than fossils found in older, deeper layers of rock.
b) Fossils appear in chronological order in the rock layers. So, probable anscestors for a species are found in
older rocks, which usually lie beneath the rock in which the later species is found.
c) Not all organisms appear in the fossil record at the same time. For example, the fossil history of vertebrates
shows that fish are the oldest vertebrates. In subsequent layers, the fossils of other vertebrates – amphibians,
reptiles, mammals, and birds – appear. This reinforces scientific evidence that amphibians evolved from
ancestral fish, reptiles evolved from ancestral amphibians, and both mammals and birds evolved form different
groups of reptiles (mammals first, and then birds).
2) Evidence from TRANSITIONAL FOSSILS (which are fossils that show intermediary links between
groups of
organisms and share characteristics common to two now separate groups). Transitional fossils link the past with
the present. For example, scientists have found fossilized whales that lived 36 to 55 million years ago. These
fossils link present-day whales to terrestrial ancestors. They had a tiny pelvis and small legs. These
characteristics would have been useless to an animal that lived an aquatic life.
 Structures that are the reduced form of structures that were functional in the organism’s ancestors are called
VESTIGIAL STRUCTURES.
3) Evidence from BIOGEOGRAPHY, which is the study of the past and present geographical distribution of
species
populations, supports the following:
a) Geographically close environments (for example: desert and forest habitats in South America) are more
likely to be populated by related species than are locations that are geographically separate but environmentally
similar (for example: desert in Africa and desert in Australia).
b) Animals found on islands often closely resemble animals found on the closest continent. This suggests that
animals on islands have evolved form mainland migrants, with populations becoming adapted over time as they
adjust to the environmental conditions of their new homes. For example, the lizards found on the Canary
Islands, off the northwest coast of Africa, are very similar to the lizards found in West Africa.
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c) Fossils of the same species can be found on the coastline of neighbouring continents. For example, fossils of
the reptile Cynognathus have been found in Africa and South America. The location of continents is not fixed –
they are slowly moving away from one another. About 510 million years ago, the continents of Africa and
South America were jointed in one supercontinent, called Gondwana.
d) Closely related species are almost never found in exactly the same location or habitat.
4) Evidence from anatomy: Vertebrate forelimbs can be used for various functions, such as flying (birds and bats),
running (horses and dogs), and swimming (whales and seals). Despite their different functions, all vertebrate
forelimbs contain the same set of bones, organized in simlar ways.
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HOMOLOGOUS STRUCTURES are those that have similar structural elements and origin but
may have a different function. For instance, the limbs have similar structures, such as the number of bones,
muscles, ligaments, tendons, and blood vessels. These structural elements are arranged to be best suited for
different functions: walking, flying, or swimming. Homologous structures are similar because they were
inherited from a common ancestor.
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ANALOGOUS STRUCTURES are structures of organisms that do not have a common
evolutionary origin but perform similar functions. For instance, types of wings vary between type of bird or
bat.
5) Evidence from EMBRYOLOGY, which is the study of early, pre-birth stages of an organism’s
development.
Embryology has also been used to determine evolutionary relationships between animals. The embryos of
different organisms exhibit similar stages of embryonic development.
 For example, all vertebrates embryos have paired pouches, or out-pocketings, of the throat. In fish and some
amphibians, the pouches develop into gills. In humans, the pouches become part of the ears and throat.
 At certain stages in the development of the embryo, the similarities between vertebrates are more apparent
than the differences.
 The similarities between embryos in related groups (such as vertebrates) point to a common ancestral origin.
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6) Evidence from DNA: Since DNA carries genetic information, scientists can determine how closely related two
organisms are by comparing their DNA. If two species have similar patterns in their DNA, this indicates that
these DNA sequences must have been inherited form a common ancestor. For example, by studying gene
sequences, scientists have determined that dogs are related to bears and that whales and dolphins are related to
ungulates (hoofed animals such as cows and deer).
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