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Taxonomy is a branch of biology that names and groups organisms according to their characteristics and evolutionary history. Aristotle (384-322 B.C.) was the first to classify living organisms. This greek philospher grouped animals into two categories. The particular organism was classified as either plant or animal origin. As time progressed, modern science and rapid research identified many new organisms. Aristotle’s classification was not sufficient. The Swedish naturalist Carolus Linnaeus (1707-1778) realized that such a system was not conducive for modern biology. Linnaeus realized that common names caused a problem as they varied from region to region. For example a jelly fish is not a fish, or a sea horse is not a horse. Linnaeus developed a classification system of organizing organisms into hierarchal categories. He did this be using the physical appearances of organisms to group them. Viruses are not in ANY of these kingdoms, scientists do not classify them as 'alive' Linnaeus made two main contributions to taxonomy. He was instrumental in developing hierarchal categories and binomial nomenclature. Hierarchal classification includes seven different levels of organization. Kingdom, Phyla, Classes, Orders, Families, Genera, and Species are included in the seven. A mnemonic device used to remember this could be Kings Play Calgary On Friday Gretzky Scores. Kingdom Phylum Class Order Family Genus Species Animalia Chordata Mammalia Primates Hominidae Homo Sapiens Humans Animalia Chordata Mammalia Carnivora Felidae Felis Domesticus House cat We identify organisms by using their Genus name and their species name. These names are either going to be in Greek or Latin. For example, humans would be known as Homo Sapiens. This two-name system is known as binomial nomenclature. This system allows scientists to have universal communication among different speaking countries. For many years, most biologists favored a Five-Kingdom system consisting of Kingdoms Monera, Protista, Fungi, Plantae, and Animalia. They were placed into categories by their type of cell (prokaryotic or eukaryotic), level of organization (unicellular or multicellular), and how they acquire their nutrition. It is suggested that protests evolved from the Monerans who are the simplest organism. Fungus, Plants and Animals evolved from the Protists in three separate evolutionary lines. The five kingdom system of classification is based on structural differences and also on modes of nutrition among the eukaryotes. A New Proposal: The Three Domains of Life In the 1970’s scientists began to find evidence for a previously unknown group of prokaryotic organisms. These organisms lived in extreme environments such as the Dead Sea, acid lakes, and salt evaporation ponds. These are environments that scientists never suspected of maintaining any life. Because they appeared prokaryotic, they were considered bacteria and named "archaebacteria" ('ancient' bacteria). However, became obvious from biochemical characteristics and DNA sequence analysis that there were numerous differences between these archaebacteria and other bacteria. Before long, it was realized that these archaebacteria were more closely related to the eukaryotes than to bacteria. Today, these bacteria have been renamed Archaea. From this work scientistspropesed that there should be a new caterogy of classification of life - the Domain, a classification category above Kingdom. The traditional 5 Kingdom system says nothing about how organisms within Kingdoms or between kingdoms may be related to each other via evolutionary relationships among the kingdoms. A New Proposal is the Three Domains of Life. The three domains are Bacteria, Archaea, and Eukarya. The three-domain system of classification is based on biochemical differences that show they are three vastly different groups of organisms. See table 27.3 on page 561 titled Classification Criteria for the Three Domains. How was the 'Three Domain' Classification of Life Determined? Scientists used the nucleotide sequence of Ribosomal rRNA (the small subunit) and other RNA and protein sequences as an “Evolutionary Chronometer” – an evolutionary time clock. What makes rRNA (or another sequence) a good ‘chronometer’? 1. It is universally distributed across group chosen – all organisms have rRNA 2. It is functionaly similar between organisms – rRNAs all participate in protein synthesis 3. Its sequence changes slowly - good for looking across long periods of time 4. The rRNA sequences can be aligned, or matched up, between 2 organisms 1. What are the names of the 3 Domains? Who is Carl Woese and what was his role developing the concept of 3 Domains? 2. What is rRNA and why was did scientists choose it as an Evolutionary Chronometer? 3. Bacteria: What is the significance of the Proteobacteria and the Cyanobacteria in the development of eukaryotic organelles? 4. Archaea: Distinguish between the three groups. Why are thay called 'extremeophiles'? 5. Eukarya: Be able to list the 4 Kingdom within Eukarya and give 1 representative organisms for each. (You do not need to know dates, numbers of species, or other facts provided in the quotes). Horse Phylum: Chordata Class: Mammalia Order: Perissodactyla Family: Equidae Species: Equus caballus Amoeba proteus