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Table of Contents Preface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . x INVITED CONTRIBUTIONS 1 Non-random random mutations: a signature of evolution of evolution (EVOEVO) . . . . . . . . . . . . . . . . . . Paulien Hogeweg 1 How do the origins of life sciences influence 21st century design thinking? . . . . . . . . . . . . . . . . . . . . . . . Rachel Armstrong 2 Hybrid Forms (2015) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Andy Lomas 12 Do Endothelial Cells Dream of Eclectic Shape? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Katie Bentley 13 SPECIAL SESSION: SLIME MOULD COMPUTERS 14 Physarum Polycephalum changes polyaniline properties . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Alice Dimonte, Tatiana Berzina, Victor Erokhin 14 Majority Gates and Circular Computation in Slime Mould . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Genaro J. Martı́nez, Andrew Adamatzky, Richard Mayne, Fangyue Chen, Qinbin He 18 Slime mould actin sensoriactuation networks: topology, dynamic transformations and models of self-assembly . . Richard Mayne, Andrew Adamatzky 25 Mimicking the Exploration of 3D Terrains by Physarum with Cellular Automata Models . . . . . . . . . . . . . . Michail-Antisthenis I. Tsompanas, Georgios Ch. Sirakoulis 33 SPECIAL SESSION: MORPHOGENETIC ENGINEERING 41 Behavioral Diversities of Morphogenetic Collective Systems . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hiroki Sayama 41 Morphogenesis and Replication of Multi-Cellular Organisms with Evolved Variable Length Self-Modifying Genomes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Stefano Nichele, Gunnar Tufte 42 Modelling the Role of Trail Pheromone in the Collective Construction of Termite Royal Chambers . . . . . . . . . Nicholas Hill, Seth Bullock 43 Proceedings of the European Conference on Artificial Life 2015 iii From tadpole to frog: artificial metamorphosis as a method of evolving self-reconfiguring robots . . . . . . . . . . Michał Joachimczak, Reiji Suzuki, Takaya Arita 51 Partial Redundancy and Morphological Homeostasis: Reliable Development through Overlapping Mechanisms . Micah Brodsky 59 MecaCell: an Open-source Efficient Cellular Physics Engine . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Jean Disset, Sylvain Cussat-Blanc, Yves Duthen 67 SPECIAL SESSION: QUANTIFYING EMBODIMENT 68 Musculo-Skeletal Models as Tools to Quantify Embodiment . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Daniel F. B. Haeufle, Michael Günther, Syn Schmitt 68 Task Dynamics & the (Ecological) Information They Create . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Andrew D Wilson, Sabrina Golonka 69 Quantifying Morphological Computation based on an Information Decomposition of the Sensorimotor Loop . . . Keyan Ghazi-Zahedi, Johannes Rauh 70 Quantifying Self-Organizing Behavior of Autonomous Robots . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Georg Martius, Eckehard Olbrich 78 MAIN ECAL TRACK 79 Modelling Conflict within the Social Networks of Large Multi-Vendor Software Projects using Communicating Stream X-Machines . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Richard Alun Williams 79 Dynamic Homeostasis in Packet Switching Networks . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mizuki Oka, Hirotake Abe, Takashi Ikegami 80 How social networks shape collective behaviours . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Daniel W. Franks, A. Jamie Wood, Nikolai W.F. Bode 81 At the root of sociality: Working towards emergent, permanent, social affines . . . . . . . . . . . . . . . . . . . . . Ovi Chris Rouly 82 The Circular Logic of Gaia: Fragility and Fallacies, Regulation and Proofs . . . . . . . . . . . . . . . . . . . . . . Inman Harvey 90 Conservation of Matter Increases Evolutionary Activity . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Simon Hickinbotham, Susan Stepney 98 Evolving Robot Controllers Using Carbon Nanotubes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 106 Maktuba Mohid, Julian F. Miller A Minimal Model for the Emergence of Cooperation in Randomly Growing Networks . . . . . . . . . . . . . . . . 114 Steve Miller, Joshua Knowles The Kinetic Basis of Morphogenesis . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 122 Yuri Shalygo Environmental Factors and the Emergence of Cultural Technical Innovations . . . . . . . . . . . . . . . . . . . . 130 Peter Andras Proceedings of the European Conference on Artificial Life 2015 iv Where Does (Co)evolution Lead to? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 Jan Paredis A New View of Protocell Metabolism . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 Ben Shirt-Ediss, Ricard Solé, Kepa Ruiz-Mirazo Ribosome synthesis and construction of a minimal cell using a cell-free expression platform . . . . . . . . . . . . . 147 Filippo Caschera, Michael C. Jewett The Universality of Peer-Influence in Social Networks . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 149 Flávio L. Pinheiro, Marta D. Santos, Francisco C. Santos, Jorge M. Pacheco Programs as Polypeptides . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 Lance R. Williams Practical Fault Tolerant 2D Cellular Automata . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 158 Steven Janke, Matthew Whitehead Evaluating the Effect of a Flexible Spine on the Evolution of Quadrupedal Gaits . . . . . . . . . . . . . . . . . . . 166 Jared M. Moore, Craig P. McGowan, Philip K. McKinley Evolving Collective Behaviors With Diverse But Predictable Sensor States . . . . . . . . . . . . . . . . . . . . . . . 174 Payam Zahadat, Heiko Hamann, Thomas Schmickl Learning by Stimulation Avoidance as a Primary Principle of Spiking Neural Networks Dynamics . . . . . . . . . 175 Lana Sinapayen, Atsushi Masumori, Nathaniel Virgo, Takashi Ikegami Contagion on Networks with Self-Organised Community Structure . . . . . . . . . . . . . . . . . . . . . . . . . . 183 Alberto Antonioni, Seth Bullock, Christian Darabos, Mario Giacobini A Passive Mechanism for Goal-Directed Navigation using Grid Cells . . . . . . . . . . . . . . . . . . . . . . . . . . 191 Vegard Edvardsen An Integrated Neuromechanical Model of Steering in C. elegans . . . . . . . . . . . . . . . . . . . . . . . . . . . . 199 Eduardo J. Izquierdo, Randall D. Beer CriPS: Critical Particle Swarm Optimisation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 207 Adam Erskine, J. Michael Herrmann Embodied Evolution of Artificial Cells in a Hybrid Wet/Hard-ware Platform . . . . . . . . . . . . . . . . . . . . . 215 Juan Manuel Parrilla Gutierrez, Leroy Cronin Ontogeny and adaptivity in a model protocell . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 216 Eran Agmon, Alexander J. Gates, Randall D. Beer Chemical Architectures for Self-Replicating Proto-Cells . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 224 Erwan Bigan, Jean-Marc Steyaert, Stéphane Douady A population of information processing objects as a model of evolutionary dynamics . . . . . . . . . . . . . . . . . 225 Richard J Carter, Stephen Mann, Karoline Wiesner Evolving strategies for single-celled organisms in multi-nutrient environments . . . . . . . . . . . . . . . . . . . . 226 Dominique Chu, David J. Barnes Emergence-focused design in complex system simulation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234 Chris Marriott, Jobran Chebib Proceedings of the European Conference on Artificial Life 2015 v The Lattice of Chemical Organisations . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 242 Pietro Speroni di Fenizio Coevolution of cooperation and layer selection strategies in multiplex networks . . . . . . . . . . . . . . . . . . . . 249 Katsuki Hayashi, Reiji Suzuki, Takaya Arita Volatility and spatial distribution of resources determine ant foraging strategies . . . . . . . . . . . . . . . . . . . 256 Drew Levin, Joshua P. Hecker, Melanie E. Moses, Stephanie Forrest Applying homeostatic neural controller to multi-legged robot and adaptivity to novel disruptions . . . . . . . . . . 264 Hiroyuki Iizuka, Hiroki Nakai, Masahito Yamamoto Investigating a cellular automata model that performs three distance diffusion on a robot path planning . . . . . . 271 Gina M. B. Oliveira, Patrı́cia A. Vargas, Giordano B. S. Ferreira The Evolution of Assortment with Multiple Simultaneous Games . . . . . . . . . . . . . . . . . . . . . . . . . . . . 279 S. Tudge, A. Jackson, R. Watson, M. Brede Exploring the organisation of complex systems through the dynamical interactions among their relevant subsets . 286 Alessandro Filisetti, Marco Villani, Andrea Roli, Marco Fiorucci, Roberto Serra Environmental bias forces parasitism in Tierra . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 294 Simon Hickinbotham, Susan Stepney Distributed vs. Centralized Particle Swarm Optimization for Learning Flocking Behaviors . . . . . . . . . . . . . 302 Iñaki Navarro, Ezequiel Di Mario, Alcherio Martinoli Unraveling the genotype-phenotype map of evolving digital organisms . . . . . . . . . . . . . . . . . . . . . . . . . 310 Miguel A. Fortuna, Luis Zaman, Charles Ofria, Andreas Wagner Incorporation of Emotions in the Orphibs Agent Architecture . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 311 Nuno Barreto, Luı́s Macedo, Penousal Machado, Licı́nio Roque Cooperative Coevolution of Morphologically Heterogeneous Robots . . . . . . . . . . . . . . . . . . . . . . . . . . 312 Jorge Gomes, Pedro Mariano, Anders Lyhne Christensen Time Series Evolution for Integrating Developmental Processes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 320 Sebastian von Mammen, Melanie Däschinger Heredity in Messy Chemistries . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 325 Nathaniel Virgo, Nicholas Guttenberg Following Strategies Reduces Accidents, but Makes Outcomes Worse: Evidence from Simulated Treefrog Mating Scenarios . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 333 Giordano B. S. Ferreira, Matthias Scheutz Incremental Neuroevolution of Reactive and Deliberative 3D Agents . . . . . . . . . . . . . . . . . . . . . . . . . . 341 Adam Stanton, Alastair Channon Achieving Compositional Language in a Population of Iterated Learners . . . . . . . . . . . . . . . . . . . . . . . 349 Lewys Brace, Seth Bullock, Jason Noble The Hunger Games: Embodied agents evolving foraging strategies on the frugal-greedy spectrum . . . . . . . . . 357 Nathanaël Aubert-Kato, Olaf Witkowski, Takashi Ikegami Wallace: An efficient generic evolutionary framework . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 365 Christopher Steven Timperley, Susan Stepney Proceedings of the European Conference on Artificial Life 2015 vi Emergence of Sense-Making Behavior by the Stimulus Avoidance Principle: Experiments on a Robot Behavior Controlled by Cultured Neuronal Cells. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 373 Atsushi Masumori, Norihiro Maruyama, Lana Sinapayen, Takeshi Mita, Urs Frey, Douglas Bakkum, Hirokazu Takahashi, Takashi Ikegami Omnigram Explorer: A Simple Interactive Tool for the Initial Exploration of Complex Systems . . . . . . . . . . 381 Tim Taylor, Alan Dorin, Kevin Korb The Digital Aquarist: An Interactive Ecology Simulator . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 389 Julian Schikarski, Oliver Meisch, Sarah Edenhofer, Sebastian von Mammen Toward a behavior-based approach to the origins of life and the genetic system . . . . . . . . . . . . . . . . . . . . 397 Tom Froese An Efficient Ant Colony System for Edge Detection in Image Processing . . . . . . . . . . . . . . . . . . . . . . . . 398 Yara Khaluf, Syam Gullipalli Abnormality Detection in Robots Exhibiting Composite Swarm Behaviours . . . . . . . . . . . . . . . . . . . . . . 406 Danesh Tarapore, Anders Lyhne Christensen, Jon Timmis The Overshoot Curriculum: Artificial Life, Education and the Human Predicament . . . . . . . . . . . . . . . . . 414 Barry McMullin Emergence of Competition between Different Dissipative Structures for the Same Free Energy Source . . . . . . . 415 Stuart Bartlett, Seth Bullock Planarity as a driver of Spatial Network structure . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 423 Garvin Haslett, Markus Brede Emotional modulation of peripersonal space impacts the way robots interact . . . . . . . . . . . . . . . . . . . . . 431 Marwen Belkaid, Nicolas Cuperlier, Philippe Gaussier Simulating self-replicating, chemically immersed, microchip swarms . . . . . . . . . . . . . . . . . . . . . . . . . . 438 Uwe Tangen, Harold Fellermann, Steen Rasmussen Epigenetic inheritance speeds up evolution of artificial organisms . . . . . . . . . . . . . . . . . . . . . . . . . . . . 439 Yoram Vadée-le-Brun, Jonathan Rouzaud-Cornabas, Guillaume Beslon Probing models of minimal swimming vehicules in vivo with microalgae phototaxis. . . . . . . . . . . . . . . . . . 447 David Colliaux, Lia Giraud, Claude Yéprémian, Pierre Bessière, Jacques Droulez Towards Failure-Resistant Mobile Distributed Systems Inspired by Swarm Intelligence and Trophallaxis . . . . . 448 Arles Rodrı́guez, Jonatan Gómez, Ada Diaconescu Autonomous lablet locomotion and active docking by sensomotory electroosmotic drive . . . . . . . . . . . . . . . 456 Abhishek Sharma, John S. McCaskill A Predator-Prey Scenario in a Virtual Ecosystem . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 463 Nesrine Ouannes, NourEddine Djedi, Yves Duthen, Hervé Luga Cooperative Object Transport Using Evolutionary Swarm Robotics Methods . . . . . . . . . . . . . . . . . . . . . 464 Muhanad H. Mohammed Alkilabi, Chuan Lu, Elio Tuci Closed-loop acquisition of behaviour on the Sphero robot . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 472 Oswald Berthold, Verena V. Hafner Proceedings of the European Conference on Artificial Life 2015 vii How to play the Syntax Game . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 479 Luc Steels, Emilia Garcia-Casademont How Much Should You Select for Evolvability? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 487 Andrew M. Webb, Julia Handl, Joshua Knowles Evolution of Cooperation in Evolutionary Robotics: the Tradeoff between Evolvability and Efficiency . . . . . . . 495 Arthur Bernard, Jean-Baptiste André, Nicolas Bredeche An Investigation of Square Waves for Evolution in Carbon Nanotubes Material . . . . . . . . . . . . . . . . . . . . 503 Odd Rune Lykkebø, Stefano Nichele, Gunnar Tufte Apparent actions and apparent goal-directedness . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 511 Martin Biehl, Daniel Polani Evolutionary Progress in Heterogenous Cellular Automata (HetCA) . . . . . . . . . . . . . . . . . . . . . . . . . . 512 David Medernach, Jeannie Fitzgerald, Simon Carrignon, Conor Ryan Simpsons Paradox, Co-operation and Individuality in Bacterial Biofilms . . . . . . . . . . . . . . . . . . . . . . . . 520 Alexandra Penn Emergent Robustness in Software Systems through Decentralized Adaptation: an Ecologically-Inspired ALife Approach . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 521 Franck Fleurey, Benoit Baudry, Benoit Gauzens, André Elie, Kwaku Yeboah-Antwi Steering a Complex Adaptive System: A Complexity Science Design Methodology Applied to an Industrial Ecosystem in the Humber Region, UK . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 529 Alexandra S. Penn Recombination Is Surprisingly Constructive for Artificial Gene Regulatory Networks in the Context of Selection for Developmental Stability . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 530 Yifei Wang, Yinghong Lan, Daniel M. Weinreich, Nicholas K. Priest, Joanna J. Bryson Searching for non-regular neighborhood cellular automata rules applied to scheduling task and guided by a forecast dynamical behavior parameter . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 538 Tiago I. Carvalho, Gina M. B. Oliveira Investigating the Impact of Communication Quality on Evolving Populations of Artificial Life Agents . . . . . . . 546 Sadat Chowdhury, Elizabeth I. Sklar Artificial life programming in the robust-first attractor . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 554 David H. Ackley, Elena S. Ackley Towards a methodology for describing the relationship between simulation and reality . . . . . . . . . . . . . . . . 562 Eric Schneider, Elizabeth I. Sklar, M. Q. Azhar, Simon Parsons, Karl Tuyls Achieving Closure in Enzymes in Artificial Chemistries . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 570 Matteo Monti, Tim Hutton, Piet Hut How Intrinsic Motivation can Speed Up Language Emergence . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 571 Miquel Cornudella, Paul Van Eecke, Remi van Trijp Coordination of collective behaviours in spatially separated agents . . . . . . . . . . . . . . . . . . . . . . . . . . . 579 Rob Mills, Payam Zahadat, Fernando Silva, Damjan Mlikic, Pedro Mariano, Thomas Schmickl, Luı́s Correia Corpus-taught Evolutionary Music Composition . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 587 Csaba Sulyok, Andrew McPherson, Christopher Harte Proceedings of the European Conference on Artificial Life 2015 viii Does self-replication imply evolvability? . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 595 Thomas LaBar, Christoph Adami, Arend Hintze Effects of Social Network Size and Topology on Evolutionary Decision Making . . . . . . . . . . . . . . . . . . . . 603 Hiroki Sayama, Shelley D. Dionne, Francis J. Yammarino Soft-Body Muscles for Evolved Virtual Creatures: The Next Step on a Bio-Mimetic Path to Meaningful Morphological Complexity . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 604 Dan Lessin, Sebastian Risi Cascading Transitions in Coupled Complex Ecosystems . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 612 Iain S. Weaver Evolving an optimal group size in groups of prey under predation . . . . . . . . . . . . . . . . . . . . . . . . . . . 620 Patrick B. Haley, Randal S. Olson, Fred C. Dyer, Christoph Adami A Metamodel for the Evolution of Evolution . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 621 Paul Andrews, Susan Stepney Informational parasites in code evolution . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 629 Andrés C. Burgos, Daniel Polani Simulation-Based Analysis of in Situ Cellular Motility . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 637 Mark Read, Jon Timmis, Tatyana Chtanova Simulating the Influence of Diet on the Intestinal Microbiome Composition . . . . . . . . . . . . . . . . . . . . . . 638 Mark Read, Andrew J. Holmes, Madison Hartill-Law, Samantha Solon-Biet, David Raubenheimer, Stephen J. Simpson Evolutionary change precedes extinction in eco-evolutionary dynamics based on a 3D virtual predator-prey system 639 Takashi Ito, Marcin L. Pilat, Reiji Suzuki, Takaya Arita Optimal Mutation Rate Control under Selection in Hamming Spaces . . . . . . . . . . . . . . . . . . . . . . . . . 640 Elizabeth Aston, Alastair Channon, Roman V. Belavkin, Rok Krasŏvec, Christopher G. Knight An Agent-Based Model of Avascular Tumor Growth . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 648 Ana Victoria Ponce Bobadilla, René Doursat, François Amblard Spatial Computing in Synthetic Bioware: Creating Bacterial Architectures . . . . . . . . . . . . . . . . . . . . . . 656 Jonathan Pascalie, Martin Potier, Taras Kowaliw, Jean-Louis Giavitto, Olivier Michel, Antoine Spicher, René Doursat Do it yourself (DIY) liquid handling robot for evolutionary search exploration . . . . . . . . . . . . . . . . . . . . 657 Asbjørn Müller, Aleksandra Amaldass, Kliment Yanev, Steen Rasmussen Human Swarms, a real-time method for collective intelligence . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 658 Louis B. Rosenberg Author Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 660 Proceedings of the European Conference on Artificial Life 2015 ix