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Evolution 2.1 Darwin's Voyage on the HMS Beagle Darwin's voyage on the HMS Beagle allowed him the opportunity to study a large variety of animal and plant species across the globe. Years after his voyage, he published the book On the Origin of Species, in which he proposed natural selection as the mechanism that underlies the process of evolution (figure 2.3) (134.0K) . The theory of evolution through natural selection is overwhelmingly accepted by scientists but is viewed as controversial by some in the general public. 2.2 Darwin's Evidence Darwin observed fossils in South America of extinct species that resembled living species. On the Galápagos Islands, Darwin observed finches that differed slightly in appearance between islands but resembled finches found on the South American mainland. These observations laid the groundwork for his proposal of evolution through natural selection (figure 2.4) (153.0K) and (figure 2.5) (101.0K) . 2.3 The Theory of Natural Selection Key to Darwin's hypothesis of evolution by natural selection was the observation by Malthus that the food supply limits population growth. A population, be it plants, animals, etc., grows only as large as that which can live off of the available food (figure 2.6) (83.0K) . Using Malthus's observations and his own observations, Darwin proposed that individuals that are better suited to their environments survive to produce offspring, gaining the opportunity to pass their characteristics on to future generations. Therefore, future generations may be different from ancestral populations, a process known as evolution. Spurred by a similar proposal put forth years later by Alfred Wallace, Darwin published his book in 1859. Darwin's Finches: Evolution in Action 2.4 The Beaks of Darwin's Finches By observing the different sizes and shapes of beaks in the closely related finches of the Galápagos Islands and correlating the beaks with the types of food consumed by the different birds, Darwin concluded that the birds' beaks were modified from an ancestral species based on the food available, each suited to its food supply (figure 2.10) (116.0K) . 1 Research has since supported Darwin's hypothesis that natural selection influences beak size in island finches based on the available food supply 2.11) (68.0K) . (figure 2.5 How Natural Selection Produces Diversity The 14 species of finches found on the islands off the coast of South America evolved from a mainland species that adapted to different niches, a process called adaptive radiation (figure 2.12) (60.0K) . Ecology 2.6 What Is Ecology? Ecology is the study of how organisms interact with each other and with their physical environment. There are five levels of ecological organization: populations, communities, ecosystems, biomes, and the biosphere. 2.7 A Closer Look at Ecosystems An ecosystem is a physical environment that contains a community of various organisms. These organisms interact with each other and with their physical environment, extracting energy and raw materials from it. Similar ecosystems found throughout the world are called biomes (figure 2.14) (106.0K) . 2.8 How Species Evolve to Occupy Different Niches Within an Ecosystem Resource partitioning is the process whereby two competitive species coexist by utilizing different portions of the habitat or different resources such as food. The varying beak sizes seen in Darwin's finches suggest that the bird species evolved in part due to resource partitioning (figure 2.15) (109.0K) and (figure 2.16) (118.0K) . Populations and How They Grow 2.9 Patterns of Population Growth Populations grow exponentially until limited by environmental conditions. A population will stabilize at a size, called its carrying capacity, that can be supported by its environment (figure 2.18) (75.0K) . Organisms exposed to an ever-changing environment tend to reproduce and grow rapidly, exhibiting an r-selected life history. Organisms exposed to more stable environmental conditions tend to reproduce and grow more slowly, exhibiting a kselected life history. 2 2.10 Human Populations Technology has allowed the human population to grow exponentially for the last 300 years to a current population of over 6 billion (figure 2.21) (71.0K) . Human populations grow at different rates, with underdeveloped countries' populations growing more rapidly than developed countries' populations. However, it takes more resources to support populations in developed countries versus those in underdeveloped countries (figure 2.22) (94.0K) and (figure 2.23) (83.0K) . 3