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Chapter 1 – Classifying Life’s Diversity 1.1 – Identifying, Naming, and Classifying Species species: a group of organisms that can interbreed in nature and produce fertile offspring Identifying Species: Using Species Concepts - Biologists use the morphological species concept, the biological species concept, and the phylogenetic species concept to define species. - The morphological species concept focuses on morphology – body shape, size, and other structural features. - The biological species concept defines species on the basis of whether two organisms can produce fertile offspring. - The phylogenetic species concept examines the phylogeny, or evolutionary history, of organisms. Species Concept Morphological Concept Is Based On appearance Biological behaviour and ability to mate and produce fertile offspring Phylogenetic evolutionary relationships Advantages and Disadvantages simple, but there is often significant variation in appearance within a species widely used, but cannot be used with many extinct species or separated species can be used with extinct species and makes use of DNA evidence, but evolutionary relationships are not always known Classifying Species - TAXONOMY is the branch of biology that identifies, names, and classifies species. - Each species is named using two words. Example: Felis domesticus (house cat). It is often referred to as the scientific name. - The system of using a two-word name for each species is called BINOMIAL NOMENCLATURE and was found nearly 300 years ago by Carolus Linnaeus. The first word is the name of the genus in which the organism is classified and the first letter is capitalized. The second word is the name identifies a particular species. - All species are classified by being in eight nested ranks (taxonomic categories). The broadest category is the domain, continuing to narrow to kingdom, phylum, class, order, family, genus, and finally species, which is the narrowest category) - Domains are the largest taxa (millions of species) and species is the smallest taxa (one organism). (TABLE 1.2, pg 15 – Taxonomic Classification of the Grey Wolf) Rank (Taxonomic Category) Domain Kingdom Phylum Class Order Family Genus Species Grey Wolf Taxon Eukarya Animalia Chordata Mammalia Carnivora Canidae Canis Canis lupus Number of Species in Taxon 4 – 10 million 2 million 50 000 5 000 270 34 7 1 - Here’s are a couple of mnemonics to help you remember the correct order of taxa: “Deaf King Philip Came Over From Germany Swimming.” OR “Does Kim Play Chess Or Fix Great Sandwiches?” - Homework: pg 16 Work on Questions #1 – 11 1.2 – Determining How Species Are Related - The goal of modern classification is to assign species to taxa so that the classification reflects both morphological similarities among organisms as well as hypotheses about their phylogeny (evolutionary history). It is thought that the more a species shares it evolutionary history with another species, the more closely related they are. Evidence of Relationships Among Species - There are three main types of evidence used to determine how closely related different species are. These are: anatomical, physiological, and DNA evidence. - Anatomical evidence includes comparing the structure and form of organisms, including bones. - Physiological evidence includes comparing the biochemistry of organisms, including proteins. - DNA evidence includes comparing organisms’ DNA sequences. Phylogenetic Trees - Once scientists have studied the features of organisms and learned more about their evolutionary histories, they often use a tool called a phylogenetic tree to represent a hypothesis about the evolutionary relationships among groups of organisms. (See Figure 1.5, pg 17 and Figure 1.12, pg 21) - Like a family tree, the roots or the base of the phylogenetic tree represents the oldest ancestral species. The upper ends of the branches represent present-day species that are related to the ancestral species. - Forks in each branch represent the points in the past at which an ancestral species split – evolved, or changed over time – to become two new species. - Understanding phylogeny can help scientists trace the transmission of disease and develop and test possible treatments. - Understanding the evolutionary relationships among species and groups of organisms can also have important consequences in agriculture and in the conservation of biodiversity. - Homework: pg 23 #1 - 10 1.3 – Dichotomous Keys - Taxonomists use dichotomous keys to make choices between pairs of options to narrow down identifications. dichotomous key: - an identification tool consisting of a series of two-part choices that lead the user to a correct identification A dichotomous key is a device used to easily and quickly identify unknown organisms. The key is a sequence of choices between two statements (couplets) based on characteristics and lead to couplets indicating progressively specific characteristics. If the organism falls into one category, you go to the next indicated couplet. By following the key and making the correct choices, you should be able to identify your organism. Constructing Your Own Dichotomous Key - Start with the most general characteristics and progress to increasingly more specific characteristics. Indent each couplet or leave a space between each couplet to make the key easier to read. Use constant characteristics rather than variable ones. Use measurements rather than terms like “large” and “small”. Use characteristics that are generally available to the user of the key rather than seasonal characteristics or those seen only in the field. Make the choice a positive one – something “is” instead of “is not”. If possible, start both choices of a pair with the same word. If possible, start different pairs of choices with different words. Precede the description with the part that is being described – “leaves are red” instead of “red leaves present”. SAMPLE DICHOTOMOUS KEY: IDENTIFYING CANADIAN MONEY 1a 1b Composed of Paper Composed of Metal Go to 2 Go to 3 2a 2b Sir John A. MacDonald is Pictured Sir Wilfred Laurier is Pictured $10 Bill $ 5 Bill 3a 3b Metals in Coin Appear Uniform More Than One Type of Metal is Visible Go to 4 Twoonie 4a 4b The Edges Have Ridges The Edges Are Smooth Go to 5 Go to 6 5a 5b Smaller than 20 mm in Diameter Larger than 20 mm in Diameter Dime Quarter 6a 6b Silver in Colour Colour Other Than Silver Nickel Go to 7 7a 7b Copper in Colour Brass in Colour Penny Loonie 1.3 – KINGDOMS and DOMAINS The Six Kingdoms - All plants, animals and other organisms can be organized into groups sharing similar characteristics. - Classification is a valuable way of learning how organisms are related to one another. - Through this we can make sense of the amazing diversity of life on Earth. - Over 2 000 years ago, Greek philosopher Aristotle, divided all living things into two Kingdoms, either Plantae or Animalia, based on certain characteristics. - A problem arose because some organisms had characteristics of both kingdoms (ex. corals and sponges that are non-motile but do not photosynthesize). - In the 17th century, when the microscope was invented, classify organisms became more difficult because a new world of organisms was revealed called microorganisms (ex: euglena that swim and photosynthesize). - In 1866, Ernest Haeckel proposed a new kingdom, Protista, one-celled organisms with a nucleus. - In the first half of the 1900’s, some single-celled organisms were found to be extremely small and without a cell nucleus, so a new kingdom, Bacteria, was created for them, bringing the total to four. - In the 1960’s, scientists added a fifth kingdom, Fungi because fungi were so different that they needed their own kingdom. - During the 1990’s bacteria that live in extreme environments because of their unique structures were researched and scientists classified these into a sixth kingdom, Archaea. - Figure 1.3, pg 24 Two Major Cell Types - There are two major types of cells: prokaryotic cells and eukaryotic cells. Prokaryote: organism with cells lacking a true nucleus and most other types of organelles; bacteria and archaea Eukaryote: organism with cells containing nuclei and other types of membrane-bound organelles; protists, fungi, plants and animals Differences Between Prokaryotes and Eukaryotes Prokaryotes Eukaryotes (bacteria, archaea) (protists, plants, fungi, animals) small (1 – 10 μm) large (100 – 1000 μm) DNA circular, not bounded by membrane DNA in nucleus bounded by membrane genome made up of a single chromosome genome made up of several chromosomes cell division not by mitosis and meiosis cell division by mitosis and meiosis asexual reproduction common sexual reproduction common multicellular forms rare most forms are multicellular mitochondria and other membrane-bound organelles absent mitochondria and other organelles present many are anaerobic (do not require oxygen) most are aerobic (require oxygen) - Fossil evidence shows that the first forms of life were prokaryotic organisms similar in appearance to bacteria. First prokaryotes appear 3.5 billion years ago. - The first eukaryotes appear 1.5 billion years ago (2 billion years later). - The first multicellular organisms appear 700 million years ago. The Three Domains - There are three domains: Bacteria, Archaea, and Eukarya. - Organisms in the domains Bacteria and Archaea are unicellular and prokaryotic. - Organisms in the domain Eukarya have eukaryotic cells and are unicellular or multicellular. There are four kingdoms in the domain Eukarya: Protists, Plantae, Fungi, and Animalia. Table 1.5, pg 29 summarizes some of main characteristics of kingdoms and Figure 1.17, pg 29 shows examples of organisms in each kingdom. HOMEWORK: pg 30 # 1 – 8 1.4 - Classifying Types of Biodiversity - Biodiversity is the variety of life in the world. It is often used to gauge the health of biological systems. Valuing Biodiversity - - Ecosystems that have a high biodiversity tend to be healthier and more resilient to changes in the environment. Biodiversity helps to maintain life-sustaining services, conditions, and processes: Insects, bats, birds, and other animals pollinate flowering plants and crops. Micro-organisms make nutrients available and break down toxic substances in water and soil. Ecosystems cycle carbon, nitrogen, and oxygen. Ecosystems clean air, purify water, control erosion, prevent floods, and modify climate. Often the presence or absence of a single keystone species can have a dramatic impact on an ecosystem. Keystone species are species that have a disproportionately large effect on the ecosystems in which they live, such as the honeybee. Discuss role of honeybee (Pearson, pg 5) Levels of Biodiversity - There are three levels of biodiversity: genetic diversity, species diversity, and ecosystem diversity. Genetic diversity genetic diversity: the variety of heritable characteristics (genes) in a population of interbreeding individuals - The variation among individuals in a population is largely a result of the differences in their genes. Genetic diversity within a population is known as the gene pool. In other words, the gene pool is the sum of all the versions of all the genes in a population. As genes determine hereditary characteristics, these small differences make each organism unique. They also allow populations to adapt to changes in environmental conditions and evolve over time. Genetic diversity is especially important in disease resistance. Populations that lack genetic diversity are more susceptible to disease than those that have high diversity. The entire population could be eliminated which could eventually lead to the extinction of the species. Genetic diversity also allows populations and species to survive changing environmental conditions, such as a change in resource availability, climate change, a change in a predator population, or the introduction of non-native species. Species Diversity species diversity: the variety and abundance of species in a given area - A greater variety of species in ecosystems allows those ecosystems to survive environmental changes, such as drought, plagues, or disease outbreaks. Each species has a certain set of conditions in which it can survive. Each species also contributes to the whole ecosystem and, without the actions of each species, the ecosystem and other organisms may not be as successful. Ecosystem Diversity ecosystem diversity: - the variety of ecosystems in the biosphere Recall that ecosystems are made up of two components – biotic factors (interacting populations of species) and abiotic factors (altitude, latitude, geology, soil nutrients, climate, and light levels). Ecosystems can range in size from a small plant that grows on another plant to an entire biome, such as a tropical rainforest or Canada’s boreal forest. Ecosystem services are the benefits experienced by organisms, including humans, which are provided by sustainable ecosystems. Earth would lose most of the services that ecosystems provide if there was no ecosystem diversity. Wetlands provide several important ecosystem services, including storing water (reduces the risk of floods), filtering water (removes pollutants) and providing habitat for commercially important species of fish and shellfish. Because wetlands are so valuable, government agencies and non-governmental organizations often work together to preserve and protect them. Ecosystem Function and Species Diversity - Ecosystems with greater species diversity have higher resilience. resilience: the ability of an ecosystem to remain functional and stable in the presence of disturbances to its parts - In experiments conducted at the University of Minnesota from 1982 to 1993, researchers concluded that greater biodiversity in an ecosystem results in at least three beneficial patterns: increased plant cover, more resistance to invasive species, and more disease resistance. (See Fig 1.21, pg 35) Ecosystem Services and Human Actions - In Ontario, bass-free lakes have been stocked with smallmouth bass. While this improves the bass population and recreational fishing, ecologists have noticed a decline in the number of native fish species such as stickleback and dace. This has a ripple effect on other species. With fewer small fish, trout must then consume less nourishing food, resulting in slower growth, smaller fish, and decreased populations. This leads to a decline in species diversity and affects the diversity of an ecosystem. Homework: pg 37 # 1 - 11