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SEED PRODUCTION AND TECHNOLOGY What is a seed? Seeds are mature, fertilized ovules. Ovules are structures of seed plants containing the female gametophyte with the egg cell, all being surrounded by the nucellus and 1-2 integuments. A seed (in some plants, referred to as a kernel), therefore, is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm (conifers and related families or clades) and angiosperm (flowering plants) which occurs after fertilization and some growth within the mother plant. The formation of the seed completes the process of reproduction in seed plants (started with the development of flowers and pollination), with the embryo developed from the zygote and the seed coat from the integuments of the ovule. A seed consists of an embryo, stored food and a seed coat. The term seed also has a general meaning that predates the above; anything that can be sown or any propagation material. For example in Irish potato, what is sown or propagated is tuber; in sweet potato, is vine; in cassava, is stem; in banana, is sucker; in ornamentals, is stem cuttings. Seeds are the principal means of regeneration of most woody plants. They serve as the delivery system for the transfer of genetic materials from one generation to the next. The part of a tree’s life cycle that involves seed formation, maturation, dissemination, and germination is a complex yet fascinating chain of events, many of which are still poorly understood. However, some knowledge of these events is necessary for successful collection, production and utilization of seeds. Importance of Seed Introduction Seeds have been an important development in the reproduction and spread of flowering plants, relative to more primitive plants like mosses, ferns and liverworts, which do not have seeds and use other means to propagate themselves. This can be seen by the success of seed plants (both www.naarocom.com Page 1 gymnosperms and angiosperms) in dominating biological ecological niches on land, from forests to grasslands both in hot and cold climates. Crop productivity Seeds are the principal means of regeneration of most annual and perennial flowering plants. They serve as the delivery system for the transfer of genetic materials from one generation to the next. Seeds play a pivotal role in agriculture. Increase in agricultural production is the key to economic growth in many countries. Seed constitutes the main propagule for plant growth. Modern/new agricultural information is disseminated through seed. For example, new seed is released with the recommended crop husbandry (from land preparation, planting, control of diseases, insects and weeds, harvesting, post harvest handling and marketing). Agricultural research findings are disseminated through seed. Seed is the main vehicle for the dissemination of plant pests. Seed-borne pathogens, such as fungi, bacteria and viruses are serious constraints to crop productivity. In worst-case scenario, seed-borne diseases can be disastrous and even life threatening. Infected seeds are at risk of being contaminated with mycotoxins and undergo changes in nutritional value. Seed-borne diseases can be harmful in several ways: 1. Seed-borne inoculum can reduce seed germination to various degrees, give rise to preand post emergence seedling mortality and progressive disease development in the field and thereby reduce the yield and quality of the crop 2. Imported seed lots carrying disease propagules may introduce devastating diseases or new strains of a pathogen into new regions 3. Infected seeds have reduced shelf life Nutritional and Economic www.naarocom.com Page 2 Many seeds are edible and the majority of human calories come from seeds, especially from cereals, legumes and nuts. Seeds also provide most cooking oils, many beverages (e.g. coffee) and spices and some important food additives. In different seeds, the seed embryo or the endosperm dominates and provides most of the nutrients. The storage proteins of the embryo and endosperm differ in their amino acid content and physical properties. For example, the gluten of wheat, important in providing the elastic property to bread dough is strictly an endosperm protein. Seeds are also eaten by animals, and are fed to livestock. Many seeds are used as chicken feed. Some seeds are also poisonous. One of the deadliest poisons, ricin, comes from seeds of the castor bean. Other poisonous seeds are those of yew, apple, chestnut and peach. The world's most important clothing fiber grows attached to cottonseed. Other seed fibers are from kapok and milkweed. Many important nonfood oils are extracted from seeds. Linseed oil is used in paints. Oil from jojoba is in the cosmetic industries. Seeds are the source of some medicines including castor oil, tea tree oil and the discredited cancer drug, Laetrile. Many seeds have been used as beads in necklaces and rosaries including, Chinaberry and rosary pea. However, the latter two are also poisonous. Resin from Clusia rosea seeds are used to caulk boats. www.naarocom.com Page 3 CHAPTER 1: REPRODUCTIVE PROCESSES IN PLANTS Introduction Plant growth originates within the buds in regions known as meristems. In the meristems, cell division and elongation occur, and these processes produce tissues that soon develop into specific plant parts. Vegetative meristems give rise to parts such as stems, leaves, and roots, while reproductive meristems give rise to floral organs that ultimately produce fruits and seeds. Within any meristem are minute primordia that resemble knobby outgrowths or ribbed inverted cones. Although hardly distinguishable to the naked eye, the configurations of the primordia become visible when the bud scales are removed and examined under magnification. As growth proceeds, the configurations enlarge and differentiate into recognizable plant organs. FLORAL INDUCTION Vegetative growth of plants occurs until they are ready to flower. At that stage, certain external stimuli can trigger floral induction, a physiological change that permits the development of reproductive primordia. This change may precede actual flowering by several days, weeks, or even months. Temperature Stimuli For floral induction to occur, many plants require exposure to low temperatures. This process has been called vernalization. In its narrowest sense, vernalization means the promotion of flowering in some winter cereals by cold treatment of the moistened or germinating seeds. In its broader sense, vernalization means the induction of flowering in any winter annual, biennial, or even perennial species through exposure to low temperature. For example, rye (Secale cereale), a winter annual, and perennial ryegrass (Lolium perenne) both must undergo prolonged exposure to low temperatures before they can produce flowers. Sugar beets and carrots are examples of biennial species that grow vegetatively the first year, after which they are vernalized by exposure to winter temperatures. www.naarocom.com Page 4 In the chrysanthemum and tomato, floral induction is accomplished by repeated exposure to low night temperatures, separated by periods of higher temperature. This phenomenon occurs in many plants and has been called thermoperiodism. Day-Length Stimuli In many species, floral induction occurs in response to day length, or photoperiod. Thus, plant species have been categorized according to their day-length requirements as short-day, long-day, intermediate-day, or day-neutral; however, it is really the length of the night, or dark period, that is the critical factor that influences flowering. Long-day plants include most cereal grains, clover, hibiscus, and Irish potato. Some short-day plants are chrysanthemum, tobacco, poinsettia, sweet potato and strawberry. Intermediate day plants include climbing hemp weed, wild kidney bean, Indian grass, and broom corn. Among the day-neutral plants are tomato, field bean, zinnia, dandelion, and chickweed. The photoperiod requirements for flowering may be qualitative or quantitative. Some short-day species such as the Biloxi variety of soybean are unable to flower except under short-day treatments; in other short-day species, such as sunflower, flowering is hastened by the appropriate short-day conditions, although it eventually occurs without them. Chemical Stimuli Certain natural and synthetic chemical substances can cause floral induction. Some are auxinlike compounds-for example, indoleacetic acid, naphthalene acetic acid, or the common herbicide, 2,4-dichlorophenoxyacetic acid (2,4-D). At certain concentrations, gibberellic acid may also cause floral induction. It promotes flowering of long-day plants held in short-day conditions; however, it inhibits flowering of short-day plants under the same conditions. Other substances known to cause flowering or to increase flower production are cytokinins, ethylene, acetylene and ethylene. In contrast, maleic hydrazide inhibits flowering. www.naarocom.com Page 5 This is a SAMPLE (Few pages have been extracted from the complete notes:-It’s meant to show you the topics covered in the full notes and as per the course outline Download more at our websites: www.naarocom.com To get the complete notes either in softcopy form or in Hardcopy (printed & Binded) form, contact us on: Call/text/whatsApp +254 719754141/734000520 Email: [email protected] [email protected] [email protected] Get news and updates by liking our page on facebook and follow us on Twitter Sample/preview is NOT FOR SALE www.naarocom.com Page 6