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Name _____________________________________________________ Test Date ________________ UNIT X - EVOLUTION I. THE THEORY OF EVOLUTION (pp. 369-386) The theory of evolution is one of the most fundamental concepts in Biology. Evolution is defined as ________________ in a _______________________________________________ over time. The scientist considered to be the founder of modern evolutionary theory is _____________________________________. A. History of Evolutionary Theory – During the 1700s, several scientists began challenging the idea of a world in which changes did not occur. These scientists and their hypotheses were very important to Darwin’s work. 1. Gradualism - _________________ first suggested that the planet was much ________________ than previously thought; began to find evidence that _________________ were slowly, but constantly taking place. 2. Malthus – Published an essay that had a huge impact on Darwin. Proposed that organisms ___________________ - reproduce; in other words, reproduce at a ______________________ rate than resources can supply. 3. Lamarck Lamarck was one of the first scientists to propose a mechanism for evolution; that is, the ___________ in a _________________________ of organisms over ____________. His major hypotheses included: Tendency Toward Perfection – Stated that organisms were continually changing in order to _____________________________________. Use and Disuse – Changes in ____________ and/or ______________ of a structure in an organism was a response to use or disuse. Structures used extensively ____________________ and structures used less frequently ________________________________________________. Inheritance of _________________ Traits B. Charles Darwin (1809 – 1882) 1. Darwin’s History Darwin’s data was collected on a 5-year journey around the world on the HMS ________________. He made observations and collected data throughout the journey. He used this data to propose a ______________________ to explain the diversity he saw. The area that had the greatest impact on Darwin was the _________________________ Islands due to the differences he saw in the same animals living on different islands. 2. Darwin’s Observations Members of a population often vary greatly in their ________________. Traits are inherited from __________________ to _________________. All species are capable of producing _________ offspring than ____________________________ Due to a lack of ____________ or other resources, many of the offspring ___________________. 3. Darwin’s Theories – Based on his observations and the hypotheses of other scientists Organisms with favorable _______________ tend to survive and ______________________; thereby leaving more descendents than other individuals This will result in an accumulation of these traits in the __________________________, changing the original make-up of the population 4. Darwin’s Legacy Did not publish his findings for years Alfred Wallace – formed identical _________________ based on his research. Sent his manuscript to Darwin, and finally Darwin was persuaded to publish his own conclusions Released “___________________________________________”, still considered one of the greatest scientific studies ever II. THE PRINCIPLES OF EVOLUTIONARY THEORY A. Evolution occurs because of natural selection - a mechanism for change that occurs when organisms with ______________ characteristics for a particular environment _______________________, ____________________, and pass these characteristics on to _______________________________. B. The ability of an organism to survive and reproduce in its environment is known as ________________. C. Fitness is based on _______________________. An adaptation is any trait that aids in the __________________ and _____________________ of an organism. Examples of adaptations are ___________________________________________________________________ D. As organisms ____________ and adapt, _________________ may occur. Speciation is the formation of new species - a group of similar organisms that ___________ with one another and produce __________________________________. Isolating Mechanisms allow for the gene pools to become ______________ so they can form a new ___________. Reproductive Isolation-as a new _______________evolves, population become _________________ isolated from each other. Behavioral Isolation-Population is capable of ________________________, but have different ______________________________ rituals. EX: _____________________________________ Geographic Isolation-__________ populations are separated by geographic barriers and evolve where they can no longer _________________________. EX: ____________________________________ Temporal Isolation-______________________ reproduce at different times even though capable. EX:__________________________________________________________________________ E. The failure of an organism to _______________ to changes in its environment will ultimately lead to its _______________ because of _________________________________. III. EVIDENCE FOR EVOLUTION A. Fossils – Fossils are _________________________________________________________________ Fossils provide a record of earlier life and evidence that evolution has occurred. B. Biogeography – Variations are seen in the same types of animals based on their _________________. In addition, there are some organisms that live in very different locations but they have similar characteristics because _________________________________________. C. Homologous Structures – Scientists use anatomical studies of different organisms for evidence of evolutionary relationships. For example, appendages that are very similar in structure, but differ in function are known as _______________ structures. Examples of homologous structures are _________________________________________________________________________________. D. Vestigial Structures - A structure that is reduced in function in a living organism, but may have been used by an ancestor is known as a ________________ structure. A structure may become vestigial when an organism changes in form or behavior. Examples are _____________________________________________________. E. Embryology – Similarities in the structures of developing _______________ of different organisms are considered to be proof of a close evolutionary relationship. F. DNA Sequencing – Scientists use DNA studies to determine the evolutionary relationship between organisms. The more similar the DNA, __________________________________________________ IV. MECHANISMS FOR EVOLUTION Evolution does not occur in an individual; instead it refers to _______________ __________________ that occur in a _________________ over time. There must be mechanisms available for _________________ changes to occur: A. Mutation – A mutation is a __________________________________. Although mutations are most often ______________________, sometimes the resulting change in ______________________ may be beneficial to an organism under certain conditions. If the change occurs in the ________________, this change will be passed onto the offspring of that organism. A positive mutation that provides a survival advantage is known as an __________________________. B. Diploidy – Most organisms are diploid, which means ________________________________________. This allows for increased genetic variation in a population. Heterozygote Advantage – Seen in _______________________________ and _______________. Recombination – Leads to increased genetic variation as a result of _____________________________ during ___________________________ of ______________________. C. Gene Flow – Gene flow occurs when organisms from one community migrate to another. This introduces new _________________ into the ____________________ which can lead to a change in the genetic make-up of the population. D. Genetic Drift – This describes a situation in which change in a population is magnified because the population size is very ______________. Causes of genetic drift include Bottleneck effect – large portion of population ______________________________________ Founder effect – segment of population moves to new _______________ E. Single Gene Traits-Natural selection on __________ ________ traits can lead to changes in ___________ frequencies and therefore to evolution. F. Polygenic Traits-Natural selection can affect the distributions of phenotypes in 3 ways: Directional Selection-Individuals at ____________ end of the bell curve have higher fitness than those in the middle or other end. EX:___________________________________________________________________ Stabilizing Selection-Individuals in the _______________ have the highest fitness causing the curve to narrow. EX:____________________________________________________________________________________ Disruptive Selection-when individuals at the ends of the curve have the higher fitness. If lasts long enough can cause the curve to split in two and create _____ distinct phenotypes. EX: _____________________________ V. MACROEVOLUTION- refers to large-scale evolutionary pattern and processes that occur over long periods of time. A. Extinction-More than _______% of all species that have ever lived are now ___________________, which means the species has died out. Darwin proposed possible reasons with competition for ____________________________________________________________________________________________ _____________extinction has occurred several times, wiping out entire _______________________. ____________________, volcanic action, and __________________________movement has been blamed. Mass extinctions clear the way for _______________________ of other species. EX:____________________ B. Adaptive radiation-the process where a _______________species has evolved through _________________________ into diverse forms that live in ___________________________ways. EX: ____________ C.____________________ _____________________can have an evolutionary “side effect”, producing unrelated species that ________________ similar to one another. This process is known as ______________________________________ and occurs in ________________________ and animals. EX:_______________________________________________ D. ______________________________ is when a change in one organism leads to a ____________________________ change in another organism. EX: ______________________________________________________________________ E. _________________________ Equilibrium is another pattern of evolution. Unlike ___________________________ punctuated equilibrium is characterized with long periods of _________________________ interrupted by brief periods of _________________change. It is controversial but known that evolution does occur at different rates. VI. Modern Evolutionary Classification-Linnaeus tried to group organisms according to biologically important characteristics, but his system had limitations and problems. A. Darwin’s ideas about descent with modification gave rise to the study of _________________________, or evolutionary relationships among organisms. Biologists now group organisms into categories that represent lines of ______________________________, or phylogeny, not just physical similarities. B. Cladistic Analysis-identifies and considers only those characteristics of organisms that are ___________________________________________ -new characteristics that arise as lineages evolve over time. These recent characteristics are called ____________________________________________. C. Derived characters can be used to construct a _____________________________________, a diagram that shows the evolutionary relationships among a group of organisms. Cladograms-are useful tools that help scientists understand how one lineage ____________________________ from another in the course of evolution. Think of them as a family tree showing _______________________________relationships among a group of organisms. D. Similarities in Genetic Material _________ and _________ are so similar across all forms of life, these molecules provide an excellent way of comparing organisms at their most basic level-____________________________________________. ____________________________________ in DNA can be used to help determine classification and _____________________________________relationships. DNA_________________________________ using ________________________________________ is a technique that can be used to analyze DNA and determine these evolutionary relationships. ________________________________________________ or reading of the DNA of organisms allow for comparison of the DNA of different organisms to trace the history of genes over ________________ of years. E. _______________________________- Comparisons of ___________can also be used to mark the passage of evolutionary____________________________. A model known as a molecular clock uses DNA comparison to ________________________ the length of time that_____ species have been evolving _________________________________________________. The molecular clock depends on a repeating process to mark time-___________________________________. ________________________ mutations (ones that cause no changes) accumulate in the DNA of________________________________________ species at about the same rate. Comparing these sequences, with these mutations, can tell scientist how _______________________ the genes are and in turn how long ago the two species shared a __________________________________________.