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THE SENSES CHAPTER 9 Receptors and Sensations Five general types of receptors are recognized: 1. Chemoreceptors: Are sensitive to changes in chemical concentrations. Sense of smell and taste fall into this category. 2. Nociceptors: These are pain receptors that detect tissue damage. 3. Thermoreceptors: Sensitive to temperature changes. 4. Mechanoreceptors: Respond to changes in pressure or movement. 5. Photoreceptors: Respond to light energy. SENSATIONS Sensations are feelings that occur when the brain receives sensory impulses from the peripheral nervous system. The impulses are relatively the same; it is the brain’s interpretation that makes them seem different and unique. SOMATIC SENSES Somatic senses involve receptors associated with the skin, muscles, joints, and visceral organs. If they occur at the body surface, they are called exteroceptive senses and include touch, pressure, and temperature. Sensations associated with changes occurring in visceral organs are called visceroceptive senses. TOUCH AND PRESSURE SENSES Touch results from sensation of tactile receptors just beneath the skin. Pressure sensations result from stimulation of receptors in deeper tissue. Each sensation is triggered by a degree of displacement of tissues involved. Types of Touch/Pressure Sensors Three types: 1. Free ends of Sensory Nerves 2. Meissner’s Corpuscles are receptors for light touch. They are also used when a person wants to judge the texture of something. Are found in the dermal papillae of the skin. Are abundant in hairless areas like lips, clitoris, palm. 3. Pacinian Corpuscles: Detect deep pressure or vibrations. Located in subcutaneous tissues, deep submucosal tissues, and serous membranes. Found around joints, tendons, muscles, mammary glands, external genitalia, pancreas, and urinary bladder. Temperature Senses Thermal sensations are perceptions of degrees of warmth and coolness (extreme heat or cold are experienced as pain). Receptors include two groups of free nerve endings located in the skin: 1. Heat Receptors are sensitive to temperatures above 77F and become unresponsive above temperatures of 113F – pain receptors kick in and report the sensation of burning. 2. Cold receptors respond to temperatures between 50F and 68F. If temperatures are below 50F, a freezing sensation is reported by pain receptors. Both heat and cold receptors adapt quickly. SENSE OF PAIN Pain receptors consist of free nerve endings that are stimulated when tissues are damaged. Protect the body by signaling damage and identifying changes that may cause longterm damage. Aid medical professionals in diagnosing specific diseases or malfunctions of the body. Pain receptors, called nociceptors, are widely distributed throughout the skin and internal tissues. Tissue injury releases chemicals called prostaglandins and kinins that stimulate nociceptors. Pain persists after initial tissue injury because these chemicals linger and there is little to no adaption for these receptors. PAIN – ACUTE OR CHRONIC ACUTE: Occurs very rapidly and is typically not felt in deeper tissues of the body. Is usually described as sharp or stabbing pain. Acute pain fibers are thin, myelinated fibers that carry impulses rapidly and cease when the stimulus stops. CHRONIC: Has a slower onset and builds slowly in intensity over a period of seconds or minutes. Often described as burning, aching, or throbbing. Chronic pain fibers are thin, unmyelinated fibers that conduct impulses slowly and continue sending impulses long after the stimulus stops. VISCERAL PAIN Visceral pain receptors are the only receptors in the viscera that produce sensations. Response of pain receptors is quite different from those associated with surface tissues. Localized stimulation of the viscera does not evoke much of a response, but widespread stimulation may elicit a strong response. Stretching of visceral tissues or a decrease in blood flow elicits the most painful responses. Visceral pain often seems to come from somewhere other than its actual origin. This is called referred pain. It occurs because of common nerve pathways: the two areas are served by the same segment of the spinal. Example, an MI is felt as pain in the arm. Stretch Receptors Stretch receptors are proprioceptors that deal with sensations from the lengthening and stretching muscles. The two main types are Golgi tendon organs and muscle spindles. Golgi Tendon Organ: Found in tendons at the point of attachment to their respective muscles. Are stimulated by increased tension in the skeletal muscle. Golgi Tendon Organ (Cont’d) Sensory impulses from these receptors produce an inhibiting reflex for muscle contraction. This helps to prevent the muscle from pulling away from its insertion point and to maintain posture. MUSCLE SPINDLES Muscle spindles are located in muscles near the origin of the tendons that serve them. Each spindle is composed of 3-10 specialized muscle fibers called intrafusal muscle fibers. The central region of the spindle has few or no actin or myosin filaments, but the ends do contain a few. This central region cannot contract. However, there are two types of afferent (sensory) fibers inside the spindle – they have dendrites that wrap around the central area of the intrafusal fiber. When stretched, the spindle stimulates the dendrites, which send a nerve impulse toward the spinal cord. Muscle spindle fibers respond to the rate and degree of length change in a skeletal muscle. The info is transmitted to the cerebrum for perception of limb position and to the cerebellum for coordination of muscle contraction. Structures of the Eye ADNEXA – accessory structures Orbit Eye Muscles Eyelids Eyelashes Conjunctiva Lacrimal Apparatus www.ipo.tue.nl/.../seminar-2%20human/ sld015.htm Pg. 195 The Lacrimal Apparatus – structures that produce, store, and remove tears Lacrimal Glands: secrete lacrimal fluid to maintain moisture on the anterior surface of the eyeball Lacrimal Canaliculi: ducts at the inner canthus of each eye – collect tears and drain them into the lacrimal sac Lacrimal sac: an enlargement of the upper portion of the lacrimal duct Lacrimal duct: passageway that drains lacrimal fluid into the nose Functions of the Eyes – receive images and transmit to brain Optic: pertaining to eye or sight Ocular: pertaining to the eye Extraocular: outside the eyeball Intraocular: within the eyeball ORBIT Frontal Sphenoid Ethmoid Maxilla Zygoma Lacrimal Palatine www.lau-verlag.de/anatom/ skeletal-system.htm Muscles of the Eye Superior/Inferior Rectus Superior/Inferior Oblique Lateral/Medial Rectus Eyelids: protect the eyeball form foreign matter, excessive light, and impact Canthus: angle where upper/lower eyelids meet Inner Canthus Outer Canthus Conjunctiva: lines the underside of each eyelid and provides protective covering over exposed surface of eyeball The Eyeball globe www.optelec.com/ lv_ref.php Made up of 3 layers Sclera Choroid Retina Interior of eye is divided into anterior/posterior segments The Sclera – the white of the eye Outer layer of eye Maintains shape of the eye and protects the delicate inner layers of tissue Cornea: transparent anterior portion of the sclera Provides most of the optical power of the eye The Uveal Tract The vascular layer of eye Choroid: opaque (light cannot pass through it) middle layer of the eyeball – provides blood supply for entire eye Iris: colored layer that surrounds pupil – it’s muscles control amount of light entering eye – decrease muscles contract making opening smaller (visa versa) Pupil: black circular opening in center of iris – permits light to enter eye Lens: focuses images on the retina – located behind iris and pupil Ciliary Body: located within the choroid, set of muscles and ligaments that adjust lens to refine the focus of light rays on the retina Near-by objects = thicker Distance objects = thinner Nerve layer located between the posterior chamber and the choroid layer at the back of the eye Contains light sensitive rods (black/white receptors) and cones (color receptors) Receive images and convert them into nerve impulses The Retina Optic disk: (blind spot) – contains no rods/cones – nerve endings of retina gather to form optic nerve which transmits nerve impulses from the retina to the brain Retina Macula lutea – In the central region of the retina lies a yellowish spot called the macula lutea. Within the center of this structure lies a small depression called the central fovea. This region of the retina is the area of the highest visual acuity. Visual Pathways Optic Chiasma – As the axons of the ganglion cells of the retina leave the eye, they converge to form the large optic nerve. These nerves eventually cross at the base of the brain (just in front of the pituitary gland) in a formation referred to as the optic chiasma. Nerve fibers continue into the posterior portion of the thalamus of the brain and enter nerve tracts that “radiate” to the occipital lobe of the brain. Some fibers continue from the lateral geniculate nucleus to the brainstem where they contribute to simultaneous eye movements and control of visual reflexes. The Anterior Segment – front 1/3 of eye Divided into anterior and posterior chambers Anterior chamber Posterior chamber behind the cornea in front of the iris Between the back of the iris and the front of the lens These chambers filled with aqueous humor (fluid) Nourishes intraocular structures Constantly filtered and drained which regulates intraocular pressure (IOP = btwn 12 & 21 mm Hg) The Posterior Segment – posterior 2/3 of eye Aids in maintaining the shape of the eye Contains vitreous humor Lined with retina and it’s related structures Normal Action of the Eye Accommodation: the eyes make adjustments for seeing at various distances Includes constriction or dilation of the pupil, movement of the eye, and changes in the shape of the lens Convergence: simultaneous inward movement of both eyes – in an effort to maintain single binocular vision as an object comes nearer Visual Acuity: the ability to distinguish object details and shape at a distance Normal vision = 20/20 Snellen Chart: used to measure visual acuity Summary of Sight The eye functions somewhat like a camera: it receives and focuses light on a photosensitive receiver, the retina. Light rays are bent (called refraction) and brought to focus as they pass through the cornea and the lens. Once an image is focused on the retina, stimulated photoreceptors alter the light signal into receptor potentials. Pathology of the Eyes EYELIDS Blepharoptosis (A) Ectropion (B) Entropion (C) Hordeolum: (D) A C B D Functions of the Ears Receive sound impulses and transmit them to the brain Inner also helps maintain balance Auditory: pertaining to the sense of hearing Acoustic: relating to sound or hearing Structures of the Ear The Outer Ear The Middle Ear The Auditory Ossicles The Eustachian Tubes The Inner Ear The Outer Ear Pinna: auricle – External portion Catches sound waves and transmits them into the external auditory canal (EAC) EAC: transmits sound waves from pinna to middle ear What is the name for the sticky yellow-brown substance that functions to prevent bacteria and dust from entering the middle ear?? CERUMEN (earwax) The Middle Ear Tympanic Membrane (eardrum) Transmits sound by vibrating Surrounded by hollow air spaces – mastoid cells (which can easily become involved in a middle ear infection) The Auditory Ossicles 3 small bones in the middle ear Transmit sound waves from the eardrum to the inner ear by vibration Named for their shape Malleus: hammer Incus: anvil Stapes: stirrup The Inner Ear- Contains sensory receptors for hearing and balance labyrinth Cochlea: spiral shaped passage that leads from the oval window Cochlear duct: filled with fluid that vibrates when sound waves strike it Organ of Corti: receptor site that receives vibrations and relays them to the auditory nerve fibers that transmit them to the auditory center of the cerebral cortex, where they are interpreted and heard Semicircular canals: helps maintain equilibrium Normal Action of the Ears Air conduction: sound waves enter the ear through the pinna, travel down the auditory canal, and strike the TM between the outer and middle ear Bone conduction: as the eardrum vibrates, it moves the auditory ossicles and these conduct sound waves through the middle ear Sensorineural conduction: sound vibrations reach the inner ear via the oval window where the structures of the inner ear receive the sound waves and relay them to the brain Sense of Equilibrium Sense of equilibrium consists of two parts: Static and Dynamic. Static equilibrium helps to maintain the position of the head and body when they are still. Dynamic equilibrium maintains balance when the head and body suddenly move and rotate. Static Equilibrium Organs are located within the bony vestibule of the inner ear, inside the utricle and saccule (expansions of the membranous labyrinth). Hair cells lie inside the utricle and saccule. Gravity causes the gelatin and otoliths to shift, bending hair cells and generating a nervous impulse. Impulses travel to the brain via the vestibular branch of the vestibulocochlear nerve, indicating the head position Dynamic Equilibrium Three semicircular canals detect motion of the head and aid in balancing the head and body during sudden movement. Organs involved are called cristae ampullaris and are located in the ampulla of each semicircular canal of the inner ear. Rapid turning of the head or body generates impulses as the cupula and hair cells bend, which are interpreted and modified by the brain to maintain specific positions for the body. Mechanoreceptors associated with the joints and the changes detected by the eyes also help maintain equilibrium Pathology of the Ears Outer Ear Middle Ear Eustachitis: inflammation of the eustachian tube Mastoiditis Otosclerosis: ankylosing of the bones of the middle ear = hearing loss Otitis Media Inner Ear Otalgia: earache Otitis: inflammation of the ear Otomycosis: swimmer’s ear Meniere’s syndrome: vertigo, fluctuating hearing loss, tinnitus (ringing or buzzing in the ears) Hearing Loss Deafness Noise-Induced hearing loss Diagnostic Procedures of the Ears Audiometry: use of audiometer to measure hearing Speech audiometry: measures the threshold of speech reception and speech discrimination Monaural: testing involving one ear Binuaral: testing involving both ears Treatment Procedures of the Ears Outer Ear Middle Ear Otoplasty: surgical repair of the pinna of the ear Mastoidectomy Tympanocentesis Tympanostomy tubes Inner Ear Fenestration Labyrinthectomy Labyrinthotomy DEAFNESS Deafness is defined as any reduction of hearing and is categorized as follows: conduction-type deafness, sensorineural deafness, central deafness, mixed-type deafness, functional deafness, congenital deafness, and neonatal deafness. Conduction-Type Deafness Occurs when there is an interference with the transmission of sounds from the external or middle ear, preventing sound waves from entering the inner ear. Sensorineural Deafness Also referred to as “nerve deafness” and involves the cochlear portion of the inner ear of the cochlear division of the vestibulocochlear nerve. Central Deafness Involves the acoustic center of the cerebral cortex. Mixed-Type Deafness Involves both the conduction system and the nervous system. Functional Deafness Sometimes called “selective” deafness. Is psychogenic, with no conduction or nerve problem identified. Deafness (continued) Congenital Deafness Present at birth. Cause may be hereditary or may be due to the mother’s exposure to disease or toxic drugs during the pregnancy. Neonatal Deafness Occurs at the time of birth. Is caused by prematurity, trauma, or Rh imcompatability. SENSE OF TASTE Special organs of taste are the taste buds. They are primarily on the surface of the tongue and are located within tiny, numerous elevations called papillae. Each taste bud consists of cells called gustatory cells that function as receptors. Chemicals must be dissolved in saliva that surrounds the taste cells in order to be tasted. Four Types of Taste Buds Each taste bud is stimulated by a specific type of chemical that produces a certain taste sensation: sweet, sour, salty, and bitter. (Some recognize two additional sensations: alkaline and metallic) Areas of taste receptors on tongue: Sweet – tip of tongue Sour – lateral edge of tongue Salt – tip and upper portion of tongue Bitter – Back of tongue Taste impulses from the anterior two thirds of the tongue are relayed to the brain via the facial (VII) nerve. Taste impulses from the posterior one third of the tongue and back of the mouth are relayed via the glossopharyngeal (IX) nerve. Taste impulses from the base of the tongue and the pharynx are relayed by the vagus (X) nerve. Impulses from these nerves are transmitted to the medulla oblongata, then to the thalamus, and finally to the gustatory cortex of the parietal lobe. SENSE OF SMELL Sense of smell is referred to as olfactory senses. Humans can distinguish approximately 10,000 chemicals. Nerve pathways for the sense of smell are directly connected to older, more primitive areas of the brain (limbic system) that are associated with memory and basic primal instincts. For example, a whiff of perfume can bring up memories of an old girlfriend. Smell is an important component for attraction and sexuality. Olfactory senses and taste operate together to aid in food selection. Smell accounts for 90% of what we think we taste. Mechanics of Smell The nasal cavity has a patch of tissue the size of a postage stamp with specialized olfactory receptors. To be detected, chemicals that enter the nasal cavity must be in a gaseous state and must be dissolved in the watery fluid surrounding nasal cilia. Once olfactory receptors have been stimulated, impulses are transmitted along axons of the receptor cells of the olfactory nerves. The impulse is then transmitted to the olfactory tract and then to interpreting centers in the base of the frontal lobes. The impulse then travels along olfactory tracts to the limbic system, and lastly to the olfactory cortex within the temporal lobes Objectives: Describe the functions and structures of the eyes and adnexa Recognize, define, spell, and pronounce terms related to the pathology and diagnostic and treatment procedures of eye disorders Describe the functions and structures of the ears Recognize, define, spell, and pronounce terms related to the pathology and diagnostic and treatment procedures of ear disorders