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Outline Origin of life Major events in the history of life Classification of organisms Life on Earth Evolved Once Same Macromolecules -Same amino acids, universal code -Same genetic code -Other macromolecules are the same Cell structure is similar - plasma membrane contains genetic material Fig 18_9 Major events in the history of Life 200 kya 1.5 bya 2.5 bya Fig 19_3 Hypothetical Origins of Life: 1 - Special creation - supernatural power created life Not testable 2 - Extraterrestrial origin - life originated elsewhere, traveled to earth Difficult to test 3 - Evolution - life evolved on earth from materials on earth Testable Each are valid hypotheses, will never know which is correct without direct observation of the actual event Testing the hypothesis that life evolved on earth Miller-Urey experiment recreated conditions of early earth Fig 19_2 Results of Miller-Urey and similar experiments -Readily form all amino acids, nucleic acids found in life, plus some -Altering gases increases diversity of molecules produced -Long chains of amino acids degrade in water -Easy to form RNA chains which can act as enzymes - can duplicate selves (inheritance) and catalyze other reactions Wikipedia -Lipids form small spheres (cells) in turbulent water - act like cell membranes Plausible scenario: correct combination of organic molecules trapped in lipid sphere, persisted and replicated We will never know for certain how life arose, plausible scenarios are best guesses Stromatolites - representatives of the earliest organisms Modern stromatolites in Austrailia Fossilized stromatolites -Mats of bacteria, trap sediment -Date back 2.75-3.45 billion years -Sharp fall off in the fossil record 1 billion years ago When life arose, organisms changed the environment “morphological” evolution “metabolic” evolution Banded Iron Formations - 2.5-1 billion years ago -Formed from Iron oxydized by O2 -Becomes insoluble in water and precipitates out The problem with oxygen….. -Life arose in, and was like adapted to, anoxic conditions -Oxygen can combine with atmospheric gases and eliminate green house gases Mass extinctions in the fossil record The “cambrian explosion” -“sudden” appearance of all the major groups of animals 560-530 MYA -Presents problem for “gradualism” mode of evolution -Lead Wolcott to hypothesize a time period preceeding the cambrian that was under represented in the fossil record Modern life on earth arose from a single common ancestor, at least 2.5 bya There are multiple hypothetical origins of life Origin of life via organic evolution is plausible Once life arose, it changed earths environment into what it is today Massive extinctions happen repeatedly Animals make a near sudden appearance in the fossil record Millions of kinds of organisms exist, interact with humans Naming/classifying avoids confusion Aristotle, Romans used long descriptive names - changed frequently Linnaeus used simple, unique, fixed names - Genus species Scientific names are hierarchical -Levels of classification (genus, family, order, etc.) are inclusive -Organisms with similar characteristics are grouped together at each level QuickT ime™ and a TI FF (Uncompressed) decompressor are needed to see this picture. Fig 18_3 QuickTi me™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Fig 18_9 Shared characteristics are due to shared evolutionary history Phylogenies are evolutionary family trees QuickT ime™ and a TI FF (Uncompressed) decompressor are needed to see this picture. Fig 18_5 Phylogenies are built using shared characters A phylogeny is a hypothesis of relationships 2 out of 3 Domains are prokaryotes Bacteria: Archea: -Most well known prokaryotes -Poorly understood -Many cause disease -Well known are “extremophiles” -Cyanobacteria, various heterotrophs -Methanogens, extremophiles Prokaryotes are the dominant life form -A spoonful of soil contains ~ 2.5 billion prokaryotic cells -There are more prokaryotic cells living on your body than cells within your body Nodules in soybean roots house nitrogen fixing bacteria -Multicellular life depends on prokaryotic life for many processes such as decay and nitrogen fixation Bacteria can obtain energy and materials by metabolizing many types of materials Oil metabolizing bacteria Fig 19_6 Cyanobacteria are responsible for atmospheric oxygen -Evolved photosynthesis -Altered environment from “reducing” to “oxidizing” Fig 19_8 Stromatolites in Shark Bay, Australia (some of the oldest fossils are stromatolites) -Atmosphere changed from trace amounts of oxygen to 21% oxygen, which radically altered the course of evolution -Still responsible for 20-30% of oxygen Heterotrophic bacteria contribute to decay Obtain materials and energy from organic sources Help recycle materials Animals and Fungi are also heterotrophs and contribute to decay Archea diversity Extremophiles in Yellowstone Fig 19_7 Methane (natural gas) is produced from methanogens in hypoxic environments Methanogens produce methane gas from H rich sources (chemoautotrophs) Eukaryotes compared to Prokaryotes