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Indianpipe –Monotropa uniflora Tetraphis pellucida Range size and abundance • Ecologists have long suspected that a species that is widespread in its distribution may tend to be more common or abundant in its population size • What causes such patterns? • 1. sampling error • 2. ecological specialization • 3. local population model Biomes • A Biome is a large, relatively distinct terrestrial ecosystem characterized by particular climate, soil, plants and animals, regardless of where it occurs on earth • A biome's boundaries are determined by climate more than by any other factor Tundra Tundra • Tundra occurs in extreme northern latitudes where the snow melts seasonally (southern hemisphere has no equivalent) • Tundra has long, harsh winters and very short summers - growing season is 50 to 160 days depending on location – summer days are long • Tundra soils are young - formed after last ice age, nutrient poor, little organic matter - have permafrost - permanently frozen layer of soil Taiga Taiga • Taiga - evergreen forests of the north • Taiga is found in northern America, Asia and Europe - covers 11% of land • Winters cold and severe - growing season longer than tundra - so greater than 160 days - little precipitation, about 50 cm (20 inches) per year • Soil is acidic, mineral-poor, deep layer of poorly decomposed pine and spruce needles at surface • Numerous lakes and ponds Temperate rain forests Temperate rain forests • Occurs on northwest coast of North America, western Australia, southern South America, Japan • very high precipitation - 200 to 380 cm (80 to 152 inches), much condensation from coastal fogs little seasonal temperature variation - winters mild and summers cool • Nutrient poor soil, but may have high organic content Temperate deciduous forests Temperate deciduous forests • Occur where temperate zone precipitation ranges from 75 to 125 cm (30 to 50 inches) • Typically have hot summers and cold winters • soil has rich topsoil with much organic matter, deep clay-rich lower layer Temperate grasslands Temperate grasslands • Occur where annual precipitation is 25 to 75 cm (10 to 30 inches) • Summers hot, winters cold, rainfall is often sporadic • soil is rich in organic matter, minerals build up in top layer of soil, dead organic matter from annual die-back of grasses builds up soil organics, grass roots may form turf or sod • shortgrass prairies may be called steppes more rainfall than deserts but not much - 10 to 15 inches Mediterranean ecosystems Mediterranean ecosystems • Mediterranean - thickets of evergreen shrubs and small trees • some temperate areas have climates with mild winters with abundant rainfall and very dry summers - called Mediterranean climates - occur around Mediterranean, California, western Australia, Chile, South Africa • In California we call this ecosystem chaparral - soil is very thin and nutrient poor • Fires are common, frequent in late summer, early autumn • plants usually dense growth of evergreen shrubs, but may have short scrubby pines and oaks - often have sclerophyllous leaves - hard, small, leathery leaves that resist water loss Deserts Deserts • Deserts are very dry habitats of both temperate and tropical zones - low water content of the desert atmosphere leads to wide daily temperature range • deserts vary greatly depending upon amount of precipitation - usually less than 25 cm (10 inches) per year • desert soil poor in organics, rich in minerals • some deserts so dry have virtually no plant life Namib and Atacama desert Tropical savanna Tropical savanna • Savannas are a tropical grassland which typically has widely scattered trees • either low rainfall or seasonal rainfall with prolonged dry periods - often 85 to 150 cm (34 to 60 inches) • Temperature doesn't change much - often in 80's or 90's • Savanna soil is low in mineral nutrients • Savanna found in Africa, South America and northern Australia Tropical rainforest Tropical rainforest • Tropical rain forest occurs where temperatures are high throughout the year and precipitation occurs almost daily - annual precipitation may be 200 to 450 cm (80 to 180 inches) - much of the rainfall is locally recycled as water that enters atmosphere from transpiration quickly falls out again • Soil is usually very old and nutrient poor, low in minerals and organics - decomposition and recycling is very quick due to high temperature and high availability of water • extremely diverse forests Distribution of Biomes in Current Climate Possible Distribution of Biomes in Climate with CO2 Doubled