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Flowers, Fruits, and Seeds Chapter 8 Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display Outline • • • • • Dicots versus Monocots Structure of Flowers Fruits Fleshy Dry Fruit and Seed Dispersal Seeds Germination Longevity Dicots versus Monocots • • • • • • • Dicots (Magnoliopsida) Two cotyledons Flower parts in fours or fives Leaves with distinct vein network Vascular cambium present Vascular bundles in ring Pollen grain with three apertures • • • • • • • Monocots (Liliopsida) One cotyledon Flower parts in threes Leaves with parallel primary veins. Vascular cambium absent Vascular bundles scattered Pollen grain with one aperture Structure of Flowers • Each flower, which begins as an embryonic primordium that develops into a bud, occurs as a specialized branch at the tip of a peduncle which may have branchlets of pedicles. Pedicle swells at its tip into a small pad (receptacle). - Other parts of the flower are attached to the receptacle. Structure of Flowers • • Outermost whorl typically consists of three to five sepals. Sepals (calyx) may be fused together. Next whorl consists of three to many petals (corolla). Calyx and corolla form the perianth. Structure of Flowers • Several to many stamens are attached to the receptacle around the base of the pistil. Each stamen consists of a filament with an anther at the top. - Pollen grains developed and disseminated in anthers. Structure of Flowers • Pistil consists of Stigma, Style, and Ovary. Superior Ovary - Calyx and corolla are attached to the receptacle at the base of the ovary. Inferior Ovary Receptacle grows up and around the ovary. - Calyx and corolla appear to be attached at the top. Gymnosperms flowers Sago palm, however not a true palm: The generic name comes from Greek Koikas, and means "a kind of palm". About 95 species are currently accepted in the cycad family Cycadaceae. Gymnosperms flowers Seed conesfemale strobili Polen conesmale strobili Pollen grains Monocot flowers Dicot flowers Structure of Flowers • Inflorescences - Group of several to hundreds of flowers Fruits • Fruit is an ovary and its accessory parts that have developed and matured. Usually contains seeds. All fruits develop from flower ovaries and accordingly are found exclusively in flowering plants. Fruits • Fruit Regions Exocarp - Skin Endocarp - Inner boundary around seed(s). Mesocarp - Fleshy tissue between exocarp and endocarp. - Three regions are collectively called the pericarp. Fruits • Fleshy Fruits Simple fleshy fruits develop from a flower with a single pistil. - Drupe - Simple fleshy fruit with a single seed enclosed by a hard, stony endocarp, or pit. E.g,; coconuts,apricot, peach,plum, almond Fruits • Berry - Usually develops from a compound ovary and often contains more than one seed. Three types of berries: True berry is a fruit with a thin skin and a relatively soft pericarp.E.g.;tomatoes, grapes, peppers, eggplants, blueberries, cranberries, pomegrantes. Fruits Pepos - Relatively thick rinds (Pumpkins). Fruits Hesperidium – Berry with a leathery skin (citrus family) Fruits • • Pomes - Bulk of flesh comes from enlarged floral tube or receptacle that grows up around the ovary. (Apples) –Acessory fruits: Fruits developed from more than an ovary alone i.e. Pomes, pepos and some berries Fruits • Aggregate Fruits Derived from a single flower with several to many pistils. - Individual pistils mature as a clustered unit on a single receptacle Raspberries, Strawberries. Aggregate fruits/Raspberries, blackberries Aggregate fruits/Raspberries, blackberries Individual pistils Fruits • Multiple Fruits Derived from several to many individual flowers in a single inflorescence. - Pineapples, Figs, Mulberries Multiple fruits/Pineapples, figs Fruits • Dry Fruits That Split at Maturity (Dehiscent) Follicle - Splits along one side or seam (Milkweed). Legume - Splits along two sides or seams (Beans, peas,garbanzo beans, lentils,carob, peanuts!). Silique - Splits along two sides or seams, but seeds are borne on central partition, exposed when the two halves separate (Cabbage, broccoli). Capsules - Consist of at least two carpels, and split in a variety of ways (Poppies, Lilies). Follicles-Milkweed) Silicles-Lunaria (Dollar plant) Capsule-Orchid Fruits • Dry Fruits That Do Not Split at Maturity (Indehiscent) Achene (Sunflower) Nut (filberts=Hazelnut) Grain (Poaceae fam.) Samara( ashes, elms) Schizocarp (parsley, carrots) Fruits • Dry Fruits That Do Not Split at Maturity (Indehiscent) Schizocarp: Twin fruit (parsley, carrots, anis, dill). Upon drying twin fruits break into two one-seeded segments called mericarps Fruit and Seed Dispersal • • • • Wind Dispersal Small and Lightweight seeds. Animal Dispersal Seeds pass through digestive tract. Fruits and seeds catch in fur or feathers. Oils attract ants. Water Dispersal Some fruits contain trapped air. Mechanical Ejection of Seeds Seed dispersal Pine Life Cycle Seeds Embryo shoot: Plumule Embryo root: radicle SEED GERMINATION (DICOTS) • Ovule-ovary attachment point Structure Cotyledons - Food storage organs that function as first seed leaves. Plumule - Embryo shoot. Epicotyl - Stem above cotyledon. Hypocotyl - Stem below attachment point. Radicle - Stem tip developing into a root. SEED GERMINATION (MONOCOTS) Germination • Germination is the beginning or resumption of seed growth. Seed must be viable. - Some require a period of dormancy.To break the dormancy artificial techniques might be used such as: - Scarification: Reduce thickness of the coat - After Ripening: Remove inhibitors - Stratification: Cold requirement mimic Favorable Environmental Factors - Imbibe water - Temperature: Above freezing but below 45 - Light: Required in some (lettuce) but unwanted in others (California poppy) Longevity • Viability of most seeds is significantly extended when the seeds are stored under conditions of low temperatures and kept dry. Aquatic lotus-1000 years - Arctic tundra lupine-10 000 years - Willow,orchids, cottonwood and tea-viable for only a few days - Cantoloupe, squash, cucumber- several years - Onion, leek- 2-3 years - Wheat with proper storage:30 years with 30 % viability 1879-William J. Beal- Experiment with 1000 seeds from 20 species of weeds - every 5 years, after 1920 every 10 years - In 1884 –most seeds of weeds germinated - In 1960-evening primrose, curly dock, moth mullein seeds germinated - In 1980 29 moth mullein, 1 mullein and 1 mallow seeds germinated - Six of original boxes are remaining and scheduled to be open in 2040 - A few species produce seeds with no period of dormancy. - Vivipary Rhizophora mangle (Vivipary) Review • • • • • Dicots versus Monocots Structure of Flowers Fruits Fleshy Dry Fruit and Seed Dispersal Seeds Germination Longevity Copyright © McGraw-Hill Companies Permission Required for Reproduction or Display Oriental Sacred Lotus/ Nelumbo nucifera