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Chapter 5: Architectural Aesthetics, Emotions And Cultural Evolution Introduction The behavioral repertoire of humans â unlike that of most nonhuman animals â relies heavily on social learning (Henrich & McElreath, 2003), which has conferred considerable adaptive advantages. Specifically, the capacity to accumulate and retain locally adaptive cultural innovations has enabled humans to spread around the globe and occupy a larger range than any other terrestrial vertebrate (Boyd, Richerson & Henrich 2011). Apart from culturally adapting to new environments, niche construction theory captures the fact that humans also actively modify their environment, which in turn feeds back on cultural and genetic evolution (Odling-Smee, 2010). In humans, as in many other animals, architecture (i.e., structures built mainly for the purposes of providing a home and for intraspecific communication) is an important part of that modified selective environment (Hansell, 2005; Odling-Smee & Turner, 2012). Given the feedback between genes, culture and constructed environments and considering the fact that social learning is adaptive in humans, in this chapter we aim to explore how human architecture might have evolved to support â or even to galvanize â social learning. More specifically we will zoom in on one general kind of architecture, i.e., religious monumental architecture, and try to shed light on how the specific awe-like emotions that can be elicited by such architecture might have impacted cultural evolution. We will distinguish between two pathways through which awe-evoking religious monumental architecture (RMA) may have enhanced cultural transmission between individuals. First, it may have stimulated cognitive performance underlying learning, leading to â for example â enhanced memory, and in so doing, it supported the reception and retention of cultural variants. Second, it may have provided a social background against which social learning could occur. Specifically, we propose that awe-evoking RMA may have caused individuals to become more prosocially oriented towards others (which increased opportunities for social learning) as well as to have contributed to establishing a learning relationship between religious leaders and commoners. 103