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The development of vegetation F.6 Geography The classification of plants 1. Life form of plants Trees (perennial 常年) Shurbs Lianas (藤本) Herbs (草本) Vegetation structure Forest Woodland Lichens Factors influencing Development 1. Plant habitats (生存環境) It is affected by landforms and soil 2. Soil Water availability Xerophytes ( 旱生植物) Hygrophytes (濕生植物) Mesophytes (中生植物) Minor classification Tropophytes ( 旱濕並生植物) Deciduous plants (落葉植物) Evergreen plants (常綠植物) Sclerophyllous plants (硬葉植物) 3. Temperature Affect rate of photosynthesis, flowering, fruiting ,seed germination Too hot will increase evaporation of plants Too cold will damage cell tissue Frontier : it means the plants can live within this boundary Imply : Every plants have their limit of distribution 4. Time - Ecological succession (生態演替) Plant and animal will succeed one another until the environment is stable for growth Climax vegetation (頂極植物) - It means the plants reached the stable community Sublimax plants (次頂極植物) It mean a stable situation by non-climatic controls Reasons for succession ~ Plants compete for suitable environment to survive ~ As environment always change , it favours different plants at different time Process of succession 1. Pioneers ( 先鋒植物) first ~ usually annual herbs and weeds 2. Grass and shurbs move in (10 – 20 yrs) 3. Pine seedlings (幼苗), form a shaded cover, changing climate 4. Pine forest mature (50yrs) ,board leave deciduous tree come (oak and hickory forest) 5. It form a mature forest finally , the climax forest is formed. (become stable and unchanging ) Common features found in climax stage Mature soil Height of plants increase and strata form Biomass and productivity increase Number of species increase New microclimate form within forest Different species replace other until a climax stage form Final community is stable than the first one In TRF, plant succession occur rapidly (50yrs) In tundra ,slow Other influence on succession Forest fires Severe storms Insects Disease Human activities If a forest is disturb by these factors , a subclimax vegetation is formed Human impacts on vegetation Clearing forest disturb climax vegetation The invasion of a plant disease by other countries cause extinction of particular plants Farming, mining, addition of fertilizer may change soil structure or affect plants growth Case study (China) Increasing population ,greater demand for cultivated land all increase pressure of cutting forest This happened seriously in North China Plain Also, during war time , many people migrate to South China ,therefore, increase pressure for farmland there Today, only 12% remained as forest in China Global and local plant environment relationship TRF ~ Amazon Basin in South America ~ Congo Basin in Africa ~ The Malay Archipelago in South East Asia ~ Central America , west coast of Africa, India, Sri Lanka, Australia Characteristics of vegetation Lack of seasonal variation ,plants lost leave in different times Vegetation layer ~ usually have five layer Leaves ~ uniform , dark green, glossy (光滑), leathery, oval and broad-leave ~ leathery due to thick cuticle (reduce transpiration) Drip-tips (滴水葉尖) ~ Allow plants to shed excess water Butress root (板根) ~ support the great weight Cauliflory (莖花) ~ the trunks or smaller trees bear flowers and fruit directly (example : cacao tree) Types of vegetation Tree species About 6000 species in Congo Mainly hardwoods, mahogany(紅木), ebony(烏木), wild rubber tree Climbing plants Plants have twining habit Example: lianas Epiphytes (附生植物) ~ ferns ,orchids, mosses and lichens ~ mainly live underground ,so called undergrowth ~ cause no harm to host Parasites (寄生植物) ~ extract water and nutrients from host Stranglers ~ originally growth like epiphytes , then put out long pendant roots to get nutrients from soil ~ when they growth up ,their root thicken and increase in number , kill the host finally Tree ferns ~ a kind of herb, growth mainly on shaded sites and often six metres high Undergrowth ~ can live in lower light environment Saprophytes (腐生植物) ~ no chlorophyll (葉綠素) ~ using symbiotic (共生根) fungi in roots ~ can live on dead organic matter Vegetation of tropical shores Mainly mangroves near tropical shores Have prop-roots(支柱根), sharply arched or growing straight up from the mud The radicle (胚根) stuck in the mud to form a firm root Plant succession and human impacts High nutrients stored in plants , but little in soil and litter Shifting cultivation increase pressure to clear land for planting Secondary forest may appear ,xerophytic plants may appear , lower dominant trees, more undergrowth and less clear strata than the primary type Need long time to restore (250 yrs) Tropical Desert vegetation Sahara Desert Arabian Desert Atacama Desert Namib Desert Kalahari Desert West-Central Australian Desert Characteristics of the vegetation Morphological characteristics ~ development of extensive root system which may be vertical or horizontal ~ deeply penetrating roots (e.g acacias, mesquite) ~ horizontal root are common in sandy or shallow soil overlying harder substrates, it may extend for 5-20 metres,just below surface ~ low shoot to root ratio 莖根比例( 1:3.5 to 1:6) Reduce the transpiration surface, plants have small leaves or even no leaves, shed foliage, roll leaves,etc. Some leaves reduced to spines Anatomical 組織characteristics ~ heavy cuticularisation (角質化過程) and cutinisation (角質化) produces a watertight, varnish-like covering to reduce water loss ~ lignification (木質化) Low, round shapes , like those of cushion plants and small cacti, can reduce structural damage from even strong wind Many hairs Stomata are arranged in recesses(凹處) or grooves (細槽) to help in retaining water Few organic matter as will eaten by termites or blown by wind Low biomass , so lack of competition Each stand is very far away Types of vegetation ~ Ephemeral annuals (短生植物) ~ 50-60% of desert plants ~ 6 – 8 weeks ~ remain as seed in the dry seasons ~ desert plantains and desert fescue ~ small size and shallow roots ~ winter annuals appear in temp 15-18C ~ summer annuals appear after the first heavy rain of summer at 26 – 33C Succulent perennials (多年生植物) Enlarge parenchyma (薄壁組織) Their stems or leaves allow the accumulation and storage of water during rainy seasons ,e.g saguaro cacti The stomata are closed at day and open at night Non-succulent perennials Plants that are adapted to arid climate Herbs, grasses, shurbs, trees The seeding produces the first leaves ,then above-ground growth stops for several months while the root system penetrates deep into the ground Can found in wadis, oases or near perennial river Examples include tamarisks, acacias grasses and palms Plants succession Cultivation cause soil erosion in desert Cutting and grazing cause dominance of plants The selection of some species for medical purposes cause extinction of certain plants The expansion of a desert that a desert ecosystem over regions that are not naturally deserts Climax vegetation in desert only happen for a short time because the soil are too mobile Tundra Vegetation 1. 2. 3. 4. 5. 6. 7. Characteristics of the vegetation Adaptation to the low temperature Adaptation to the short growing seasons Adaptation to strong wind Adaptations to permafrost Adaptation to physiological drought Decay slowly in tundra Food Storage Types of vegetation Few hundred found Small, stunted, xerophytic characteristic Trees- spruce, larch, pine Mosses and lichens Grass and sedges (most common ) - Heathers, reindeer moss, cotton grass Flowering plants – poppies Local plant-environment relationships Altitude zone vegetation characteristics ~ temperature drops ~ precipitation usually increase ~ relative humidity increase due to the cooling of air ~ light intensity and day time warming increase ~ outgoing radiation also increase , greater diurnal range in temperature Clouds form at great height because dew point is easily reached Orographic effect cause different rainfall on different sides Permanents snowcaps may exist on high mountains Winds also increase ,leading to windy exposed upper slopes South facing slopes , drier. North-facing slopes, under the shape for a long time Great changes in the environment over short distances (steep environmental gradient ) The relationship of biomes to Ecoclines Environment gradient (環境梯度)or ecocline (生態變異)is a gradient along which the communities and environment change Gradients analysis (梯度分析)looks at the changing condition across an ecosystem boundary An ecotone (群落過度帶)is a transition zone or overlap between two ecosystems The main biomes Tropical rain forest Savanna or tropical grassland Desert Temperate forest Coniferous forest Tundra