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Three-domain system From Wikipedia, the free encyclopedia A phylogenetic tree based on rRNA data, showing the separation of bacteria, archaea, and eukaryotes The three-domain system is a biological classification introduced by Carl Woese in 1977[1][2] that divides cellular life forms into archaea, bacteria, and eukaryote domains. In particular, it emphasizes the separation of prokaryotes into two groups, originally called Eubacteria (now Bacteria) and Archaebacteria (now Archaea). Woese argued that, on the basis of differences in 16S rRNA genes, these two groups and the eukaryotes each arose separately from an ancestor with poorly developed genetic machinery, often called a progenote. To reflect these primary lines of descent, he treated each as a domain, divided into several different kingdoms. Woese initially used the term "kingdom" to refer to the three primary phylogenic groupings now referred to as "domains," until the latter term was coined in 1990[2]. Classification The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the five-or-six-kingdom systems. This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as archaea appear to be more closely related to eukaryotes than they are to their fellow prokaryotic bacteria.The current system has the following kingdoms in the three domains: Archaea Domain – prokaryotic, no nuclear membrane, distinct biochemistry and RNA markers from eubacteria, possess unique ancient evolutionary history for which they are considered some of the oldest species of organisms on Earth; traditionally classified as archaebacteria; often characterized by living in extreme environments Kingdom Archaebacteria Examples: Methanogens – metabolize hydrogen and carbon dioxide into methane Halophiles – thrive in salt Thermoacidophiles – thrive in acid and high temperatures (up to 110 degrees Celsius) Bacteria Domain – prokaryotic, no nuclear membrane, traditionally classified as bacteria, contain most known pathogenic prokaryotic organisms (see [3] for exceptions), studied far more extensively than Archaea Kingdom Eubacteria Examples: Cyanobacteria – photosynthesizing bacteria Spirochaete – Gram-negative bacteria that include those causing syphilis and Lyme disease Firmicutes – Gram-positive bacteria including Bifidobacterium animalis which is present in the human large intestine Eukarya Domain – eukaryotes, nuclear membrane Kingdom Fungi or fungi Examples: Kingdom Plantae or plants Examples: Saccharomycotina – includes true yeasts Basidiomycota – includes shiitake mushrooms Bryophyta – mosses Magnoliophyta – flowering plants Kingdom Animalia or animals Examples: Arthropoda – includes insects, arachnids, and crustaceans Chordata – includes vertebrates and, as such, human beings Kingdom Protista or protists (recognized to be paraphyletic, and thus subject to dissolution and/or redefinition) Examples: Rhodophyta – red algae Chromalveolata – includes dinoflagellates