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Plants Angiosperms Tracheophytes • Reproductive structures of the tracheophytes: • Ferns create sporangia. • These hold and eventually release spores. • Spores grow to create a new generation of fern. • Gymnosperms create strobili (or cones). • These create pollen (male strobili) or eggs (female strobili). • The pollen carries half of the genetic information necessary for the creation of a new plant to the egg, which holds the other half, in a process called pollination. • Fertilization occurs when the genetic information in the pollen is merged with the other half of genetic info. in the egg. • The fertilized egg is the seed, the embryonic plant. • In gymnosperms, the seeds are eventually released from the female cones into the wild to create the next generation of that plant. • Angiosperms create flowers. Angiosperms • Flowers have certain parts that are considered “male” or “female:” • Anthers – the male parts of the flower that generate pollen • Ovaries – the female parts of the flower that hold the egg • The male and female parts of the flower can be: • On the same flower (a perfect flower), or • On separate (male or female) flowers. • Sometimes, these separate male and female flowers will be on the same plant (called being monoecious) ; • Sometimes these separate flowers will be on separate (male or female) plants (called being dioecious). Angiosperms • Flowers depend on other creatures, usually animals (often bees) or on the wind for pollination to occur. • Bees and flowers have a complex, symbiotic relationship. • The bees get food (nectar and/or pollen) from the flowers. • The flowers become pollinated by the bees while they are gathering nectar and/or pollen from the flowers. • This type of symbiosis is called mutualism (because both creatures benefit from the relationship). Parts of Flowers Mature Perfect Flower • Flowers produce pollen and nectar for the purpose of eventually creating a seed. Nectaries • Pollen is produced by the anthers. Nectar Producing • Nectar is produced in the nectaries, glands that Glands in Flower can be found in various locations depending on the flower type. • Ovules (eggs) are produced in the ovaries. • • • • There may be a single ovary in a flower, or There may be multiple ovaries in a single flower. A single ovary may produce a single egg within, or A single ovary may produce multiple viable eggs within itself. Parts of Flowers MicrosporangiumMature Perfect Flower produces microspores – Microspores become Pollen Grains Anthers Male Parts of Flower produce pollen Pollen Grains will will burst burst out out of of microsporangium microsporangium Pollen grains carry plant gametes (sperm cells). Parts of Flowers Pistil of a Flower • Pollen grains carry gametes, or sex cells, which are the “male” sex cells, called sperm cells. • The “female” sex cells are the ovules, or eggs. • Each of the gametes carry half of the genetic information needed to create a new plant. • Pollen grains from different plants can look very different . . . • When the pollen grain gets to the female flower’s pistil, it germanates, creating a pollen tube through which the sperms cells travel to reach the female flower’s ovules. Pollen Grains will will burst burst out out of of microsporangium microsporangium Parts of Flowers Ovaries Female Parts of Flower Produce Ovules (Eggs) Mature Perfect Flower Ovary Produces Produces Ovule(s) Ovule(s) Ovule (egg) Once eggs are fertilized by a sperm cell from a pollen grain, you have a seed. A seed is an embryonic plant (beginning of a new plant generation). Bees • Bees are amazing creatures with a complex societal structure & an extreme interdependence on flowers. • Bees may collect several things from the flowers in their environment: • Nectar • Nectar is turned into honey by bees for use as food. • A bee honey stomach and its saliva contains enzymes that alter the chemical structure of the nectar. • Bees also encourage evaporation of the water in nectar, so that the solution goes from about 80% water to about 16% as honey. • People also eat honey that has been created by bees. • Pollen • Pollen is collected and eaten and also stored for later use as food during the months of the year when flowers are not available. Bees • Bees are designed with organs and physical structures specifically for their interactions with flowers. • Bees have a proboscis (a hollow tongue) which they extend from their mouths into a flower to get the nectar. • Bees have a honey stomach (also called a crop) into which they funnel nectar for storage until they return to the hive. • Bees have special structures on one section of their legs called pollen baskets (or corbicula—Latin for “basket”) which help them collect pollen. • Bees have structures on other parts of their legs (mostly comb-like hairs) to help them pull pollen off of other parts of their bodies and move it into their pollen baskets. • Bees can digest pollen for protein and honey for energy. • Bees know how to turn nectar into honey . . . Bees – Covered in Pollen Bees – Proboscis Proboscis retracted Proboscis extended Bees – Honey Stomach Bees – Pollen Basket • Honey bees have pollen baskets, or corbicula, on their hind legs. • They will use their tongue to wet the hairs on their forelegs and any pollen there with nectar and then use them to wipe the pollen that they got all over them back toward their hind legs and into the corbicula. • The pollen becomes hard-packed, smooth, and shiny as it is packed into the corbicula of a honey bee. Bees – Pollen Basket Full Pollen Baskets Rear Leg – Pollen Basket (Corbicula) Bees – Scopa Bumblebee carried in a scopa • Note that the with onlypollen species of honey bee in North America has pollen baskets (corbicula), however . . . • Bumblebees and other bee species may instead have a scopa, which is a collection of hairs in a similar location but which holds pollen differently. • Pollen carried in a scopa is not wet or compressed. • It is left fluffy and dry, jammed in between the hairs on the leg or pinned through by them, rather than packed into a “basket.” Bees – Nectar & Pollen Storage • Bees store nectar—transformed into honey—and Beehive in tree trunk pollen back in their hives. • Hives are usually made in cavities in trees in nature. • However, they will also build between walls, eaves, or rooves of people’s homes. • They create hexagonal cells from wax and put the honey and pollen—and even baby bee eggs—in these wax cells. • A honeycomb is a structure created from many of these cells. • The bees themselves create the wax. • It is generated under their abdomens when they are a certain age. • Bees will collect this wax from their fellow bees, build the honeycomb, and then fill the cells with what they need. Bees – ofNectar Pollen Different & Pollen Storage Kinds Stored in Cells Bee Larvae Honey Cells Sealed with Wax Bees • Bees also may collect propolis (prop-a-liss) from plants. Propolis sealing edges of • Propolis is collected from either: a bee hive. • Flower buds • Sap flows from coniferous gymnosperms (!) • Other botanical sources • Uses bees have for propolis: • Reinforce the structural stability of the hive • Reduce vibration • Make the hive more defensible by sealing alternative entrances • Prevent diseases and parasites from entering the hive • To inhibit fungal and bacterial growth • Preventhoney putrefaction hive. Africanized bee within theBrazilian bees sealing • Should a small lizard or mouse finds its way into the hive and carrying green large opening in hive die, beespropolis may be unable to carry it out. native with greeninstead, propolis. • If to so, Brazil. they would seal the carcass in propolis mummifying it and making it odorless and harmless. Bees • Propolis is actually a mixture bees create when they need it. • They mix tree sap with pollen and wax to create propolis. • It is a thick, resin-like substance. • Many people have counted on propolis for medicinal purposes for millennia, including: Oral health Preventing or fighting off various types of infections Healing wounds and burns As an anti-fungal agent As an anti-parasitic To fight cancer and associated illness As an anti-inflammatory agent For wart removal To reduce cold sores & other similar, related and unrelated viral infection symptoms • As a preservative (was used in mummification by the Egyptians) • • • • • • • • • Bees – Random Facts • Honey bees pollinate 30% of all food that Americans consume. • They perform 90% of all pollen transfers on our orchard crops • They pollinate 85% of all flowering plants • One honey bee visits 50-100 flowers during each collection trip and can harvest several thousand flowers in a day, making 12 or more trips, gathering pollen or nectar from a single floral species each. • It takes about 556 worker bees to gather 1 pound of honey from about 2 million flowers. • It takes about 55,000 flight miles per gallon of honey made. 1 • The average honey bee will make only of a teaspoon of 12 honey in its lifetime (6 weeks). • These foragers are the oldest bees in the hive. • It is during the last two weeks of their lives that they gather nectar, pollen, water, and propolis. Bees – Random Facts • A hive can gather pollen and nectar from up to 500 million flowers in a year. • During peak nectar flows, a healthy hive can produce 2 to 5 pounds of honey per day. • The bees use about 8 gallons of water to make 1 gallon of honey. • 9 pounds of honey is synthesized to make 1 pound of beeswax. • Honey bees can fly up to 6 miles from the hive at 15 mph with their wings beating 11,400 times per minute. • Honey bees use the sun as a directional marker when leaving and returning to the hive. • The returning bees do a waggle dance on the vertical comb surfaces in a circle or figure eight pattern which shows the other bees: • Which direction to fly, and • How far to fly. Bees – Random Facts • Roughly fifty thousand worker bees (females) live in a colony along with one queen and several hundred drones (males). • During the warmer months the worker bees live about six weeks, the queen can live up to three years. • Worker bees born in the fall will live throughout the winter with the hive population being about half of what it is in the summer. • Average interior temperature of the hive's brood area is 93-95 degrees (F) regardless of the outside temperature. • In colder weather they do not hibernate, but cluster generating heat much like musk-ox and penguins. • There are approximately 2.7 million managed Honey Bee Hives in the USA. • There are actually no honey bees native to the United States. • European honey bees (German Black Bees) were first introduced into this country (in Virginia) about 1621-22. Modified from http://www.apexbeecompany.com/honey-bee-facts/ Seeds • Recall that the pollen grains: • Move from the anthers of one flower to the pistol of another flower (of the same species). • They then create a pollen tube and send the gamete cells down the pistol in search of the egg(s). • Once they reach the egg, the egg is said to be fertilized. • At this point, you will finally get a seed. • It is the male and female gametes fused into one new embryo. Seeds - Cotyledons • A cotyledon is part of the embryo within the seed of a plant. • The cotyledons are formed during the process of embryogenesis along with the roots and shoots of the plant prior to germination. • Often when the seed germinates (begins to grow), the cotyledon may become the first leaves of the seedling. • Botanists use the number of cotyledons present in the seed of a plant as a means of classification. • Monocots are seeds that have only one cotyledon. • Dicots are plants with two cotyledons. Seeds – Monocot vs Dicot • Flowers • Monocots tend to have flower parts in multiples of 3. • Dicots tend to have flower parts in multiples of 4 or 5. • Leaves • Monocots tend to have parallel veination. • Dicots tend to have net veination. • Roots • Monocots usually have adventitious roots. • Adventitious roots arise from an organ other than the actual root. • Usually, from a stem, but sometimes a leaf. • They are especially numerous on underground stems. • The formation of adventitious roots makes it possible to vegetatively propagate many plants from stem or leaf cuttings. • Dicots usually have tap roots. • Stems • Monocots’ vascular bundles are usually spread throughout the cross-section of the stem. • Dicots usually have vascular bundles spread to the outside. Apple Trees Malus domestica Orchard Apple or Table Apple Integrated Taxonomic Information System (ITIS) Classification: Domain: Kingdom: Division: Class: Order: Family: Genus: Species: Eukaryota Plantae Tracheophyta Magnoliopsida Rosales Rosaceae Malus domestica Apple Tree in Bloom ! Apple Trees Malus domestica Orchard Apple or Table Apple National Center for Biotechnology Information (NCBI) Classification: Domain: Kingdom: Phylum: Class: Order: Family: Genus: Species: Eukaryota Viridiplantae Streptophyta Rosales Rosaceae Malus domestica ! Apple Trees • The various types of modern apples are descended most likely from the Malus sieversii, a wild apple found in various regions of Central Asia. • The species name domestica denotes that the apple in question is a modern apple with only a likely descendence from the Malus sieversii. • Actual varieties of modern apples are called cultivars, and are hybridized plants that humans have created by breeding different types of apple-bearing plants. • Just the American cultivars (or those popular in the U.S.) include: Bailey Baldwin Beacon Ben Davis Beverly Hills Bottle Greening Cameo Carolina Red Juice Carter’s Blue Courtland Crimson Delight Crimson Gold Criterion Dudley Winter Empire Enterprise Esopus Spitzenburg Fireside Ginger Gold Golden Delicious Golden Russet Golden Supreme Granny Smith (AU) Green Cheese Grimes Golden Harrison Harrison Cider Honeycrisp Honeygold Idared Jonagold Jonathon Junaluska Jupiter King Lady Alice Liberty Lodi Macoun Maiden’s Blush Melinda Golden Delicious McIntosh (CA) Melon Melrose Mollie’s Delicious Mother Newell-Kimzey Newtown Pippin Nickajack Northern Spy Ozark Gold Paula Red Pink Pearl Granny Smith (AU) Pristine Prima Porter’s Pound Sweet Red Delicious Rhode Island Greening Rome Beauty Roxbury Russet Smokehouse (from PA!) Stark Earliest Stayman Sweet Sixteen Swee Tango Teser Tolman Sweet Twenty Ounce Westfield SeekNo-Further Wealthy Winesap Wolf River York Imperial McIntosh (CA) Red Delicious Apples • As the apple matures, the fertilized seeds in the ovary are enclosed in: • A fleshy fruit which the ovary grows into, and • A further layer of fleshy material which grows from other parts of the flower, not the ovary. • People generally eat the outer part of apples, not the part that grew from the ovary. • The apple core is what the ovary eventually becomes, but we generally throw this part away. • It is the sepals, petals, and stamens that were the hypanthium, or flower cup, around the base of the flower, become the part people actually like to eat. Daisies Bellis perennis Lawn Daisy or European Daisy Integrated Taxonomic Information System (ITIS) Classification: Domain: Kingdom: Division: Class: Order: Family: Genus: Species: Eukaryota Plantae Tracheophyta Magnoliopsida Asterales Asteraceae Bellis perennis ! Daisies Bellis perennis Lawn Daisy or European Daisy National Center for Biotechnology Information (NCBI) Classification: Domain: Kingdom: Phylum: Class: Order: Family: Genus: Species: Eukaryota Viridiplantae Streptophyta Asterales Asteraceae Bellis perennis ! Daisies • Daisies are another example of an angiosperm plant. • They are called Lawn Daisies, European Daisies, or even Bruisewort at one time. • Many people consider daisies to be a lawn weed. • The name is understood to be a mangling of the actual given name from long ago, Day’s Eye. • The flower appears like a little yellow sun with rays spraying out all around it. • The binomial designation Bellis perennis comes from Latin terms. • Perennis means everlasting. • Bellis either comes from: • The Latin word bella, which means beautiful—meaning the plant is beautiful all year long or every year . . . or, • The Latin word bellum, which means war—meaning there is an association with war and this plant, and it was used as healing agent for wounds after battles in Roman times and thereafter. Daisies • Daisy “flowers” are actually an inflorescence of many small flowers. • There are really 2 types of flowers involved in a daisy flowerhead, which is a capitulum type of inflorescence: • Disc florets • Each of these disc florets have both male (stamens with anthers) and female (stigmas on a style) parts (perfect flowers) and are all situated above individual ovaries. • Ray florets • Each of these ray florets have both male (stamens with anthers) and female (stigmas on a style) parts, however the male parts are often sterile, making the rays female. Each is situated over an ovary. • These florets all sprout from a receptacle at the top of a leafless stalk and are surrounded by involucre (inve-loo-ker) bracts (leaf-like structures that surround the flower from underneath). • The fruit produced is of the achene type. • These fruits are one-seeded, lance shaped, flat, & yellowish brown. • Daisies can also reproduce from creeping, underground horizontal stems (rhizomes). Daisies Disc Floret Make up Interior Part of Inflorescence Daisies Ray Floret Make up “Petals” Surrounding Interior Flowers Daisies Capitulum Type Inflorescence — Made Up of Many, Many Disc Florets in the Middle and Multiple Ray Florets Around Outer Edge Daisies Wheat Wheat Triticum aestivum Common Wheat Integrated Taxonomic Information System (ITIS) Classification: Domain: Kingdom: Division: Class: Order: Family: Genus: Species: Eukaryota Plantae Tracheophyta Magnoliopsida Lilianae Poaceae Triticum aestivum ! Wheat Triticum aestivum Common Wheat National Center for Biotechnology Information (NCBI) Classification: Domain: Kingdom: Phylum: Class: Order: Family: Genus: Species: Eukaryota Viridiplantae Streptophyta Liliopsida Poales Poaceae Triticum aestivum ! Wheat • Wheat is another example of an angiosperm plant. • The fruit of a wheat plant is technically called a caryopsis (plural caryopses). • It is a type of simple, dry fruit that is: • • • • Monocarpellate (formed from a single carpel) Indehiscent (not opening at maturity) Like an achene in appearance Different from an actual achene because the pericarp is fused with the thin seed coat • The caryopsis is popularly called a grain. • Because the fruit and the seed are essentially one unit, no one really speaks of the fruit and seed as anything other than justCaryopsis one thing: a grain. Wheat • An “ear” of wheat consists of: • A number of individual spikelets (flower heads) set one above another • These are all along a length of stem known as the rachis. • Each spikelet has a single set of glumes (sheaths) enclosing up to 8 grains (each developing from a flower); • Each grain is enclosed in its own glumes (the lemma and palea). • The glumes may have long dorsal to terminal bristles known as awns but this is now rare in modern wheat. • There are certain characteristics that aid in identifying the specific type (species) of wheat you are dealing with: • The number of flowers/spikelets • Whether or not the grains are easily released from the glumes is another: • ‘Hulled' wheats have strong glumes which hold the grains in the spikelets • ‘Naked' wheats are easily released even from the lemma and palea • The ease of release has implications for timing of harvest, ease of threshing, and resistance to disease and/or predation. http://www.farm-direct.co.uk/farming/stockcrop/wheat/wheatanat.html Wheat • Wheat is grown on more than 540,000,000 acres of farmland around the world—larger than for any other crop. • World trade in wheat is greater than for all other crops combined. • With rice, wheat is the world's most favored staple food. • It is a major diet component because of the wheat plant's: • • • • Ability to grow from near arctic regions or at the equator Ability to grow at sea level to high plains well above sea level Ease of grain storage Ease of converting grain into flour for making edible, palatable, interesting and satisfying foods • Wheat is the most important source of carbohydrate in a majority of countries. • Wheat protein is easily digested by nearly 99% of the human population. • With a small amount of animal or legume protein added, a wheat-based meal is highly nutritious. https://en.wikipedia.org/wiki/Wheat