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Water and Plant Cells • Properties of Water • Water Potential • Components of Water Potential • Water relations Water in plant life • Corn yield as a function of water availability. • Productivity of ecosystems as a function of annual precipitation. • Most water lost through transpiration. Properties of Water • • • • • • • Polar molecule Thermal Properties Specific Heat Melting Point Heat of fusion Boiling point Heat of vaporization Solvent properties • Dielectric constant • Cohesion and Adhesion Water Transport Processes • Diffusion • Pressure-driven bulk flown (Aquaporins) • Osmosis Water movement • Fick’s law of diffusion • Js = - Ds (∆Cs / ∆x) • Js is rate of transport (mol/m2s1) • Ds = diffusion coefficient through a medium • (∆Cs / ∆x) = concentration gradient of a substance separated by distance x. Water Potential • Chemical activity of water • Components of water potential • Ψw = Ψs + Ψp + Ψg Components of water potential: Solutes (osmotic potential) • Ψs = 0 for pure water Or always negative • Ψs = - R T Cs • R = 8.32 J/mol K • T = absolute temperature • Cs = solute concentration (mol/L) Pressure potential • • • • Hydrostatic pressure (Ψp ) Positive (turgor pressure) Negative (tension) Ψp = 0 for water at ambient pressure (0 MPa) Gravity • Ψg = pwgh • Pw = density of water • g = acceleration due to gravity • pwg = 0.01 MPa m-1 • At cell level we can ignore gravity or for small plants. Concepts of water potential Plant cell volume • Small changes in volume cause large changes in turgor • 10-15% change in volume, turgor goes to zero • Modulus of elasticity (ε) or rigidity of cell wall. • High ε wall is stiff, low ε wall is soft Physiological changes during drought