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Phylum Cnidaria • “From a distance I was never threatened, in fact I was infatuated with its beauty. A large, inviting, bright blue float lured me closer. As I swam nearer I could see that hidden from my previous view was an infrastructure of tentacles, some of which dangled nearly nine meters below the water’s surface! The creature seemed to consist of many individuals and I wondered whether or not each individual was the same kind of being because, when I looked closely, I counted eight different body forms! • I was drawn closer and the true nature of this creature was painfully revealed. The beauty of the gasfilled float hid some of the most hideous weaponry imaginable. When I brushed against those silky tentacles I experienced the most excruciating pain. Had it not been for my life vest, I would have drowned. Indeed, for some time, I wished that had been my fate.” • Radial or Biradial – Advantageous because it allows for sensory structures are evenly distributed around the body. • Tissue level of organization • Ectoderm forms epidermis (outer layer) • Endoderm forms gastrodermis (inner layer) • Mesoglea contains some cells but does not have true tissues. • Cnida: fluid-filled, intracellular capsule enclosing a coiled hollow tube • Operculum: lid • Cnidocil : stimulation of the cnidocil forces open the operculum and discharges the coiled tube. • Specialized cnidocyte • Used in food gathering and defense • Armed with spines that penetrate the prey and release paralyzing toxins • Other cnidocytes – Wrap around prey – Sticky secretions that help in attachment • Most Cnidarians possess two body forms in their life histories. • Polyp – Asexual and Sessile – Attaches to a substrate at aboral end – Mouth is surrounded by food-gathering tentacles • Medusa – Dioecious and free-swimming – Shaped like an inverted bowl with tentacles hanging – Mouth faces downward – Swims by pulsations of the body wall • Gastrovascular Cavity – Functions in digestion, respiration, excretion, and discharge of gametes – Feed on small crustaceans and fish – Nematocysts entangle and paralyze prey – Tentacles shorten which draws food to the mouth – Mucus and enzymes dissolve food in the gastrovascular cavity which are phagocytized and further digested. • Hydrostatic skeleton – Water or fluids in body cavity and against which contractile elements of the body wall act. – Think of a balloon – Polyps: somersault, glide or inch across the substrate – Medusa: swim or float • Nerve Net – Simple nervous system – Nerve cells located below epidermis and interconnect…this creates a nerve net. – The net conducts impulses around the body in response to a localized stimulus. – Response changes due to stimulus strength. • Most are separate sexes • Eggs and sperm are released into gastrovascular cavity • Free swimming larva called planula • Can also bud (become medusa) Animalia Cnidaria • • • • • Hydrozoa Scyphozoa Staurozoa Cubozoa Anthozoa • • • • Small, relatively common Mostly marine, some freshwater Alternation of generations Nematocysts are only in the epidermis • Gametes are released to the outside • Mesoglea does not contain cells • Colonial Polyps – Individuals are specialized for feeding, reproduction, or defending the colony • Ex: Obelia, Gonionemus, Hydra (freshwater), Portuguese Man of War • All Marine • Sessile, oral end looks like medusa • Crawling larva…very little dispersal abilities • ~100 species • All Marine • “True Jellyfish” – Dominant stage in life cycle is the medusa stage Most are harmless but some can be very dangerous to humans Page Title • Note 1 • Note 2 • Note 3 • Medusa is cuboidal • Tentacles hang from each corner • Active feeders and swimmers in tropical waters • Some possess dangerous nematocysts • • • • Colonial or Solitary Lack Medusa All Marine External – radial symmetry • Internal – biradial symmetry • Solitary, large, and colorful • Some attach to solid substrates, burrow in soft substrates, or live in symbiotic relationships • Some move by thrashing or crawling along a surface • Feed on invertebrates and small fish • Asexual Reproduction: Fragmentation • Sexual Reproduction – Monoecious (protandry) – Dioecious • Stony Coral – Look like anemones – Have a calcium carbonate exoskeleton that they can retract into when threatened • Soft Coral – Do not have the CaCO3 exoskeleton • Sea anemone/Clownfish • Coral/Zooxanthellae – Protists that undergo photosynthesis and give the coral their color – Also release carbon to build exoskeletons