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2.3.1 Flows of Energy and Matter Transfer and transformation of Energy Not all solar radiation ends up as biomass. Losses include: ◦ Reflection from leaves ◦ Not hitting chloroplasts ◦ Wrong wavelength ◦ Transmission of light through the leaf ◦ Inefficiency of photosynthesis ENERGY ENTERS THE ECOSYSTEM AS SUNLIGHT Only 2% of the light energy falling on plant is used to create energy The rest is reflected, or just warms up the plant as it is absorbed Transfer and transformation of energy. Energy comes into the ecosystem as light energy, Transfer and transformation of energy. Energy converted into chemical energy by producers. Transfer and transformation of energy. That chemical energy is transferred as organisms are eaten, with energy being lost as heat and respiration. Photosynthesis Process where plants use sun light energy to create chemical energy Photosynthesis: equation ◦ 6CO2 + 6H2O --> C6H12O6 + 6O2 Inputs: light energy, water, carbon dioxide Outputs: oxygen gas, sugar Energy transformations: Light to Chemical respiration backwards! Respiration Process by which animals create energy through consumption of organic molecules (sugars) Respiration: ◦ C6H12O6 + 6O2 --> 6CO2 + 6H2O Inputs: oxygen gas, organic molecules (sugars) Outputs: carbon dioxide, energy in ATP, waste heat Energy transformations: chemical to heat Photosynthesis backwards! Energy Transfers in Ecosystem Energy Flow Diagram Conversion of Energy Conversation of energy into biomass for a given period of time is measured as productivity Gross productivity Total energy captured or “assimilated” by an organism. Measured in joules (J) Plant (Gross Primary Productivity) ◦ GPP = sunlight energy used during photosynthesis Animals (Gross Secondary Productivity) ◦ GSP = food eaten - energy in faeces Energy is stored in leaf as sugars and starches, which later are used to form flowers, fruits, seeds, Net productivity The energy left over after organisms have used what they need to survive. All organisms have waste energy and respiratory loss given off as heat, metabolism (R) Net productivity Plants and animals have to use some of the energy they capture to keep themselves growing: ◦ They both move water and stored chemicals around ◦ Plants make flowers, fruits, new leaves, cells and stems ◦ Animals create cells and need to move muscles. Net productivity = Gross productivity - Respiration Energy or using symbols: NP = GP - R Net Primary vs. Net Secondary Productivity (NPP) vs. (NSP) Calculate Net productivity for plants and animals NPP = GPP – R PLANTS NSP = GSP – R ANIMALS NSP GSP Productivity Primary Productivity – gain by autotrophs in energy or biomass per unit area per unit time Productivity Secondary Productivity – biomass gained by heterotrophs thru feeding and absorption, measured in units of mass or energy per unit area per unit time Productivity Gross Primary Productivity – Gain in energy or biomass thru photosynthesis per unit area per unit time. Productivity Net Primary Productivity – The gain by producers in energy or biomass per unit area per unit time remaining after respiration losses. Productivity Gross Secondary Productivity (Gross Assimilation) – Gain in energy or biomass thru absorption per unit area per unit time. Productivity Net Secondary Productivity (Net Assimilation) – The gain by consumers in energy or biomass per unit area per unit time remaining after respiration losses. Productivity in Food Web In a food web diagram, you can assume that: ◦ Energy input into an organism represents the GP ◦ Energy output from that organism to the next trophic level represents the NP ◦ GP-NP = R (respiration energy ) and/or loss to decomposers ? Therefore… The least productive ecosystems are those with limited heat and light energy, limited water and limited nutrients. ◦ Example biome:_______________ The most productive ecosystems are those with high temperature, lots of water light and nutrients. ◦ Example biome:__________________ How to measure primary productivity 1. 2. 3. Harvest method – measure biomass and express as biomass per unit area per unit time. CO2 assimilation- measure CO2 uptake in photosynthesis and releases by respiration O2 production-Measure O2 production and consumption Measuring productivity continued 4.Radiosotope method-use c14 tracer in photosynthesis. 5.Chlorophyl measurement- assumes a correlation between the amount of chlorophyll and rate of photosynthesis. ◦ Complete this energy flow diagram: Label GPP, NPP and R for the primary producer Add arrows to show missing energy pathways (5 in total) Fill in the blank box to explain why some sunlight is not fixed by plant HERBIVORES PLANT SUN ……………… ……………. (~98% of energy is here) RESPIRATION DECOMPOSERS The data in the table below relate to the transfer of energy in a small clearly defined habitat. The units in each case are in kJ m-2 yr-1 Trophic Level Gross Production Respiratory Loss Loss to decomposers Producers 60724 36120 477 1° Consumer 21762 14700 3072 2° Consumer 714 576 42 3° Consumer 7 4 1 Respiratory loss by decomposers --- 3120 --- Construct an energy flow model to represent all these data – Label each arrow with the appropriate amount from the data table above. Use boxes to represent each trophic level and arrows to show the flow of energy Calculate the Net Productivity for ◦ NPP for Producers ◦ NSP for 1°Consumers, 2°Consumers, 3°Consumers ◦ NSP for Decomposers ENERGY FLOW MODEL R=36120 60724 Producers R=576 R=14700 21762 1 Consumer 714 2 Consumer 3072 R=4 7 3 Consumer 42 477 1 Decomposers R=3120 Productivity Calculations NPP of Producers:60724 - (36120+477) NSP of 1 Consumer21762 -(14700+3072) NSP of 2 Consumer714 -(576+42) NSP of 3 Consumer7 -(4+1) NSP of Consumers: 22483 -(15280+3115) NSP of Decomposers: (477+3072+42+1) =24127 kJ.m-2.yr-1 =3990 kJ.m-2.yr-1 =96 kJ.m-2.yr-1 =2 kJ.m-2.yr-1 =4088 kJ.m-2.yr-1 -3120 =472 kJ.m-2.yr-1