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9.6 Hearing and Equilibrium The Ear: • Used for hearing and equilibrium • The inner ear contains sensory cells called hair cells for both functions • Each hair cells has between 30 and 150 cilia… responding to stimuli • The ear can be divided into 3 main sections: • 1) outer ear • →The pinna • →The auditory canal • 2) middle ear • →Eardrum (tympanic membrane) • →Ossicles (malleus, incus, stapes) • →Oval window • →Eustachian tube • 3) inner ear • →Vestibule • →Semicircular canals →Cochlea The Outer Ear – The pinna: outer ear flap that collects and directs the sound – The auditory canal (ear canal): carries sound to the eardrum and is lined with specialized sweat glands that produce earwax (used to trap foreign particles, preventing them from entering the ear.) The Middle Ear • Sound is a form of energy that converts into electrical impulses for interpretation. • Sound must travel through a medium such as air (but also water and solids) to be heard. The Middle Ear • Air filled chamber that holds 3 small bones called ossicles. – malleus (hammer), incus (anvil) and stapes (stirrup) • Sound vibrations strike the eardrum and are transmitted to the malleus, then the incus and lastly the stapes. • The stapes strikes the membrane covering the oval window in the inner wall of the middle ear and amplifies the sound. • Muscles joining the bones of the middle ear protect the inner ear from excessive noise. After receiving vibrations from the ossicles, the oval window pushes inwards towards the fluidfilled cochlea. Fluid waves within the cochlea convert them into electrical impulses (sound) using the Organ of Corti in the inner ear. Equalizing pressure in the middle ear • The eustachian tube goes from the middle ear to the mouth and the chambers of the nose and helps to equalize pressure. This is the site that builds up with fluids in an ear infection. The Inner Ear: Organ of Corti • The Organ of Corti is the actual hearing apparatus in the cochlea. The organ of Corti is located in the cochlea. The cochlea is shaped like a snail’s shell and contains two rows of specialized hair cells that respond to sound waves of different frequencies and intensities and changes them into nerve impulses. • The hair cells are covered in a gelatinous coating and sit on the basilar membrane. The Inner Ear: Organ of Corti • As fluid waves pass through the cochlear canals, the basilar membrane moves up and down causing the hair cells to hit the membrane above called the tectorial membrane. • As hairs bend, messages are sent through sensory nerves at their base to the cochlear nerve and eventually the auditory nerve that leads to the brain. Organ of Corti • BBC Science & Nature Human Body and Mind Nervous System Layer Effect of Sound Waves on Cochlear Structures Loudness • Receptor hair cells on the basilar membrane trigger an action potential that is carried to the brain • The louder the sound, the more hair cells are stimulated. • Hair cells can be damaged if the sound is too loud. Pitch • Low frequency waves (bass) move to the wider more elastic area of the cochlea to vibrate. Animation: Effect of Sound Waves on Cochlear Structures (Quiz 1) Mechanical Stimulation • The basilar membrane will respond to jarring blows … particularly to the head • The vibrations resonate through the skull, and pass onto the cochlea. This can cause ‘ringing’ in your ears. Equilibrium and the Inner Ear • An area called the vestibule is connected to the middle ear by the oval window and has two small sacs: the utricle and the saccule that help establish head position. • There are three semicircular canals that are arranged at different angles that helps identify body movement with the fluid inside them. Anatomy of the Ear Learning Activity Static Equilibrium • Movement along one plane (head position) • Controlled by the saccule and utricle. Cilia from hair cells are suspended in a gelatinous material containing CaCO3 granules called otoliths. • When the head is in a normal position, otoliths do not move but when the head moves, gravity acts on otoliths, causing gelatinous material to shift and hair receptors to bend. Movement of hair receptors stimulates nerves that relay info about head position to brain. Dynamic Equilibrium: • Provides information during movement • While moving, balance is maintained by 3 semi-circular canals, each of which has a small pocket called an ampulla • Rotational stimuli cause the fluid and hence the otoliths in these canals to move.