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Plants and Our Environment Classification Introduction We live on a planet called Earth. Biologists call the Earth and its atmosphere -- the Biosphere. The Biosphere is composed of organic and inorganic matter Organic matter (stuff containing carbon) is again divided into living and non-living objects All living beings are currently divided into five Kingdoms Kingdoms of All Living Beings: Plantae - the plant kingdom, studied in the field of Botany Fungi - the kingdom of fungus and molds, studied as Mycology Animalia - the animal kingdom, the domain of Zoology Protoctista - a catch-all for all other "higher-order" organisms from singlecelled microbes to large seaweeds (algae) Monera - consists of bacteria -- smallcelled microorganisms without true cell nuclei Protoctista Green Algae (7000 species) Red Algae (4000 species) Brown Algae (1500 species) Diatoms (10,000 species) and various Plankton Animalia Spinal Cords (Chordata) Joint-Legs (Arthropoda) Soft Bodies = Mollusks (Mollusca) the Kingdom most and least familiar to us mammals, fish, and birds -- belong to a single subgroup within one of the 33 Phyla comprising the Animal Kingdom 800,000 + species have been identified in the Animal Kingdom -most of them in the Arthropod phylum Other Animalia Spiny-skinned (Echinodermata) [Marine] Stingers (Cnidaria) Fungus provide a critical part of nature's continuous rebirth recycle dead organic matter into useful nutrients Many plants depend on the help of a fungus to get their own nutrients “symbiotic relationship” Plantae: The Plant Kingdom Plants provide nourishment for our bodies and souls Plants provide the oxygen we breathe and the food that sustains us Plants provide shade over our heads and cool carpets under our feet Surround us with beautiful colors and marking the change of seasons Prominent plants determine ecological communities such as "Redwood-Tanoak Forest" or "Oak Grassland" and dictate the animals, fungi found there, and climate as well. Plant Classification • Plants are classified into 12 phyla or divisions based largely on reproductive characteristics; they are classified by tissue structure into nonvascular (mosses), and vascular plants (all others) q by "seed" structure into those that reproduce through naked seeds, covered seeds, or spores; q by stature divided into mosses, ferns, shrubs and vines, trees, and herbs. All of these higher-level groupings are decidedly lopsided: the vast majority of the 270,000 plant species are flowering herbs. Division or Phylum? Although the International Code of Botanical Nomenclature allows the use of the term "phylum" in reference to plants, The term "Division" is almost always used by botanists. Plant Divisions Green Algae (Division Chlorophyta) - most primitive members Liverworts and Mosses (have neither xylem and phloem) nor woody tissue Lycopsids (Division Lycopsida) - often called club mosses. They are not mosses at all, but vascular plants with xylem and phloem running through their roots, stems, and leaves Horsetails (Division Equisetopsida) Ferns (Division Filicopsida) Seed Plants (Division Spermatopsida) – Gymnosperm and Angiosperm CLASS Angiospermae (Angiosperms) Plants which produce flowers Gymnospermae (Gymnosperms) Plants which don't produce flowers SUBCLASS Dicotyledonae (Dicotyledons, Dicots) Plants with two seed leaves Monocotyledonae (Monocotyledons, Monocots) Plants with one seed leaf SUPERORDER Group of related Plant Families classified in the order in which they are thought to have developed their differences from a common ancestor Six Superorders in the Dicotyledonae (Magnoliidae, Hamamelidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae), Four Superorders in the Monocotyledonae (Alismatidae, Commelinidae, Arecidae, Liliidae) The names of the Superorders end in -idae ORDER Each Super order is further divided into several Orders. The names of the Orders end in ales FAMILY Each Order is divided into Families. Plant with many botanical features in common This is the highest classification normally used. Similarity between plants is often easily recognizable The names of the Families end in -aceae SUBFAMILY Family may be further divided into a number of sub-families Group together plants within the Family that have some significant botanical differences. The names of the Subfamilies end in oideae TRIBE A further division of plants within a Family, based on smaller botanical differences The names of the Tribes end in -eae SUBTRIBE A further division, based on even smaller botanical differences, often only recognisable to botanists. The names of the Subtribes end in -inae GENUS This part of the plant name that is most familiar, Normal name that you give a plant – Acer (Maple), Quercus (Oak), and so on. The plants in a Genus are often easily recognizable as belonging to the same group. The name of the Genus should be written with a capital letter. SPECIES This level defines an individual plant. Often, the name will describe some aspect of the plant – Colour of the flowers, size or shape of the leaves, or it may be named after the place where it was found. Together, the Genus and species name refer to only one plant Used to identify that particular plant Sometimes, the species is further divided into sub-species that contain plants not quite so distinct that they are classified as Varieties. The name of the species should be written after the Genus name, in small letters, with no capital letter. VARIETY A Variety is a plant that is only slightly different from the species plant but the differences are not so insignificant as the differences in a form. The Latin is varietas, which is usually abbreviated to var. The name follows the Genus and species name, with var. before the individual variety name. FORM A form is a plant within a species that has minor botanical differences Such as the color of flower or shape of the leaves. The name follows the Genus and species name, with form (or f.) before the individual variety name. CULTIVAR A Cultivar is a cultivated variety a particular plant that has arisen either naturally or through deliberate hybridization, and can be reproduced (vegetatively or by seed) to produce more of the same plant. The name follows the Genus and species name. It is written in the language of the person who described it, and should not be translated. It is either written in single quotation marks or has cv. written in front of the name. Carl Linnaeus (1707-1778) also known as: Carl von Linné Carolus Linnaeus Known as the Father of Taxonomy Taxonomy – systematic classification of plants Devised a method of hierarchical classification binomial nomenclature Named approx. 250,000 plant species Quercus falcata Michx. Linnean herbarium (S-LINN) Department of Phanerogamic Botany Swedish Museum of Natural History (S) Mosses non-vascular plants cannot transport fluids through their bodies rely on surrounding moisture to do this job for them small in stature but very important in our ecosystem prevent erosion, and contribute to the lush green appearance of many forested areas. reproduce by spores, never have flowers, and can be found growing on the ground, on rocks, and on other plants. 24,000 bryophyte species, sometimes grouped into a single phylum are now grouped in three phyla: Mosses (Bryophyta), Liverworts (Hepatophyta) and Hornworts (Anthoceraphyta) Leaf-like Liverworts Mosses Leafy Liverworts Hornworts Ferns second major step in the evolutionary sophistication of plants reproduce by spores like mosses instead of by seed Ferns and allies have a vascular system to transport fluids through their bodies The main phylum, the Ferns (Filicinophyta = Pteridophyta) includes around 12,000 species. Three other phyla are included as fern allies: the Horsetails , 40 species; Club mosses (Lycopodophyta, 1,000 species), and Whisk ferns (Psilophyta, 3 species) Conifers The gymnosperms add the next level of complexity to plant evolution reproduce from seeds instead of spores The seeds, however, are "naked" (Greek: gummnos) - not covered by an ovary Usually, the seed is produced inside a cone-like structure such as a pine cone hence the name "conifer." Some conifers, such as the Yew and Ginkgo, produce their seeds inside a berry-like structure. Conifers Cont’d fairly easy to identify In addition to cones, these trees and shrubs typically have needle-like, scale-like or awl-like leaves And they never have flowers Approximately 600 species of conifers including the pines, firs, spruces, cedars, junipers, and yew Species within the conifer ranks give us pine nuts -- pesto's magic ingredient -- as well as juniper berries for gin. Conifer allies include: Ginkgo (Ginkophyta) with a single species: the Maidenhair Tree (Ginkgo biloba); palm-like Cycads (Cycadophyta) ; and herb-like cone-bearing plants (Gnetophyta) such as Ephedra Angiosperms (Angiospermophyta, Class Dicotyledoneae) the final improvement to plant reproduction they grow their seeds inside an ovary (Greek: angeion = vessel) which is embedded in a flower After it is fertilized, the flower falls away and the ovary swells to become a fruit. Plants in the class Dicotyledoneae grow two seed-leaves (cotyledons). foliage leaves typically have a single, branching, main vein originating at the base of the leaf blade, or three or more main veins that diverge from the base. The vast majority of plants are Dicots. Most trees, shrubs, vines, and flowers belong to this group Approx. 200,000 species. Most fruits, vegetables and legumes come from this class. Angiosperms (Angiospermophyta, Class Monocotyledoneae ) Monocots start with one seed-leaf (cotyledon) the first leaf formed on a seedling) leaves with parallel veins, usually unbranched flower parts occur in multiples of three (three or six petals, three or six stamens, and so on) Approx. 30,000 plants are classified as monocots including many of the prettiest members of kingdom Plantae: orchids, lilies, irises, palms and even the Bird-of-Paradise plant. The grasses which carpet our lawns and meadows are also monocots. Monocots provide us with our primary sources of nutrition, supplying us and the animals we eat with grains such as wheat, oats, and corn, as well as fruits such as dates and bananas. Plant Classification an attempt to organize the hundreds of thousands of species of plants into a meaningful scheme that allows us to make assumptions about the life history, biology, and chemical constituents of various groups. The International Code of Botanical Nomenclature, the rules that govern the naming of plants, currently recognizes seven ranks Each of these ranks is included in one (and only one) higher rank and has one or more members of subordinate rank. International Code of Botanical Nomenclature The seven ranks of corn are: Kingdom: Division: Class: Order: Family: Genus: Species: Complete Classification of Corn The seven ranks of corn are: Kingdom: Plantae Division: Magnoliophyta Class: Liliopsida Order: Cyperales Family: Poaceae Genus: Zea Species: Zea mays The seven ranks indicates something about the corn plant The kingdom Plantae indicates that corn is a plant and probably photosynthesizes using chlorophyll The division Magnoliophyta indicates that it is a flowering plant The class Liliopsida indicates that it is a monocot The order Cyperales indicates that the seeds store starch and the flowers lack petals and sepals The family Poaceae indicates that it has a unique type of inflorescence, round stems, and a distinctive internal anatomy. And so on, down to the species level. Use of Scientific Names Smallest two categories in the classification system Binomial nomenclature Latin and Greek languages (no longer spoken, does not change) Common names confusing and inconsistent Universal recognition, one name per plant species Professional horticulturalists need to know both names Written in italics or underlined Examples: Acer rubrum Acer plantanoides Acer saccharinum Acer palmatum Acer saccharum Acer ginnala Credit is given to the founder: Betula nigra L. Variety vs. cultivar – intentional or unintentional genetic variation Ex: A. rubrum ‘Red Sunset’ or A. rubrum cv. Red Sunset Horticultural Descriptions of Plants Woody vs. herbaceous Evergreen vs. deciduous Semi evergreen Trees vs. shrubs Vines Groundcovers Annuals Biennials Perennials Hardy vs tender plants Nursery plants Greenhouse crops Bedding plants Foliage plants Native plants Exotics Naturalized Aquatic Specimen Accent Tropical How are plants identified? Use of a dichotomous key Each couplet consists of two contrasting statements Based on physical features or taxonomic characteristics of the plant Plant Identification Visual Descriptions of Plants Leaf Types Simple vs. Compound Leaves Shape Margin Lobes Base Vein pattern Apex style Simple vs compound Surface texture Trifoliate Compound Simple Palmately Compound Pinnately Compound Bi-Pinnately Compound Types of Simple Lobed Leaves Tri-Lobed Palmately Lobed Pinnately Lobed Apex 2 Notch 7 Lobe 6 Leaf Blade 1 Midrib 8 Margin 5 Petiole 4 Spine & 9 veins Base 3 Leaf Shapes Oblanceolate Obovate Ovate Lanceolate Oval Linear Orbicular Oblong Leaf Apexes Acute (sharply pointed) Cuspidate (curving to a point) Obtuse (Rounded or Blunt) Emarginate (notched at the tip) Leaf Bases Cuneate Rounded Cordate Truncate Leaf Margins Dentate Entire Serrate Doubly Serrate Crenate-Serrate Crenate Twigs Types of buds Terminal bud Lateral/axillary bud Bud arrangement Leaf petiole scar Lenticles Pith Bud scale scar T 1erminal Bud Lenticels 2 Lateral/axillary bud 3 Internode 4 (space between nodes) Petiole Scar 5 Bud Scale Scar 6 One years growth 7 Vascular Bundle Scar 8 Pith 9 Types of Buds Valvate Scaly (2 scales) Stalked Sessile Imbricate Scaly (w/o stalk) (many scales) Leaf & Bud Arrangements Fascicled Alternate Opposite (clustered on a spur) Whorled Types of piths Hollow Solid Generous Diaphragmed Stellate Chambered Fruits A fruit is the ripened ovary of the flower Different types of flowers produce different kinds of fruit To correctly identify a plant, you must have the fruit or the flower Fruits Fruits are a unique structure found only in angiosperms Fruits develop from the ovary of the flower in addition, may include various accessory structures that derive from other parts of the flower or inflorescence Angiosperm means “seeds in vessels”. The FRUIT IS the vessel Why do angiosperms make fruits? Two major functions of fruits to protect the ovules as they mature into seeds to aid in seed dispersal The chief agents in seed and fruit dispersal are wind, water and animals. Fruit Development Starts with pollination (fertilization) The flower drops its colorful parts The ovary expands to make room for growing seeds (ovules) The ovary matures into a structure known as the pericarp which often differentiates into several distinct tissues layers Perfect Flower vs Imperfect Flower EXOCARP MESOCARP ENDOCARP Fruit Classification Fruit classification is based primarily on the following criteria: Whether the fruit is dry or fleshy Whether the fruit is dehiscent or indehiscent Number of flowers/ovaries that make up the fruit Number of seeds in the fruit Ovary position Family to which the plant belongs Dehiscent Fruit Fruits that split open In a dehiscent fruit, the seeds are the unit of dispersal Dehiscent Fruit Indehiscent Fruit Fruits that do not split open are In an indehiscent fruit, the fruit and seed become an integrated dispersal unit. Indehiscent Fruit Seed Dispersion edible fruits have co-evolved with animals that can disperse seeds Animals become attracted to ripening fruits as they become softer, more fragrant, and higher in sugar Fruits change color to attract birds and mammals, animals large enough to disperse the seeds Types of Fruit Classification of Fruit Types Simple fruit - formed from a single pistil or single flower w/ a single ovary (pea, tomato, lily, apple, cucumber) Aggregate fruit - formed from a cluster of separate pistils borne in a single flower (strawberry, raspberry, magnolia) Multiple fruit - formed from the pistils of several to many flowers consolidated with other floral or inflorescence parts (pineapple, sweetgum, dogwood, fig) Accessory – develops from one or more ovaries and includes the calyx and/or receptacle Perfect Flower vs Imperfect Flower Simple Fruit Aggregate Multiple Examples of Simple, Dry, and Dehiscent Fruit Follicle - A dry dehiscent fruit developed from 1 carpel and at maturity splitting along only one seam Milkweed Examples of Simple, Dry, and Dehiscent Fruit Legume - A dry dehiscent fruit developed from 1 carpel and at maturity splitting along two seams (beans, peas) Examples of Simple, Dry, and Dehiscent Fruit Silique - a special long slender capsule of 2 carpels Splits along two seems This is typical of the mustard family Examples of Simple, Dry, and Dehiscent Fruit Capsule - A dry dehiscent fruit developed from several united (syncarpous) carpels. Lilies, evening primrose, Yucca, poppies Examples of Simple, Dry, and Indehiscent Fruit Achene - A one-seeded, dry, indehiscent fruit; the one seed is free from the pericarp except for a single point of attachment (buttercups, dandelion, sunflower). Examples of Simple, Dry, and Indehiscent Fruit Nut - similar to an achene but with the wall (pericarp) greatly thickened and hardened as in an acorn (beech, chestnut, oak) Examples of Simple, Fleshy Fruit Drupe - A one-seeded simple "stone fruit" developed from a superior ovary innermost portion of the wall, the endocarp, becomes hard and stony the outermost part, the exocarp, becomes a relatively thin skin the middle portion between the skin and the stone, the mesocarp, becomes either fleshy or fibrous (cherry, coconut, olive) Examples of Simple, Fleshy Fruit Berry - A simple fruit in which the ovary wall or at least its inner portions become enlarged and usually juicy. (grape, banana, tomato, watermelon). Examples of Simple, Fleshy Fruit Hesperidium - special type of berry a leathery rind forms; the interior of the fruit divided by septa, indicating the number of carpels. (citrus) Examples of Simple, Fleshy Fruit Pepo - special type of berry relatively hard rind is formed; the interior of fruit not divided by septa. (watermelon, gourds, squash, cucumbers). Examples of Simple, Fleshy Fruit Pome - An accessory fleshy fruit formed by a group of carpels from an inferior ovary and surrounded by and united to the floral tube or receptacle. (apple, pear, mountain ash). Examples of Aggregate Fruit formed from a cluster of separate pistils borne in a single flower The individual units may be berries, achenes, drupes, or other specific types. (raspberry, strawberry, magnolias). Examples of Multiple Fruit formed from the pistils of several to many flowers consolidated with other floral or inflorescence parts (pineapple, sweetgum, dogwood, fig, mulberry) Types of Fruit 1BERRY 4Drupe 2Pome Multiple 3 5 Aggregate 6 Legume Winged 7 Samaras 9Achene Aggregate 8 of Samaras 10 Indehiscent 11 Dehiscent Nut 13 Capsule 12 Follicle 14 Cone Achene Follicle Drupe Berry Coffee Fruit Hesperidiums Pepo Pome Aggregate Legume Capsule Samara Nut Multiple Parts of an Ovary Wall EXOCARP MESOCARP ENDOCARP Flowers What is a flower? reproductive organ of angiosperm plants especially one having showy or colorful parts plant cultivated for its blooms or blossoms The function of a flower? To produce seeds through sexual reproduction For the higher plants, seeds are the next generation Serve as the primary means by which individuals of a species are dispersed across the landscape After fertilization, a flower develops into a fruit containing the seed(s). 4 Main Parts of a Flower Petal Sepal Stamen Pistil 1 6 2 3 7 4 5 Types of Flowers Described by the position of the ovary in relation to the flower parts Described as singular (one flower per stem) Described as composite (multiple small ray and disc flowers (sunflower or daisy) Described as inflorescence (clusters of small flowers arranged on an axis (snapdragons or baby’s breath) Type of flower? Hypogynous Why? Flower parts separate Position of ovary? Superior Type of flower? Perigynous Why? Flower parts fused Position of ovary? Superior Type of flower? Epigynous Why? Flower parts fused Position of ovary? Inferior Perfect Flower vs Imperfect Flower Complete Flower vs Incomplete Flower Monoecious Plants Dioecious Plants Examples of Inflorescences Solitary terminal Solitary – Axillary Spike – w/o stalks Raceme – w/ stalks Panicle – compound raceme Catkin Cyme Corymb Umbel Compound Umbel Flower Inflorescence Corymb Umbel Compound Umbel Cyme A cyme is a flower cluster in which the central or terminal flower blooms earliest. Catkin Environmental Benefits From Trees Settle out, trap, and hold small particles such as dust, ash, pollen, and smoke that can damage lungs Absorb sulfur dioxide and other pollutants Replenish the atmosphere with oxygen Hold the soil with their roots, preventing erosion Provide homes and food for birds and other animals Keep buildings warmer in cold weather by serving as a windbreak and cooler in hot weather by providing shade- lowering energy bills in the process Muffle traffic noise Provide beauty and enjoyment Products From Trees Paper Bark: bottle corks, heat shields for spaceships, tannin to cure leather, baseballs, medicine Cellulose (wood fiber): paper, carpets, plastic film, steering wheels, toothbrush handles, ping-pong balls, wallpaper past, explosives, shampoo, luggage, pillows Gums & Resins: cosmetics, mouthwash, paint thinner, perfumes, soap, coating for pills, rubber gloves, paint Food: allspice almonds apples cinamon coconuts maple syrup olives oranges peaches pistachios walnuts cocoa Handouts Parts of an Ovary Wall 2 1 4 5 3 6 8 7 9 10 11 2 7 6 1 8 5 9 3 4 1 2 3 6 4 7 5 8 9 Leaf Type Types of Simple Lobed Leaves Tri-Lobed Palmately Lobed Leaf Shapes Leaf Apexes Leaf Bases Leaf Margins Types of Buds Leaf & Bud Arrangements Types of piths