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Speciation Text Handout pgs 336-342. 1. What is speciation? Speciation - the origin of new ________________ from pre-existing species. 2. What is a species? Biological Species = A set of naturally genetically reproductively isolated from other sets of populations. populations that are 3. Other concepts/definitions of species exist. “Species” can sometime be hard to define – e.g. _______________ 4. There are 2 main types of speciation: 1) ______________________ and 2) _____________________ (“patric” means land or area. “Allo” pertains to different locations and “sym” means same) *both followed by/involve events that isolate species and include differences in genes 5. Allopatric speciation = evolutionary change occurring in Ancestral population divides; each can undergo geographic ranges. evolutionary change. 6. Sympatric speciation = evolutionary occurring in same (overlapping) geographic ranges. May occur by: - Initial selection (e.g., different food sources). Local ecological _________________ specialization (e.g., races/ecotypes) Polyploidy mutation (chromosomes ______________________) 7. Reproductive Isolating Mechanisms (genetic) • Polyploidy = evolution of chromosome no. that is multiple of an ancestral set. • Hybridization of 2 species followed by polyploidy ----> _____________________________________________ • Polyploid hybrid from both parents. Spartina Salt Marsh Grass, Cord Grass S. maritima (Europe & Africa, native species) X S. alterniflora (e. North America, introduced to Europe) 2n = 60 2n = 62 HYBRIDIZATION (+ loss of one chromosome) Polyploid Speciat ion: S. X townsendii 2n = 62 CHROMOSOME DOUBLING (+ loss of 1 chromosome pair) S. angelica NEW POLYPLOID 4n = 122 SPECIES 8. Other Reproductive Isolating Mechanisms PRE-ZYGOTIC (pre-mating) i) Habitat isolation - differences in _______________ preference ii) Temporal isolation - differences in ____________________ of reproduction iii) Behavioral isolation - differences in behavioral ______________________ with respect to finding and attracting mates PRE-ZYGOTIC (post-mating) iv) Mechanical isolation - differences in _______________________________, don’t “fit” v) Gametic isolation - _____________/___________ incompatibility POST-ZYGOTIC vi) zygotic mortality – mating and fertilization possible, but genetic differences result in ___________ that dies. E.g. sheep and goat mate but zygote is not viable. vii) Reduced hybrid viability - embryo doesn’t . viii) Reduced hybrid fertility - hybrids develop but ________________. Tigon - Result of male tiger and female lion mating incaptivity. Offspring are ____________________. Liger - Result of male lion and female tiger mating in captivity. Offspring are ____________________. Adaptive Radiation - spreading of populations or species ____________________________________________, with adaptive evolutionary divergence. Promoted by: 1) New and varied niches (different _____________________ available) - provide new selective pressures 2) ___________________of interspecific competition - enables species to invade niches previously occupied by others Question: How can the separation of 2 populations lead to a) the formation of new species, b) a reduction in genetic diversity of the populations and a possible threat to their survival?