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Anatomy & Physiology SIXTH EDITION Chapter 17, part 1 The Special Senses Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Frederic H. Martini Fundamentals of Learning Objectives • Describe the sensory organs of smell, and trace the olfactory pathways to their destination in the brain. • Identify the accessory and internal structures of the eye, and explain their function. • Explain how light stimulates the production of nerve impulses, and trace the visual pathways to their destination in the brain. • Describe the structures of the external and middle ear and explain how they function. Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Learning Objectives • Describe the parts of the inner ear and their roles in equilibrium and hearing. • Trace the pathways for the sensations of equilibrium and hearing to their destinations in the brain. Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings SECTION 17-1 Olfaction Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Olfactory organs • Contain olfactory epithelium with olfactory receptors, supporting cells, basal cells • Olfactory receptors are modified neurons • Surfaces are coated with secretions from olfactory glands • Olfactory reception involved detecting dissolved chemicals as they interact with odorant binding proteins Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.1 The Olfactory Organs Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.1a, b Olfaction • Olfactory pathways • No synapse in the thalamus for arriving information • Olfactory discrimination • Can distinguish thousands of chemical stimuli • CNS interprets smells by pattern of receptor activity • Olfactory receptor population shows considerable turnover • Number of receptors declines with age Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings SECTION 17-2 Gustation Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Taste receptors • Clustered in taste buds • Associated with lingual papillae Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Taste buds • Contain basal cells which appear to be stem cells • Gustatory cells extend taste hairs through a narrow taste pore Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.2 Gustatory Reception Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.2 Gustatory pathways • Taste buds are monitored by cranial nerves • Synapse within the solitary nucleus of the medulla oblongata • Then on to the thalamus and the primary sensory cortex Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Gustatory discrimination • Primary taste sensations • Sweet, sour, salty, bitter • Receptors also exist for umami and water • Taste sensitivity shows significant individual differences, some of which are inherited • The number of taste buds declines with age Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings SECTION 17-3 Vision Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Accessory structures of the eye • Eyelids (palpebrae) separated by the palpebral fissue • Eyelashes • Tarsal glands • Lacrimal apparatus Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.3 Eternal Features and Accessory Structures of the Eye Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.3a, b external structures of the eye • Conjunctiva covers most of eye • Cornea is transparent anterior portion Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Lacrimal apparatus • Secretions from the lacrimal gland contain lysozyme • Tears form in the lacrimal glands, wash across the eye and collect in the lacrimal lake • Pass through the lacrimal punctae, lacrimal canaliculi, lacrimal sac and nasolacrimal duct Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings The eye • Three layers • Outer fibrous tunic • Sclera, cornea, limbus • Middle vascular tunic • Iris, ciliary body, choroid • Inner nervous tunic • Retina Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.4 The Sectional Anatomy of the Eye Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.4a, b internal structures of the eye • Ciliary body • Ciliary muscles and ciliary processes, which attach to suspensory ligaments of lens • Retina • Outer pigmented portion • Inner neural part • Rods and cones Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.4 The Sectional Anatomy of the Eye Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.4c Figure 17.5 The Pupillary Muscles Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.5 retina • Retina contains rods and cones • Cones densely packed at fovea (center of the macula lutea) • Retinal pathway • Photoreceptors to bipolar cells to ganglion cells, to the brain via the optic nerve • Axons of ganglion cells converge at blind spot (optic disc) • Horizontal cells and amacrine cells modify the signal passed along the retinal neurons Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.6 The Organization of the Retina Copyright © 2004 Pearson Education, Inc., publishing as Benjamin Cummings Figure 17.6a