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APES: Chapter 9 Population Dynamics, Carrying Capacity, Conservation Biology *INTRODUCTION: 9-1 Population Dynamics and Carrying Capacity Populations are dynamic, constantly undergoing changes in size, density, and age distribution due to environmental stress and changing environmental conditions. In general there are four factors that influence the size of a population: birth (+), deaths (-), immigration (+), and emigration (-). Different populations vary in their ability to grow (biotic potential). The intrinsic rate of increase refers to a population’s growth rate if it had access to unlimited resources. 1 Organisms with high intrinsic rates of increase typically have the following characteristics: - early reproductive age - short generation times - can reproduce several times - have many offspring each time The intrinsic rate of increase of many species depends upon having a certain minimum population size (minimum viable population). Passenger Pigeon Reading Environmental resistance consists of the factors that prevent a population from reaching its intrinsic rate of increase. 2 The carrying capacity refers to the number of individuals that can be supported in a given area in a sustainable manner. Populations with unlimited access to resources grow slowly at first and at a faster rate with time (exponential growth). When graphed this yields a J-shaped curve. As populations experience environmental resistance the rate of growth begins to slow as the carrying capacity is reached (logistic growth). When graphed this yields an Sshaped curve (see Fig. 9-4). 3 At times a population may overshoot the carrying capacity and eventually suffer a population crash or dieback (See Fig. 9-6). Human populations are not exempt from dieback although technology and cultural changes have increased the carry capacity of the earth for our species. http://www.youtube.com/watch?v=9_9SutNmfFk human population growth HW #1: page 201: #’s 2, 5, 6 (all parts), 7, 8 http://www.youtube.com/watch?v=BSVbdaubxxg section summary 4 **DENSITY FACTORS Some controls on populations have an impact independent of the population’s density (density independent controls): - floods - fire - severe weather - habitat destruction - pesticides Other controls have a greater impact if the density of the population is large (density dependent controls): - competition for resources - infectious disease (bubonic plague) - parasitism Overall there are four common patterns of population fluctuations: - stable: the population stays pretty close to the carrying capacity. - Irruptive: populations explode to high levels then return to a stable level or crash. - Chaotic: irregular fluctuations that are not predictable - Cyclic: population’s size varies on a fairly regular basis. 5 2- Predation and Population Size The role of predators in ecosystems has been studied extensively. At one time most biologists accepted the top-down control hypothesis. Predators (higher trophic level) controlled the population size of their prey (lower trophic level). This idea was used to explain the cycling of hare and lynx populations studied since the 1800’s (see Fig 9-8). 6 It is now believed the size of some populations might be better explained using the bottom-up control hypothesis. The size of a herbivore population (higher trophic level) is determined by how much food they consume or the quality of the food source (lower trophic level). Although some bottom-down controls are still believed to exist, the size of predatorprey populations are probably a combination of both factors. HW #2: page 201 #’s: 9, 10, 11, 12. ***9-3 Reproductive Patterns All organisms are constantly struggling for genetic immortality. Remember, Darwinian Fitness is a measure of the reproductive success of an organism or population. 7 There is not however one reproductive strategy that is always better than another. Asexual reproduction involves no exchange of genetic information and the offspring are genetic clones of the original cell. Binary Fission Budding Sexual reproduction involves an exchange of genetic information between organisms and is estimated to occur in 97% of all life. 8 There are many costs associated with sexual reproduction 1- time consuming 2- disease 3- genetic errors (recombination) 4- predation risks ... but these are outweighed by the advantage of generating genetic diversity. Scientists have identified two fundamental reproductive patterns: r- selected (opportunists): put most of their energy and resources into reproduction. - many small offspring but few reach sexual maturity - little if any parental care - high intrinsic rates of growth - early successional species - generalist niche K- selected: tend to do well in competitive situations when their population is near the carrying capacity. - few large offspring 9 - late reproductive age - extensive parental care - typically follow a logistic growth pattern - because the intrinsic rate of growth is low they are prone to extinction No matter what pattern organisms used their ultimate population size is determined by available suitable habitat. ****Survivorship Curves Survivorship curves indicate the number of individuals alive in a population during a given time. There are three generalized patterns: Late loss curves: common in K-selected species (most die late) 10 Early loss curves: typical for r-selected species (most die early) Constant loss curves: mortality rates are constant at all age levels HW #3: page 201 #’s: 13, 14, 15, 16. 9-4 Conservation Biology Conservation biology began in the 1970’s in attempt to use science to preserve species and ecosystems. There are several fundamental questions they ask: 1- what species are in danger of extinction? 2- what is the status of functioning ecosystems and the services they provide? 3- what actions can be taken to sustain ecosystems, their services, and their populations? 11 Conservation biology is heavily dependent upon field research and focuses upon three essential principles: 1- biodiversity is essential to all life and should be preserved 2- humans should not hasten the extinction of species or disrupt ecological processes 3- the best way to preserve biodiversity and ecological functions is to protect intact ecosystems The new discipline of bioinformatics has contributed enormously to the field of Conservation Biology by providing access to key biological information through databases. *****Human Impacts on Nature Humans have used technology to modify nature extensively in numerous ways: 1- fragmentation and degradation of habitats 12 http://www.youtube.com/watch?v=KEINq3-9Tu0 2- simplified natural ecosystems 3- use and/or waste an increasing percentage of the planet’s net primary productivity 4- strengthen some pest species 13 5- eliminate predators 6- introduce non-native species http://www.youtube.com/watch?v=2iKYcqKyXic Answer the following questions as you watch the video. 1. How did Burmese Pythons arrive in the Everglades? 2. What is the economic cost? 3. What is the ecological cost? 4. How has ballast water contributed to the invasive species problem? 5. Describe one impact (ecological/economic) zebra mussels have had on the great lakes. 7. over-harvest natural resources 8. interfere with the normal cycling of chemicals and the flow of energy in ecosystems Solutions Many biologists now believe the best way for us to live more sustainably is to learn how nature sustains itself and then to mimic these strategies. These lessons include: 1- we are totally dependent upon the earth and sun 2- everything is connected 3- we can never merely do one thing 14 4- we should reduce the damage we are doing to nature and heal some of the ecological wounds we have caused 5- we should use care, restraint, humility, and cooperation with nature as we alter it to meet our needs. APES: Chapter 12 Human Population: Growth, Demography, and Carrying Capacity 12-1 Factors Affecting Human Population Demography is the study of the changes and influences on the human population. http://www.worldometers.info/worldpopulation/ 15 http://www.npr.org/2011/10/31/141816460/visu alizing-how-a-population-grows-to-7-billion The rate at which the human population changes is frequently measured by subtracting the crude death rate (the number of deaths per 1,000) from the crude birth rate (number of births per 1,000). This is usually expressed as a percentage using the following equation: % change = birth rate- death rate X 100 1,000 which equals: birth rate- death rate 10 Even though the birth rate has fallen around the world the death rate has fallen much faster. Exponential growth has slowed in the last 40 years but the base population has still almost doubled. Calculation: student notes Problem: Calculate the number of people that would be added to a population of 3 billion people at a fertility rate of 5%: 16 Problem: Calculate the number of people that would be added to a population of 6 billion people at a fertility rate of 2.6 %: Two types of fertility rates affect a country’s population size and growth rate. 1- replacement fertility level: the number of children a couple must have to replace themselves (approximately 2.1 children). Even when (if?) this number was to be achieved the global population would continue to grow for another 50 years (assuming death rates stay the same) POPULATION MOMENTUM. 2- total fertility rate (TFR): the average number of children a women will have in her lifetime. TFR has dropped significantly since 1950 from about 5.0 to 2.6. Notice however it is still far above the replacement level of 2.1. http://www.youtube.com/watch?v=uOuzC0UMBHQ fertility rates 17 More than 95% of the human population growth by the year 2050 is expected to take place in developing countries where living in acute poverty (less than $1.00 a day) is a way of life. -----------------------------------------------------There are many factors that affect fertility rates: - importance of children in the labor force - urbanization - cost of raising and educating children - educational and employment opportunities for women - infant mortality rates 18 - average age at marriage - availability of abortions - availability of reliable birth control methods - religious beliefs, traditions, cultural norms. Death Rates are influenced by access to: food supply - medical care medical and health technology improved sanitation safe water (#1) Two common ways of measuring the overall health of people in a country are: 1- life expectancy- the average number of years a newborn can expect to live http://www.youtube.com/watch?v=p6qhwUd9uw0 life expectancy 19 2- infant mortality rates (commonly defined as death before the age of one) Although infant mortality rates are falling, almost 22,000 infants die each day due to preventable causes. 20 There are 37 countries that have lower infant mortality rates than the United States due mainly to: inadequate health care for the poor drug addiction during pregnancy high birth rates among teenage girls (the highest of any industrialized country). MINI HW #4Review Questions (page 275 #’s 2, 15, 17 12-2 Population Age Structure Diagrams Age structure diagrams show the proportion of a population (or gender) at a particular 21 age. They are typically divided into three broad categories: 1- prereproductive (ages 0-14) 2- reproductive (14- 44) 3- postreproductive (45 and up). Age structure diagrams that have a triangular shape have the potential to grow rapidly. About 30% of the world’s population is in the prereproductive group. The more rectangular the diagram becomes the slower the birth rate. 22 Rapid changes in population growth rates can have severe cultural impacts (Social Security in this country). If population decline is gradual the harmful effects can usually be managed. HIV/AIDS is having a major impact of the age structure of many African countries. Life expectancy may drop 35-40 years with the greatest impact on the most productive adults. Increases in the number of orphans and reduction in food production are expected to follow. The international community, especially developed countries, are being called upon to spearhead the effort to slow the spread of AIDS through improved education, health care, family planning, and contributing economic aid. 23 12-3 Solutions There are several proposed solutions to the problem of human population growth: 1. limiting migration (the United States allows large numbers to immigrate compared to many of the other countries of the world.) 2. reducing birth rates through economic incentives, family planning, and empowering women. Many question the need to limit birth rates arguing: a- we do not know the carrying capacity of the earth b- humans are the world’s most valuable resource when it comes to solving global problems c- it violates their private or religious beliefs. 3- Economic development typically reduces birth rates in a fairly recognizable pattern 24 known as demographic transition. You should be familiar with this model in detail. 4- A small number of countries use economic rewards or penalties to help reduce birth rates. China has instituted the most extensive and intrusive controls and arguably the most effective. Mini HW #5: Review Questions page 275 #’s: 8, 9, 12, 14. 25 26