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Created by Julia Hsu Levy – Version 1.5 Chapter 28: Animals Animal characteristics: 1. Muticellular 2. Heterotroph 3. Eukaryotes 4. NO cell wall 5. Two unique tissue types: muscle (movement) and nervous (response) 6. Mostly sexual reproduction these stages: Cleavage: many rounds of mitotic cell division Blastula: hollow ball structure Gastrulation: part of embryo pinches in, forming gastrula (pouch) Specific tissue types result (endoderm, ectoderm, sometimes mesoderm) 7. Some undergo metamorphosis (juvenile larva pupa adult) 8. Hox gene controls animal development from embryonic stage 9. Most have symmetry (except sponges that are assymetrical) 10. Most are acoelomate, pseudocoelomate, or coelomate Those with symmetrical bodies these associations: have Dorsal: top Ventral: bottom Anterior: head Posterior: tail 161 Created by Julia Hsu Levy – Version 1.5 * * * Evolutionary History of Animals * * * May have evolved from flagellated protists Classification P. Porifera P. Cnidaria P. Ctenophora P. Platyhelminthes P. Nematoda P. Annelida P. Mollusca C. Gastropoda C. Bivalvia C. Cephalopoda P. Arthropoda C. Arachnida C. Insecta P. Echinodermata C. Agnatha C. Chondricthyes C. Osteichthyes C. Amphibia C. Reptilia C. Aves C. Mammalia Marsupial Monotreme Placental Invertebrate Animal Kingdom Survey Common examples Notable traits Sponges Asymmetry, pores, filter feeder, parazoa (lacks true tissues, only animal like this) Jellyfish, sea anemone, hydra Tentacles, radial symmetry, filter feeder, nematocyst, eumetazoa (has true tissues) Comb jelly Similar to cnidarian Flatworm, tapeworm Bilateral symmetry, flat body, aceolomate, mouth only Roundworms, pinworms, Bilateral symmetry, round hookworms body, pseudocoelomate, mouth and anus Segmented worms, Bilateral symmetry, found earthworms, leeches body, coelomate, metanephridia Body has 3 parts: foot, Snails visceral mass, mantle Clams, oysters, mussels Squid, octopus Millipedes, centipedes Exoskeleton, molting Spiders, ticks Insects Malpighian tubules Starfish, sand dollar Water vascular system, tube feet Vertebrate Animal Kingdom Survey Jawless fish Parasitic on fish Shark, skates, rays Cartilaginous, paired fins Trout, bass, salmon, etc Boned fish, swim bladder, operculum Frogs, toads, salamanders, Close ties with water, respires newts through skin, metamorphic Lizards, crocodiles, alligators Amniotic egg, ectotherm Birds Light hollow bonds, feathered Mammary glands, hair Kangaroo Pouch Platypus Egg-laying Humans placenta 162 Created by Julia Hsu Levy – Version 1.5 * * * Animal Structure and Function * * * Anatomy: study of the structures of an organism Physiology: study of the functions of an organism Atoms molecules molecular structures cell tissues organs organ systems Animal tissues Epithelial: forms outer covering and organ linings Connective: sparse cells in extracellular matrix Muscle: long, excitable cells that can contract Nervous: senses stimuli and transmits electrical signals Epithelial tissue: tight junction connections between cells protects against injuries classified according to arrangement of cells Connective tissue: bind and support other tissue loose, adipose (fatty), fibrous, cartilage, bone, blood Muscle tissue: parallel bundles of microfilaments in cytoplasm (actin and myosin) most abundant tissue in animals ex: cardiac, striated, visceral Nervous tissue: neuron cell = cell body + dendrites +axons 163 Created by Julia Hsu Levy – Version 1.5 * * * Homeostasis * * * Regulating the internal environment is dependent upon control systems. two 1. Negative feedback: opposite effect of whatever that is happening (ex: human body heat) 2. Positive feedback: amplifies (increases) the same response (ex: oxytocin and labor) 164 Created by Julia Hsu Levy – Version 1.5 * * * Digestive System * * * Energy derived from food input digestion absorption catabolism Metabolic rate: total amount of energy an animal uses per unit of time (measured in kcal or CAL) measuring amount of O2 animal uses per unit time measuring animal’s heat loss per unit time minimal rate: needed to support life maximal rate: during peak activity rate determined by age, sex, weight, body temperature, etc. Endotherm: generate own body heat Ectotherm: generates body heat from metabolically environment Ex: birds and mammals Ex: fish, amphibians, reptiles, and Basal metabolic rate (BMR): metabolic invertebrates rate measured under resting, fasting, and Standard metabolic rate (SMR) stress-free conditions determined at specific temperature under resting, fasting, and stree-free conditions Why is eating important? o Chemical energy o Building blocks for other materials o Essential nutrients ANIMAL NUTRITION: Most use some combination of extracellular and intracellular digestion. 1. Ingestion: eating 2. Digestion: breaking food down (chemically or physically) into smaller particles and molecules 3. Absorption: uptake of molecules 4. Elimination: passing out of digestive system undigested materials Undernourished: not enough nutrients Overnourished: too many nutrients (severe obesity) Malnourished: improper nutrients (missing essential nutrients) 165 Created by Julia Hsu Levy – Version 1.5 166 Created by Julia Hsu Levy – Version 1.5 1. Mouth: ingestion and digestion (salivary enzymes) 2. Esophagus: peristalsis motion 3. Stomach: digestion (peptic enzymes), controlled with sphincter muscle 4. Small intestine: chemical digestion and absorption 5. Large intestine: water absorption, E. coli digestion and vitamin production 6. Anus: elimination, controlled with sphincter muscle Alimentary canal: continuous tube with two openings (mouth and anus) Feeding Mechanisms o Suspension feeders: sift small food particles from water (ex: clam, oysters) o Substrate feeders: live in or on food source (ex: leaf miners) o Deposit feeders: eating way through dirt (ex: earthworms) o Fluid feeders: sucking nutrient-rich fluids from living host (ex: mosquitoes, leech) o Bulk feeders: eat large pieces of food (ex: humans) Hydra captures prey with stinging tentacles. The prey is pushed into a gastrovascular cavity where enzymes are released. Phagocytosis and intracellular hydrolysis complete digestion. The protist Paramecium uses only intracellular digestion, where chemicals secreted break down food particles.. 167 Created by Julia Hsu Levy – Version 1.5 Crop and stomach = storage Gizzard = grinder Pharynx = throat Intestine = chemical digestion Anus = elimination Ruminents: (1) When the cow first chews and swallows a mouthful of grass, boluses enter the rumen and (2) the reticulum. • Symbiotic bacteria and protists digest this cellulose-rich meal, secreting fatty acids. • Periodically, the cow regurgitates and rechews the cud, which further breaks down the cellulose fibers. (3) The cow then reswallows the cud, which moves to the omasum, where water is removed. (4) The cud, with many microorganisms, passes to the abomasum for digestion by the cow’s enzymes. 168 Created by Julia Hsu Levy – Version 1.5 * * * Circulatory System * * * CIRCULATION: Diffusion is too slow to move materials. Circulation reduces the amount of time necessary to transport vital chemicals and gases. Open circulatory system: hemolymph (body fluid) bathes internal organs directly while moving through sinuses Closed circulatory system: blood is confined to vessels and distinct interstitial fluid present Circulatory system: heart + blood vessels + blood Heart: cardiac muscle Comparing Circulatory Systems 2-chambered Ex: fish One atrium, one ventricle 3-chambered Ex: amphibians Two atria, one ventricle 4-chambered Ex: humans Two atria, two ventricles Systole: heart contracts Diastole: heart relaxes, chambers fill with blood Heart rate: number of heartbeats per minute Sinoatrial (SA) node: controls heart muscle 1. Oxygenated blood out of left ventricle to the body. 2. O2 into body, CO2 into blood 3. Deoxygenated blood back to heart into right atrium, right ventricle, and to the lungs. 4. CO2 exhaled, O2 inhaled 169 Created by Julia Hsu Levy – Version 1.5 5. Oxygenated blood back to heart into left atrium, left ventricle (repeat #1). Arteries away from heart Veins back to heart Blood pressure: hydrostatic force that blood exerts against a vessel wall Capillary exchange: exchange of materials between blood through capillary walls and interstitial fluid Blood components: 1. Plasma: fluid for blood, electrolytes+proteins+90% water 2. Red blood cells: erythrocytes 3. Leukocytes: white blood cells 4. Platelets: clotting factors 170 Created by Julia Hsu Levy – Version 1.5 * * * Respiratory System * * * Gas exchange: movement of O2 and CO2 between animal and environment Gas exchange over entire body surface Ex: protozoan and unicellular organisms Plasma membrane of every cell has contact with outside environment Ex: sponges, flatworms, cnidarians Isolated cells from environment, respiratory surfaces are generally thin with moist skin over bed of capillaries Region of body surface with lots of folds to increase surface area for gas exchange Ex: humans Ex: earthworms, amphibians Diaphragm: muscle that raises and lowers to decrease and increase lung capacity Medulla senses pH of blood. Increased CO2 levels drops pH. Lower pH raises rate of breathing to exhale excess CO2. Hemoglobin: four subunits each with heme group that bonds O2 with an iron atom CO2 transported in blood dissolved in plasma, bound to hemoglobin, or as bicarbonate ion. 171 Created by Julia Hsu Levy – Version 1.5 * * * Immune System * * Robert Koch was the first to connect certain diseases to specific bacteria (anthrax, TB) Koch’s postulates: (1) The researcher must find the same pathogen in each diseased individual investigated, (2) Isolate the pathogen form the diseased subject and grow the microbe in pure culture, (3) Induce the disease in experimental animals by transferring the pathogen from culture, and (4) Isolate the same pathogen from experimental animals after the disease develops. 172 Created by Julia Hsu Levy – Version 1.5 * * * Controlling Internal Environment * * * Osmoconformers: animals that do NOT actively adjust internal osmolaroty Ex: saltwater animals, body fluids isotonic with the environment Osmoregulators: animals that regulate internal environment by regulating water intake and excretion Ex: freshwater animals, all terrestrial animals, many saltwater animals Planarian excretion: * * * Excretory System * * * Earthworm excretion: Human nephrons: Insect excretion: 173 Created by Julia Hsu Levy – Version 1.5 Flatworms: fluid passes through flame bulb, cilia brushes into tubule, fluid drains into excretory ducts Earthworm: fluid passes through metanephridium and empties into storage body outside the body through nephridiopore Insect: waste removed from hemolymph into tubules, waste accumulates, water follows, fluid passes to hindgut to rectum, salts and water reabsorbed, solids excreted Nitrogenous waste: 1. Ammonia: aquatic animals 2. Urea: mammals and most adult amphibians, requires lot of water to dilute or otherwise toxic 3. Uric acid: land snails, insects, birds, and reptiles (paste-like consistency) Mammalian excretory system: nephrons 1. filter blood: Bowman’s capsule, glomerulus 2. secretion: proximal and distal tubules 3. reabsorption: tubules and loop of Henle 174 Created by Julia Hsu Levy – Version 1.5 * * * Endocrine System * * * Hormones; endocrine or neurosecretory cell secretions that affect a target cell Neurosecretory ell: nrueons that receive signals from other nerve cells and release hormones as a result Target cell: cell that can respond to a particular hormone Endocrine gland: ductless glands that secrete hormones Exocrine gland: glands that produce a variety of substances and send products through ducts (ex: sweat, mucus, etc) 175 Created by Julia Hsu Levy – Version 1.5 * * * Reproductive System * * * Parthenogenesis: egg development with fertilization Hermaphroditism: solution for animals that have difficulty finding mates o Sequential hermaphroditism: an individual reverses its sex during its lifetime. (ex: clearner shrimp) Asexual reproduction (mostly in invertebrates) o Fission: asexual reproduction in which a parent separates into two or more approximately equal sized individuals. o Budding: asexual reproduction in which new individuals split off from existing ones. o Gemmules of sponges are an example of a type of asexual reproduction that involves the release of specialized cells that can grow into new individuals. o Fragmentation: the breaking of the body into several pieces, some or all of which develop into complete adults. o Requires regeneration of lost body parts. Sexual reproduction: o Internal fertilization: favorable conditions for copulation (quality over quantity) o External fertilization: moist habitat to protect eggs from drying out (quantity over quality) Pheromones: chemicals that attract mates (ex: bombykol of female moth) Sperm anatomy: Middle piece = lots of ATP to flagella supply Acrosome: enzymes that digest the outer layer of the egg during fertilization away Produced in the testes (gonad) the scrotum held in Fertilization takes place in the Fallopian tubes (oviduct) 176 Created by Julia Hsu Levy – Version 1.5 Oogenesis: Spermatogenesis: Female hormones: Male hormones: 177 Created by Julia Hsu Levy – Version 1.5 * * * Nervous System * * * 1. Resting state: sodium channels closed, + outside and - inside 2. Threshhold: all or nothing response, inside much reach certain + voltage before response can “fire” 3. Depolarization state: sodium channels open, Na+ ions flood inside the nerve cell, inside goes + while outside goes + 4. Repolarization state: sodium channels close, Na+ blocked but K+ let in, makes inside – against and outside + 5. Undershoot: Na+ continues to flood out with K+, membrane voltage becomes very negative, creates the refractory period Repeat for the the next section of the nerve axon. 178 Created by Julia Hsu Levy – Version 1.5 Synapses: spaces between two nerve cells that receive electrical or chemical signals to fire the next neuron Examples of neurotransmitters: acetylcholine, epinephrine, norepinephrine, dopamine, serotonin, gases (NO, CO) Nerve nets: Cephalization: head development 179 Created by Julia Hsu Levy – Version 1.5 Cerebral cortex: thoughts Medulla oblongata: reptilian brain (controls basic body functions – breathing, heart rate, etc) Cerebellum: balance Pituitary gland: master gland that produces hormones Hypothalamus: controls the pituitary gland 180 Created by Julia Hsu Levy – Version 1.5 * * * Skeletal-Muscular System * * * Sliding-filament model . 1. At rest tropomyosin blocks the myosin binding sites on actin. 2. When calcium binds to the troponin complex a conformational change results in the movement of the tropomyosin-tropinin complex and exposure of actin’s myosin binding sites. 3. Myosin sites on actin bind to tropomyosin, forming a cross-bridge. 4. The two filaments slide, causing a muscle contraction. 181