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Name and Date _________________________ Section # ______________________________ Lab 3. Seed, Seedling and Plant; Tissues and Vegetative Structures The plant body is composed of two integrated organ systems: the shoot system and the root system. The shoot system is designed for energy and carbon capture through photosynthesis, as well as for reproduction. Shoots are composed of stems, leaves and flowers, which all have a critical function and a unique structure. The root system is designed for anchorage, storage and the harvesting of both minerals and water from soils. The root system, too, has structurally different organs designed to fulfill a particular function. Today in lab you will start by studying the external structure (morphology) of a whole plant body from embryos in the seed to the development of mature organs systems in the seedling and mature plant. You will also study the diversity of external form in plants and novel modifications of stems and leaves. 1. Seed, Seedling and Plant Choose one of the plants available today in lab to trace the development of the root and shoot systems from its’ embryonic origin in the seed to fully developed leaves stems, flowers and roots in a mature plant. Seeds contain embryonic shoot and root axes as well as storage material for the germinating seedling. In most dicots this storage occurs in the fleshy embryonic seed leaves (cotyledons). In monocots, the storage material is often in the form of endosperm and is extra-embryonic (i.e. found outside of the embryo). The cotyledon in monocots is often used to absorb the energy stored in the endosperm and then passes into the rapidly growing seedling. Carefully dissect acorn and bean seed to observe the embryonic axis, cotyledons and storage materials. Sketch and label the dissected seeds in the space below. Seedlings show initial developmental forms, which may have several juvenile aspects to them. When compared to seedlings, mature plants may have very different leaf sizes or shapes and may show different branching patterns in both the root and shoot system. In the space below sketch both a seedling and a mature plant. Highlight developmental differences in leaf, stem and root morphology. Label your drawing. 2. Vegetative Structures Winter twigs can be used to display the morphological features found on most woody stems. Observe the different winter twigs on display in lab today and compare the variation in morphological features shared by each twig. Identify structures such as the terminal and axillary buds, nodes and internodes, leaf scars and lenticels. Notice that these species vary considerably in their outer appearance. Most shrubs and trees are unique enough for field identification based on their twig morphology alone. In the space below, sketch 2 or 3 terminal buds and leaf scars from different species. Identify the species of winter twigs with the key provided. Twig # Species 2 Leaf morphology and arrangement can vary widely between species of plants. A typical leaf will be constructed of a blade designed for photosynthesis and a stem-like structure called the petiole that attaches the blade to the stem. In the upper notch between the petiole and the stem is an axillary bud, which is a dormant shoot and is diagnostic of a leaf. Of course there are many exceptions to the typical leaf and you will be looking at the wide variation in leaf types and morphology in lab today. In the space provided below, sketch one of the leaves on display in lab today. Represent as best as you can, leaf characteristics such as venation, margin type and leaf arrangement. Observe the different examples of leaf, stem and shoot morphology on display in lab today. Can you find examples of different leaf shapes, arrangements and venation? Use the table below to organize your observations. Species arrangement Leaf shape Venation Leaf margin 3 3. Modifications of Leaves and Shoots Leaves and shoots are sometimes modified in plants to achieve a variety of functions. For example, in grapes, peas, cucumbers and many other vines, the blade or the petiole is modified into a climbing structure called a tendril. Tendrils will wrap around anything that they touch and thus will allow plants to climb other plants, trellises, fences, stakes etc. In some cases leaves are modified for protection as is the case of the spines found on cacti. Spines are modified leaves whereas other structures, such as rose prickles, are modified from the epidermis and thorns are modified from stems (such as those on Hawthorns). Carnivorous plants modify their leaves to act as insect traps. Carnivorous plants generally grow in nutrient poor soils and the trapped insects serve as an extra source of nitrogen and phosphorus. Plants will sometimes modify their stems to act as dispersal or over wintering structures. Stolons are modified stems that run along the surface of the ground and will produce new plants from the axillary buds at its’ nodes. Rhizomes are underground stems that can store starch reserves that will be utilized by the growing plant the following spring. New plants will also arise from the axillary buds located at its’ nodes. Notice the examples of modified leaves and stems. Fill in the table below with the name of the modified structure (eg stolon), its developmental origin (e.g. a modified stem) and its function (e.g. over wintering structure) Species Name of structure Modified from what structure function 4 4. Root Systems Plant roots systems fall into two general categories: tap root systems and fibrous root systems. Within each of these architectural types there is tremendous variation in the extent to which roots branch and the depth to which they penetrate soils. The categories stem from a developmental difference which is established shortly after the species germinates from its seed. When the embryonic root axis (primary root) persists and grows into a strong leader root, a tap root system develops. On the other hand if the primary root is short lived and the root system is dominated by branching lateral roots, then a fibrous root system results. In general tap root systems penetrate deeper into the soil than fibrous root systems, which are more likely to dominate in shallower soil depths. In many species roots can become swollen with storage products, usually starch. Examples of storage roots that form from tap roots include carrots, rutabagas and turnips. It is also common for fibrous roots to store starch as is the case with begonias, dahlias and sweet potatoes. Study the root systems made available in today’s lab. Look for the presence of the primary root and the relative abundance of lateral roots in each plant. Make representative sketches in the space below. Make sure to label your drawings. 5