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Rate of Photosynthesis Geoff Klein April 11 As light intensity increases, the rate of photosynthesis also increases until a certain point. At that point, the light-saturation point, the rate of photosynthesis has maxed out and can not proceed any faster. This is because of other limiting factors like amount of CO2 available, the number of enzymes available and other factors necessary for photosynthesis to occur. This graph shows the rate of photosynthesis from the affect of light by observing the amount of CO2 consumption. When there is no light, there is no activity in the cell. What this causes is for CO2 to actually leave the cell by diffusion. The intensity of light has a direct effect on the rate of photosynthesis. In terms of temperature, there is no significant effect to photosynthesis as a whole, but temperature does effect the Calvin cycle. As temperature exceeds 30 degrees Celsius, the Calvin cycle rate begins to decline fast due to the denaturing of enzymes. Between 10 and 30 degrees Celsius is the optimal temperature for maximum efficiency. The higher the oxygen concentration, the slow photosynthesis becomes. As oxygen levels increase, competition for rubisco increases because both oxygen and carbon dioxide can react with the active site. When CO2 binds, it becomes fixed into organic molecules through the Calvin cycle. When O2 is bound, photorespiration occurs and photosynthesis is reduced in activity. Photorespiration occurs when there is more oxygen than carbon dioxide in the cell. The oxygen is bound to rubisco creating glycolate then carbon dioxide. The reason rubisco can react with two different substrates is because the early prehistoric atmosphere consisted of mostly carbon dioxide. When the atmosphere started to change to mostly oxygen, the function for reacting with oxygen was created through evolution. Net CO2 uptake = Photosynthetic CO2 uptake – photorespiratory CO2 evolution – respiratory CO2 evolution Net O2 evolution = Photosynthetic O2 evolution – photorespiratory O2 uptake – respiratory O2 uptake Cellular respiration ATP Photosynthesis - ATP is used in energy investment - Used to make final product, phase of glycolysis G3P - Produced through cellular respiration - Produced in noncyclic and cyclic electron flow NAD+/ - 2 NAD+ used in glycolysis/ pyruvate - Used in stage 2 (non-cyclic NADP+ oxidation, and 6 used in Krebs cycle electron flow) when NADPH reductase reduces it to go to Calvin cycle - Produced in Calvin cycle CO2 - Produced in cellular respiration - Used in photosynthesis Glucose - Used in cellular respiration - Produced by photosynthesis as a result of G3P H+ H2O - Pumped out into inter-membrane - Pumped into thylakoid space to produce ATP lumen to create ATP - Produced in glycolysis and ETC - Used in photosynthesis Review Questions Q: What effect does increasing light intensity have on photosynthesis? A: It causes the rate of photosynthesis to increase until the light-saturation point. Q: How does the increase of oxygen concentration effect photosynthesis? A: As oxygen concentration increases, the rate of photosynthesis decreases.