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3/1/2016 Morphology form) Plant biology, perhaps the oldest branch of science, is driven by a combination of curiosity and need curiosity about how plants work need to apply this knowledge judiciously to feed, clothe, and house a burgeoning human population. (body shoot and root systems inhabit two environments • shoot (aerial) stems, leaves, flowers • root (subterranean) taproot, lateral roots vascular tissues transport materials between roots and shoots • xylem/phloem • A Root – Is an organ that anchors the vascular plant – Absorbs minerals and water – Often stores organic nutrients – Taproots found in dicots and gymnosperms – Lateral roots (Branch roots off of the taproot) – Fibrous root system in monocots (e.g. grass) (b) Storage roots (a) Prop roots (c) “Strangling” aerial roots (d) Buttress roots A stem is an organ consisting of – Nodes (could be opposite or alternate) – Internodes (e) Pneumatophores (a) Stolons Storage leaves (d) Rhizomes Stem Node Root Bulbs (c) Tubers Rhizome Root 1 3/1/2016 An The axillary bud form a lateral shoot, or branch A leaf • Is the main photosynthetic organ of most • Is a structure that has the potential to vascular plants terminal bud Leaves • Is located near the shoot tip and causes generally consist of • Blade elongation of a young shoot • Stalk • Petiole Gardening tip: Removing the terminal bud stimulates growth of axillary buds classifying angiosperms Three Tissue Systems In – Taxonomists may use leaf morphology as a criterion Petiole Axillary bud (b) Compound leaf. dermal tissue • epidermis (skin) single layer of cells that covers entire body waxy cuticle/root hairs (a) Simple leaf vascular tissue ground tissue Leaflet • xylem and phloem transport and support Petiole Axillary bud • mostly parenchyma occupies the space b/n dermal/vascular tissue photosynthesis, storage, support (c) Doubly compound leaf. Leaflet Petiole Axillary bud Carries Protects plant from: • Physical damage • Pathogens • H2O loss (Cuticle) out long-distance transport of materials between roots and shoots Consists of two tissues • xylem • phloem 2 3/1/2016 • • • Any tissue that isn’t dermal or vascular Includes various cells specialized for storage, photosynthesis, and support Pith = ground tissue internal to the vascular tissue Cortex = ground tissue external to the vascular tissue Xylem (water) dead at functional maturity tracheids- tapered with pits vessel elements- regular tubes Vessel Parenchyma St: “typical” plant cells. Most abundant Fu: perform most metabolic functions Ex: fleshy tissue of most fruit • • WATER-CONDUCTING CELLS OF THE XYLEM Anatomy (internal structure) • division of labor • cells differing in structure and function – 5 types • parenchyma, collenchyma, sclerenchyma Collenchyma Sclerenchyma St: grouped in cylinders; thick Fu: provide support but allow growth in young parts of plants Ex: celery St: hardened secondary walls Fu: specialized for support; dead Ex: fibers (hemp/flax) Meristems • perpetually embryonic tissues located at regions of growth • divide to generate additional cells (initials and derivatives) • apical meristems (primary growth- length) Phloem (food) alive at functional maturity sieve-tube members arranged end to end with sieve plates located at tips of roots and shoots • lateral meristems (secondary growth- girth) SUGAR-CONDUCTING CELLS OF THE PHLOEM Sieve-tube members: longitudinal view Tracheids Pits Companion cell Sieve-tube member Sieve plate Tracheids and vessels Nucleus Vessel element Tracheids Cytoplasm Companion cell Cortex Vascular cylinder Epidermis Key to labels Primary of Roots Growth • Includes apical meristem Plants grow in length (roots and shoots) Zone of elongation Zone of maturation • Elongation of cells • Cell differentiation • Cell become functionally Dermal Root hair Zone of differentiation Ground Vascular • New cells produces • Root cap is located in Zone of cell division • Includes apical meristem • New cells produces • Root cap is located in root Zone of cell division root Zone of elongation Zone of maturation Zone of elongation • Elongation of cells • Cell differentiation • Cell become Apical meristem functionally mature Zone of cell division Root cap mature 100 µm 3 3/1/2016 Primary Growth in Shoots Epidermal Tissue • upper/lower epidermis • guard cells (stomata) Apical meristem (1, 7) Located at buds of shoots Site of cell division that allows plant to grow in length Ground Tissue • Mesophyll • palisade/spongy parenchyma Vascular Tissue • Veins • xylem and phloem Leaf Anatomy Guard cells Key to labels Dermal Ground Vascular Cuticle Stomatal pore Epidermal cell Sclerenchyma fibers 50 µm (b) Surface view of a spiderwort (Tradescantia) leaf (LM) Stoma Upper epidermis Palisade mesophyll Bundlesheath cell Lateral Meristems Spongy mesophyll Lower epidermis Cuticle Guard cells Xylem Phloem (a) Cutaway drawing of leaf tissues Vein Guard cells The vascular cambium As a tree or woody shrub ages Vein Air spaces Guard cells (c) Transverse section of a lilac (Syringa) leaf (LM) 100 µm • vascular cambium + produces secondary xylem/phloem (vascular tissue) • cork cambium + produces tough, thick covering (replaces epidermis) • secondary growth + occurs in all gymnosperms; most dicot angiosperms – Develops from parenchyma cells • The older layers of secondary xylem, the heartwood, no longer transport water and minerals The outer layers, known as sapwood • Still transport materials through the xylem 4 3/1/2016 Periderm - protective coat of secondary plant body cork cambium produces cork cells secondary growth commences farther down the shoot transforms older regions first 5