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UNIT 5: Population Dynamics 12.5: Interactions between Individuals pg. 616 - 628 In ecosystems, species form populations, there are more then on species living in an ecosystem, therefore there are many populations. These populations interact and form communities: therefore there are intraspecific and interspecific relationships occurring. These interactions can be beneficial, harmful, or neutral, may or may not affect the individual’s ability to survive or reproduce, and is a product of long-term evolutionary adaptation. When individuals of two different species exert selective pressures (reproductive success) on each other, influencing their evolution is known as coevolution. Co-evolution – is a process in which one species evolves in response to the evolution of another species. Co-evolutionary relationships can take on many forms, for example: Predation – is the interaction between a predator and a prey where the predator gains nutrients and energy form the prey while killing it. Herbivory – is the interaction between herbivorous animals and the plants that they eat. Mutualism – is an interaction n which both partners benefit. Parasitism – is an interaction in which on species benefits and the other is harmed. Commensalism – is an interaction in which one species benefits and the other is unaffected. Table 1: Population Interactions and their Effects Interaction Effects on Inte4racting Example Populations Predation Herbivory Mutualism Parasitism +/- Predators gain nutrients and energy; prey are killed or injured +/- Herbivores gain nutrients and energy; plants are killed or injured +/+ Both populations benefit +/- Parasites gain nutrients; hosts are injured or killed -/- Both competing populations lose access to some resources Commensalism -/0 One population benefits; the other is unaffected Competition Northern lynx (predator) and snowshoe hare (prey) White-tailed deer (herbivore) and foliage Honey bee (pollinator) and flowering plants Mistletoe, which attaches to a tree and takes water and nutrients from its host; usually stunts growth but can kill the tree with heavy infestation Trees in a forest competing for light Moss, which grows on a tree, getting the light and nutrients it needs while the tree is unaffected Predation and Herbivory Feeding relationships, food chain, and food webs indicate the movement of nutrients and energy from producer through to top carnivore. Animals acquire their nutrients and energy by consuming other organisms, predation and herbivory are factors of co-evolution, which has allowed predators and herbivores to evolve characteristics that enable them to feed effectively. Food abundance affects food choice. If a prey is scares, then the predator will take what it can get, creating a higher cost-to-benefit ratio. If there is an abundance of food then, the predator will specialize, and select prey with largest energy return. Defence Mechanisms Defence mechanisms include camouflage, behavioural defences, chemical defences, spines and armour, and mimicry. Any adaptation that is used by prey increases its chance of survival, giving the prey an advantage against its predators. Camouflage allows an organism to mimic the patterns of its environment, making it hard to be identified by its predator Behavioural defences can passive or active. Standing still is a passive approach while fleeing is an active response. Chemical defences make the organism unattractive to the predator. The prey can release noxious odours or releasing poisonous chemicals. Spine and Armour, are used by some plants and animals. The use of hard thorny like structures prevents the predator from harming its prey. Mimicry occurs when an organism mimics another organism which the predator finds unpalatable. There are two types of mimicry; Batesian and Mullerian. Batesian, a palatable organism mimics an organism which in unpalatable. For Mullerian, two or more unpalatable organisms that share common predators mimic each other. Mimicry – is a form of defense in which one species evolves an appearance that resembles the appearance of another species. Population Cycles A predator prey relationship is an interspecific interaction and dependent on the density of population. As the density of the prey population increases, so will the predator population follow, since there is more food to support an increase. As the prey population decreases through over feeding, the predator population will decrease because of lack of resources and an increase in competition for those resources. The populations will be dynamic and continue to fluctuate. There is a lag in response, the prey population will either increase or decrease and the predator population will lag in responding to these changes. There are other factors that can affect the cycling of a population, such as; catastrophic natural events, changes in climate, and human interference, non-native species introduction and habitat loss. Figure 4: a) A model of the predator-prey cycle; b) Lynx and hare populations through time. Competition When different species use the same limiting resources will experience interspecific competition, this type of interaction will increase mortality rates and decrease reproduction. Population size and growth rates of each of the competing species populations are reduced over time. There are two main forms of interspecific competition: interference competition and exploitative competition. Interference Competition –is competition in which all the individuals have equal access to resources; the fitness of some individuals is reduced by the presence of others. Exploitive Competition – is competition in which all the individuals have equal access to resources; some have superior ability to gather resources. Biological fitness, also called Darwinian fitness, means the ability to survive to reproductive age, find a mate, and produce offspring. Basically, the more offspring an organism produces during its lifetime, the greater its biological fitness. Competition and Niches Alfred J. Lotka and Vito Volterra both proposed a model of interspecific competition, independently, describing the effects of competition between two species, where if one specie increased in population size, it would negatively affect the growth rate of the other species. G.F. Gause, tested the model experimentally, using two different specie of Paramecium; aurelia and caudatum. He grows each paramecium species in separate cultured medium, controlling their environments similarly. Each specie of paramecium showed rapid population growth until they met their carrying capacity, than maintained population size. Gause then mixed the two specie of Paramecium, aurelia and caudatum, in one cultured medium. Aurelia competed with caudatum. The aurelia population increased, while the caudatum population did not. He identified the results as the Competitive Exclusion Principle. ”Populations of two or more specie can not coexist indefinitely if they rely on the same limiting resources and exploit these resources in the same way. One species inevitably harvests resources more efficiently; (fitness) produces more offspring than the other, and by its actions, negatively affects the other species.” Ecological niche is an organism’s biological characteristics, including use of and interaction with abiotic and biotic resources in its environment. This concept allows ecologists to visualize resource use and the potential for interspecific competition in nature. The niche includes food, shelter, and nutrients, as well as the abiotic conditions, such as light, and temperature. Ecologists can study competition between two populations by plotting their fundamental niches and realized niches, if their fundamental niches overlap, then they may compete for resources. Fundamental Niche – is the range of conditions and resources that a population can possibly tolerate and use. Realize Niche – is the range of conditions and resources that a population actually uses in nature. Figure 5: G.F. Gause performed experiments to observe interspecific competition in Paramecium. a) and b) When each species was raised alone, both species reached a stable population size, c) When the species were raised together, Paramecium aurelia persisted while the population of Paramecium caudatum declined Resource Partitioning Resource Partitioning – is a situation in which several species that live in the same place use different resources or use the same resources in different ways. Organisms may compete for resources, or they may avoid competition by partitioning resources, which allows for the co-existence of different species in the same ecosystem. Figure 8: a demonstration of competition between two species of barnacles. A) Before the experimental treatments, both species exist in their realized niches. B) In the absence of Balanus, Chthamalus occupies both shallow water and deep water. C) In the absence of Chthamalus, Balanus still occupies only deep water. Symbiosis In biology there are three types of symbiotic relationships recognized; Mutualism, commensalism, and parasitism. Mutualism (+/+) – both partners will benefit from their relationship, for example the relationship between plants and insects. Bees support the pollination of plants while the bees obtain the nectar secrete from the flower. In this situation both organisms exploit each other for their common good. Commensalism (+/0) – One partner benefits while the other neither benefits or is harmed. Commensalism is rare in nature because the interaction between two organisms rarely leaves one unaffected. Parasitism (+/-) – During this type of interaction one organism will benefit while the other is negatively affected, and receives no benefit. Parasitic relationships could also be considered specialized predator – prey relationships, one organism is feeding on the other. Disturbing Ecosystem Relationships Human activities, such as the introduction of non-native species and habitat destruction are upsetting the dynamic equilibrium in many ecosystems. The relationships between organisms in an ecosystem are dynamic, organism compete for resources, are parts of food chains and food webs. Interactions between two organisms will affect not only themselves but also other species within the ecosystem. The greatest disruption in an ecosystem is created by the introduction of a non-native invasive species or the loss of a species. Non-native invasive species enter an ecosystem where they do not have a natural predator to control their population growth. The can disperse quickly, establish themselves, and dominate the ecosystem. They will compete for limiting resources, which will negatively impact native species, decreasing their population or possible extinction. Human impact, such as habitat destruction is currently ranked as the primary cause of species extinction worldwide. Because of the interaction between species in an ecosystem, when one organism becomes extinct it has a direct impact on the entire ecosystem. Chapter 12: Summary pg. 632 Chapter 12: Self-Quiz pg. 633 Chapter 12: Review pg. 634 - 639