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PHYLUM NEMATODA y Nematodes, also known as round worms, are among the simplest animals to have a complete digestive system; ie: they have both a mouth and an anus. y Nematoda is the second largest of the animal phyla, and its members are more numerous than those of any other phylum of multicellular animals. Habitat y Most are free – living and are found in virtually all parts of the earth. Many are parasites that affect almost every kind of plant and animal. The variety of habitats occupied is greater than all other animal groups except arthropods. Structure y Elongate, cylindrical body that contains a fluid-filled body cavity called the pseudocoelom between the body wall and the digestive tract. y The pseudocoelom acts as both a hydrostatic skeleton upon which the muscles can act, and a transportation system for nutrients, oxygen and wastes. y Tapered body, more blunt at the front end, is covered with a tough cuticle. Ingesting Food y Free-living forms are often carnivorous and catch and eat small animals with their muscular sucking pharynx. Some feed on algae, fungi, and pieces of decaying organic matter in the soil. Others are parasitic and feeding on living plant and animal tissues. y May have a piercing stylet for sucking out and digesting cell contents, or sharp teeth and cutting plates for digesting intestinal linings. Digesting Food y Food is broken down both chemically and physically in the straight digestive tract that opens through the mouth and exits through the anus. Food moves through the intestine by ingestion of more food and the general movements of the worm’s body, rather than by muscular contractions. Elimination y Undigested food exits via the anus. Muscles surrounding the anus relax, and the waste is forcefully expelled. Very efficient unidirectional movement of food through the digestive system is a significant evolutionary step. Internal Transport y Nutrients, oxygen and wastes are transported through the fluid in the pseudocoelom. No internal transport system exists, so the roundworms depend on diffusion to move materials through their body. Respiration y Breathe through their body walls; rely on diffusion, no internal transport system. Excretion y The digestive tract seems to play a major role in excretion, and the excretory system probably function in osmotic regulation. Excretory wastes will also move by diffusion through the pseudocoelom. Locomotion y Depends on the opposing forces of the longitudinal muscles, elastic cuticle, and hydrostatic skeleton. Range of motion is limited by the lack of circular muscles and the way the nervous system controls muscle contraction: the body only bends in the dorsoventral plane resulting in a whip like thrashing motion. Sensing Environment y A variety of sense organs in the form of tactile bristles and chemosensory pits can be found. Simple photoreceptors have been found in some marine and freshwater nematodes. Coordination y Nervous system consists of a nerve ring and ganglia, around the pharynx, from which nerves run both forward and backward. Nerve ring is the site of most sensory integration. Usually 4 major nerve cords. Reproduction y Reproduce sexually. Most species have separate males and females, but a few are hermaphroditic. Fertilization takes place inside the body of the female. y As is usual, parasitic species have complex life cycles that can involve several hosts.