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Name _____________________________________________________ Test Date _____________________ UNIT XI – EVOLUTION (chapter 15) I. THE THEORY OF EVOLUTION The theory of evolution is one of the most fundamental concepts in Biology. Evolution is defined as _____________ in a ________________________________ over time. The scientist considered to be the founder of modern evolutionary theory is Charles Darwin. A. History of Evolutionary Theory – During the 1700s, several scientists began challenging the idea of a world in which changes did not occur. These scientists and their hypotheses were very important to Darwin’s work. 1. Gradualism - ________________ first suggested that the planet was much _____________ than previously thought; began to find evidence that ________________ were slowly, but constantly taking place. 2. Malthus – Published an essay that had a huge impact on Darwin. Proposed that organisms ___________- reproduce; in other words, reproduce at a ____________________ rate than resources can supply. 3. Lamarck Lamarck was one of the first scientists to propose a mechanism for evolution; that is, the ______________ in a _______________ of organisms over ___________. His major hypotheses included: Tendency Toward Perfection – Stated that organisms were continually changing in order to _______________________________________________. Use and Disuse – Changes in _____________ and/or _____________ of a structure in an organism was a response to use or disuse. Structures used extensively _____________________ and structures used less frequently __________________. Inheritance of ________________Traits B. Charles Darwin (1809 – 1882) 1. Darwin’s History Darwin’s data was collected on a 5-year journey around the world on the HMS _________________ He made observations and collected data throughout the journey. He used this data to propose a ________________ to explain the diversity he saw. The area that had the greatest impact on Darwin was the _______________ Islands due to the differences he saw in the same animals living on different islands. Ex: _______________________________________________________________________________. He began to suspect that populations from the mainland changed after reaching the Galapagos. Upon his returned he talked to animal breeders & called what they did to direct breeding to produce offspring with the desired traits ________________________, same as what we called selective breeding last unit. 2. Darwin’s 4 Basic Principles (Observations) Members of a population often vary greatly in their _______________. Variations (traits) can be inherited from __________________ to __________________. All species are capable of producing _____________ offspring than _________________________________ Variations that increase reproductive success will have a greater chance of being passed on than those that do not increase reproductive success. 3. Darwin’s Theories – Based on his observations and the hypotheses of other scientists Organisms with favorable ____________ tend to survive and ______________; thereby leaving more descendents than other individuals This will result in an accumulation of these traits in the ________________, changing the original _____________ make-up of the population 4. Darwin’s Legacy Did not publish his findings for years Alfred Wallace – formed identical ___________________ based on his research. Sent his manuscript to Darwin, and finally Darwin was persuaded to publish his own conclusions Released “___________________________________________________” still considered one of the greatest scientific studies ever II. THE PRINCIPLES OF EVOLUTIONARY THEORY A. Evolution occurs because of natural selection - a mechanism for change that occurs when organisms with ________________ characteristics for a particular environment __________________, _______________, and pass these characteristics on to __________________. B. The ability of an organism to survive and reproduce in its environment is known as _________________. C. Fitness is based on ___________________. An adaptation is any trait that aids in the ______________ and __________________ of an organism. Examples of adaptations are _______________________________________________________________________ D. As organisms _______________ and adapt, ________________ may occur. Speciation is the formation of new species - a group of similar organisms that ________________ with one another and produce ______________________________________________________________________!. For speciation to occur population must diverge & then be reproductively isolated. E. Two types of Speciation: Allopatric speciation: physical barrier divides on population into two or more: EX: ___________________________________________________________ Sympatric speciation: species evolves into new one without a physical barrier. Seen in insect & plant species. F. Isolating Mechanisms allow for the gene pools to become _________________ so they can form a new _____________. Reproductive Isolation-as a new ________________ evolves, populations become ____________________ isolated from each other. Prezygotic –prevents reproduction by making _________________________ because of geographic, ecological, behavioral, temporal or other differences EX: __________________________________ have overlapping ranges & are similar in appearance but use different mating songs & do not interbreed. EX: Fireflies- similar species but mate at different _______________________________ EX: Trout- different species in same stream but mate diff. time of year Postzygotic- when fertilization has occurred but a _______________ offspring _____________________ or reproduce; prevents offspring survival or reproduction EX: Mule, liger G. The failure of an organism to __________________to changes in its environment will ultimately lead to its __________________ because of __________________________. III. EVIDENCE FOR EVOLUTION A. Fossils – Fossils are _ _______________________________________________ Fossils provide a record of earlier life and evidence that evolution has occurred. Fossils also provide evidence about the earth’s _________________, geography, and _________ forms. Almost _____% of Earth’s history occurred during the __________________ Time. There are two major classes of Traits when studying transitional fossils:derived traits-newly evolved features such as feathers, that do not appear in the fossils of common ancestors. Ancestral traits are more primitive features such as teeth & tails, that do appear in ancestral one B. Biogeography – Variations are seen in the same types of animals based on their ______________________ In addition, there are some organisms that live in very different locations but they have similar characteristics because _____________________________________________________________ C. Comparative anatomy – Scientists use anatomical studies of different organisms for evidence of evolutionary relationships. 1. Homologous structures- For example, appendages that are very _______________ in structure, but differ in ___________________ are known as _____________________ structures. Examples of homologous structures are __________________________________________________ 2. Vestigial Structures - A structure that is ______________ in function in a living organism, but may have been used by an ancestor is known as a ______________ structure. A structure may become vestigial when an organism changes in form or behavior. Examples are ______________________________________________ 3. Analogous structures- Not all similar features are evidence of common ancestry. Analogous structures can be used for the ________________ & can be superficially similar in construction but are ________________ from a common ancestor. They show that functionally similar features can evolve independently in similar environments. Ex. Wings of birds vs. insects D. Embryology – Similarities in the structures of developing ___________________ of different organisms are considered to be proof of a close evolutionary relationship. E. Comparative biochemistry – Scientists use DNA studies to determine the evolutionary relationship between organisms. The more similar the DNA, _____________________________________ IV. MECHANSIMS FOR EVOLUTION Evolution does not occur in an individual; instead it refers to _______________________ that occur in a population over time. There must be mechanisms available for _____________________ changes to occur: A. Mutation – A mutation is a _____________________. Although mutations are most often _________________, sometimes the resulting change in ________________ may be beneficial to an organism under certain conditions. If the change occurs in the __________________ this change will be passed onto the offspring of that organism. A positive mutation that provides a survival advantage is known as an _____________________ B. Diploidy – Most organisms are diploid, which means ___________________________________ This allows for increased genetic variation in a population. Heterozygote Advantage – Seen in ________________________ and _______________. Recombination - Leads to increased genetic variation as a result of ___________________ during ______________ of ______________. C. Gene Flow – Gene flow occurs when organisms from one community migrate to another. This introduces new ________________ into the _______________________ which can lead to a change in the genetic make-up of the population. D. Genetic Drift – This describes a situation in which change in a population is magnified because the population size is very ___________. Causes of genetic drift include: Bottleneck effect – large portion of population ______________________________ Founder effect – segment of population moves to new __________________________ E. Single Gene Traits-Natural selection on ___________________ traits can lead to changes in _________________ frequencies and therefore to evolution. EX: _____________________________________________ F. Polygenic Traits-Natural selection can affect the distributions of phenotypes in 3 ways: Directional Selection-Individuals at ______ end of the bell curve have higher fitness than those in the middle or other end. ____________________________________________________________________________. Stabilizing Selection-Individuals in the _________ have the highest fitness causing the curve to narrow. _______________________________________________________________________________________. Disruptive Selection-when individuals at the _______ of the curve have the higher fitness. If lasts long enough can cause the curve to split and create ______ distinct phenotypes. _________________________________ _______________________________________________________________________________________ V. MACROEVOLUTION- refers to large-scale evolutionary pattern and processes that occur over long periods of time. A. Extinction-More than ______% of all species that have ever lived are now ___________, which means the species has died out. Darwin proposed possible reasons with competition for _________________________________________ ________ extinction has occurred several times, wiping out entire ___________________________ ________________, volcanic action, and _______________________ movement have been blamed. Mass extinctions clear the way for _____________________ of other species. EX: B. Adaptive radiation or divergent evolution- the process where a __________ species has evolved through ______________________ __________ into diverse forms in short period of time that live in different ways. Often follows mass extinctions EX: ___________________________________________________________________________________ C. _______________________________ when unrelated species evolve similar traits even though they live in different parts of the world because of similar ecology & climate ___________________________________________________________________ D. __________________________ is when a change in one organism leads to a ________________________________change in another organism. EX: ___________________________________ E. ________________________ Equilibrium is another pattern of evolution. Unlike ____________________________, punctuated equilibrium is characterized with long periods of ____________________ interrupted by brief periods of ____________________ change. It is controversial but known that evolution does occur at different rates. 1.